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<document id="992B01331162005736D9BF5EE45EB6F4" ID-CLB-Dataset="8531" ID-DOI="10.11646/zootaxa.4851.2.5" ID-GBIF-Dataset="7beefd33-c83a-4e05-82d1-7522b65c112c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4407593" ID-ZooBank="C0027037-9243-40E5-AAE1-22A9A272BD14" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609433360249" checkinUser="plazi" docAuthor="Park, Kyu-Tek &amp; Bae, Yang-Seop" docDate="2020" docId="03D987EDD946E0464BFEFA5C798AFC30" docLanguage="en" docName="zootaxa.4851.2.5.pdf" docOrigin="Zootaxa 4851 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torodora cordisigna Park &amp; Bae 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="313" masterDocId="FFE0FF95D940E04E4B69FFB37810FFDC" masterDocTitle="Genus Torodora Meyrick (Lepidoptera, Lecithoceridae, Torodorinae) in Laos with descriptions of five new species and two newly recorded species" masterLastPageNumber="318" masterPageNumber="305" pageNumber="311" updateTime="1698850697182" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="625B6EC379D60BF1EB8F0497F962FA4B">Genus Torodora Meyrick (Lepidoptera, Lecithoceridae, Torodorinae) in Laos with descriptions of five new species and two newly recorded species</mods:title>
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<mods:namePart id="94EAAEA663E3216E0613F7BB24249A75">Park, Kyu-Tek</mods:namePart>
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<mods:affiliation id="660FF79ADAAC31653849CE4B7F8CF1E9">BioResource and Environmental Center, Division of Life Sciences, Incheon National University, Incheon, 22012 Korea ktpark 02 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9933 - 4497</mods:affiliation>
<mods:nameIdentifier id="1A9E64BBD4484686E3D6C66740FB1975" type="email">ktpark02@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6431093D54E16759D11162DC70F1D2CD">Bae, Yang-Seop</mods:namePart>
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<mods:affiliation id="8B4A14E9218EE522A178B0CCD4BBB278">BioResource and Environmental Center, Division of Life Sciences, Incheon National University, Incheon, 22012 Korea ktpark 02 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9933 - 4497 &amp; Division of Life Sciences, Incheon National University, Incheon, 22012 Korea</mods:affiliation>
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<mods:date id="A268643F30C3EB03879667A924F73FCF">2020</mods:date>
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<treatment id="03D987EDD946E0464BFEFA5C798AFC30" ID-DOI="http://doi.org/10.5281/zenodo.4407961" ID-GBIF-Taxon="177203856" ID-Zenodo-Dep="4407961" LSID="urn:lsid:plazi:treatment:03D987EDD946E0464BFEFA5C798AFC30" httpUri="http://treatment.plazi.org/id/03D987EDD946E0464BFEFA5C798AFC30" lastPageId="8" lastPageNumber="313" pageId="6" pageNumber="311">
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<paragraph id="8BCF36FBD946E0484BFEFA5C7A2FF9D5" blockId="6.[151,575,1519,1582]" box="[151,575,1519,1546]" pageId="6" pageNumber="311">
<heading id="D0878197D946E0484BFEFA5C7A2FF9D5" bold="true" box="[151,575,1519,1546]" fontSize="11" level="1" pageId="6" pageNumber="311" reason="1">
<emphasis id="B904EAE9D946E0484BFEFA5C7A2FF9D5" bold="true" box="[151,575,1519,1546]" pageId="6" pageNumber="311">
<taxonomicName id="4C704D78D946E0484BFEFA5C79C4F9D5" ID-CoL="9FPX8" authority="Park" authorityName="Park &amp; Bae" authorityYear="2020" box="[151,468,1519,1546]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="311" phylum="Arthropoda" rank="species" species="cordisigna" status="sp. nov.">
<emphasis id="B904EAE9D946E0484BFEFA5C799DF9D6" bold="true" box="[151,397,1519,1546]" italics="true" pageId="6" pageNumber="311">Torodora cordisigna</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="A2375792D946E0484A88FA5C7A2FF9D5" box="[481,575,1519,1545]" pageId="6" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<paragraph id="8BCF36FBD946E0484BFEF9A07933F9F2" blockId="6.[151,575,1519,1582]" box="[151,291,1555,1582]" pageId="6" pageNumber="311">
(
<figureCitation id="134B2A7ED946E0484BF6F9A078FBF9F2" box="[159,235,1555,1582]" captionStart="FIGURE 5" captionStartId="7.[151,250,1883,1907]" captionTargetBox="[160,1426,186,1842]" captionTargetId="figure-19@7.[151,1436,181,1859]" captionTargetPageId="7" captionText="FIGURE 5. Torodora cordisigna Park, sp. nov.: A, holotype; B, labial palpus in lateral view; C, hind tibia; D, forewing venation; E, female genitalia, gen. slide no. CIS-8077, holotype, with close-up signum. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407608" httpUri="https://zenodo.org/record/4407608/files/figure.png" pageId="6" pageNumber="311">Figs. 5</figureCitation>
A-F)
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D946E0484BFEF9E878C7F961" pageId="6" pageNumber="311" type="materials_examined">
<paragraph id="8BCF36FBD946E0484BFEF9E878C7F961" blockId="6.[151,1436,1627,2014]" pageId="6" pageNumber="311">
<emphasis id="B904EAE9D946E0484BFEF9E8795CF9A9" bold="true" box="[151,332,1627,1653]" pageId="6" pageNumber="311">Type material.</emphasis>
<materialsCitation id="3B183CA6D946E0484A33F9E878C7F961" ID-GBIF-Occurrence="3028414312" collectingDate="2015-11-09" collectorName="YS Bae &amp; Bae, BS &amp; Park, SM &amp; Na, JW &amp; Kim, DJ" country="Laos" elevation="68" latitude="18.503056" location="Phou Khao Khouay National Protected Area" longLatPrecision="21" longitude="102.96833" pageId="6" pageNumber="311" specimenCount="1" stateProvince="Bolikhamxai" typeStatus="holotype">
<typeStatus id="54CB8859D946E0484A33F9E879D7F9A9" box="[346,455,1627,1653]" pageId="6" pageNumber="311" type="holotype">Holotype</typeStatus>
: female,
<collectingCountry id="F367766BD946E0484950F9E87A96F9A9" box="[569,646,1627,1653]" name="Laos" pageId="6" pageNumber="311">LAOS</collectingCountry>
:
<collectingRegion id="49B4F819D946E04849FEF9E87B60F9AA" box="[663,880,1627,1654]" country="Laos" name="Bolikhamxai" pageId="6" pageNumber="311">Bolikhamsai Prov.</collectingRegion>
,
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,
<geoCoordinate id="EE44503CD946E0484BFEF9CC7933F945" box="[151,291,1663,1689]" degrees="18" direction="north" minutes="30" orientation="latitude" pageId="6" pageNumber="311" precision="15" seconds="11" value="18.503056">18°3011N</geoCoordinate>
<geoCoordinate id="EE44503CD946E0484A45F9CC79DCF945" box="[300,460,1663,1690]" degrees="102" direction="east" minutes="58" orientation="longitude" pageId="6" pageNumber="311" precision="15" seconds="06" value="102.96833">102°58 06E</geoCoordinate>
,
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<elevation id="005DD1C8D946E0484AB0F9CC7A05F946" box="[473,533,1663,1690]" metricMagnitude="1" metricUnit="m" metricValue="6.8" pageId="6" pageNumber="311" unit="m" value="68.0">68 m</elevation>
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,
<date id="FFCE103BD946E048494BF9CC7A81F946" box="[546,657,1663,1690]" pageId="6" pageNumber="311" value="2015-11-09">
<collectingDate id="EF8AE9D3D946E048494BF9CC7A81F946" box="[546,657,1663,1690]" pageId="6" pageNumber="311" value="2015-11-09">9 xi 2015</collectingDate>
</date>
, leg.
<collectorName id="2685532DD946E04849A4F9CC7B37F946" box="[717,807,1663,1690]" pageId="6" pageNumber="311">YS Bae</collectorName>
, BS
<collectorName id="2685532DD946E0484837F9CC7C9EF946" box="[862,1166,1663,1690]" pageId="6" pageNumber="311">Park, SM Na, JW Kim, DJ</collectorName>
Lee; gen. side no. CIS- 8077.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D946E0484BAEF9747CA1F801" pageId="6" pageNumber="311" type="diagnosis">
<paragraph id="8BCF36FBD946E0484BAEF9747CA1F801" blockId="6.[151,1436,1627,2014]" pageId="6" pageNumber="311">
<emphasis id="B904EAE9D946E0484BAEF9747952F93D" bold="true" box="[199,322,1735,1761]" pageId="6" pageNumber="311">Diagnosis.</emphasis>
This new species is characterized by the forewing venation: R
<subScript id="17F434BED946E0484898F9697BE9F934" attach="left" box="[1009,1017,1754,1768]" fontSize="6" pageId="6" pageNumber="311">3</subScript>
and R
<subScript id="17F434BED946E0484F56F96A7C49F93B" attach="left" box="[1087,1113,1753,1767]" fontSize="6" pageId="6" pageNumber="311">4+5</subScript>
unusually with long stalk, R
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and R
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short stalked, arising near from R
<subScript id="17F434BED946E048493EF94D7A4FF8D0" attach="left" box="[599,607,1790,1804]" fontSize="6" pageId="6" pageNumber="311">3</subScript>
at base. The forewing color pattern is similar to the following new species,
<taxonomicName id="4C704D78D946E0484BFEF8BC7982F8F5" authority="Park &amp; Bae, 2020" authorityName="Park &amp; Bae" authorityYear="2020" box="[151,402,1807,1833]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="311" phylum="Arthropoda" rank="species" species="multispinata" status="sp. nov.">
<emphasis id="B904EAE9D946E0484BFEF8BC7982F8F5" box="[151,402,1807,1833]" italics="true" pageId="6" pageNumber="311">Torodora multispinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2375792D946E0484AF3F8A379E1F8F6" box="[410,497,1808,1834]" pageId="6" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
with a minute, yellowish-white costal patch beyond 3/4 of the costa as in the latter, but it can be distinguished from a narrow basal line on the fringe of the forewing. The hind tibia has more distinct white scales at apex. The female genitalia are differentiated from those of any congeners by having a heart-shaped signum in the female genitalia. It is more or less similar to that of
<taxonomicName id="4C704D78D946E048481DF8CF7CBDF849" authority="Wu &amp; Liu, 1994" authorityName="Wu &amp; Liu" authorityYear="1994" box="[884,1197,1915,1942]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="311" phylum="Arthropoda" rank="species" species="sciadosa">
<emphasis id="B904EAE9D946E048481DF8CF7BE1F849" box="[884,1009,1915,1941]" italics="true" pageId="6" pageNumber="311">T. sciadosa</emphasis>
<bibRefCitation id="EFE14B0AD946E048489EF8CF7CBDF849" author="Wu, C. S. &amp; Liu, Y. Q." box="[1015,1197,1915,1942]" pageId="6" pageNumber="311" pagination="155 - 173" refId="ref6443" refString="Wu, C. S. &amp; Liu, Y. Q. (1994) A study of the Chinese Torodora Meyrick, 1894 and descriptions of new species (Lepidoptera, Lecithoceridae). Sinozoologia, 11, 155 - 173." type="journal article" year="1994">Wu &amp; Liu, 1994</bibRefCitation>
</taxonomicName>
which was described from
<collectingCountry id="F367766BD946E0484BBCF82C790BF865" box="[213,283,1951,1977]" name="China" pageId="6" pageNumber="311">China</collectingCountry>
, but the abdominal sternite VIII gently emarginated, whereas in the latter, deeply emarginated; signum more elongated anteriorly, length about 1.5x of width; and the distributional pattern of spines.
</paragraph>
</subSubSection>
<caption id="DF0F6673D947E0494BFEF8E87CB0F84B" ID-DOI="http://doi.org/10.5281/zenodo.4407608" ID-Zenodo-Dep="4407608" httpUri="https://zenodo.org/record/4407608/files/figure.png" pageId="7" pageNumber="312" startId="7.[151,250,1883,1907]" targetBox="[160,1426,186,1842]" targetPageId="7">
<paragraph id="8BCF36FBD947E0494BFEF8E87CB0F84B" blockId="7.[151,1436,1883,1944]" pageId="7" pageNumber="312">
<emphasis id="B904EAE9D947E0494BFEF8E87903F8AF" bold="true" box="[151,275,1883,1907]" pageId="7" pageNumber="312">FIGURE 5.</emphasis>
<taxonomicName id="4C704D78D947E0494A73F8E87A32F8AF" authority="Park" authorityName="Park &amp; Bae" authorityYear="2020" box="[282,546,1883,1907]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="312" phylum="Arthropoda" rank="species" species="cordisigna" status="sp. nov.">
<emphasis id="B904EAE9D947E0494A73F8E879FAF8AF" box="[282,490,1883,1907]" italics="true" pageId="7" pageNumber="312">Torodora cordisigna</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="A2375792D947E0494945F8E87A90F8AF" box="[556,640,1883,1907]" pageId="7" pageNumber="312" rank="species">
<emphasis id="B904EAE9D947E0494945F8E87A68F8AF" bold="true" box="[556,632,1883,1907]" pageId="7" pageNumber="312">sp. nov</emphasis>
.
</taxonomicNameLabel>
: A, holotype; B, labial palpus in lateral view; C, hind tibia; D, forewing venation; E, female genitalia, gen. slide no. CIS-8077, holotype, with close-up signum. Scale bar: 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C36A6570D948E0464BAEFF257968FC5C" pageId="8" pageNumber="313" type="description">
<paragraph id="8BCF36FBD948E0464BAEFF257B2CFD74" blockId="8.[151,1437,150,1004]" pageId="8" pageNumber="313">
<emphasis id="B904EAE9D948E0464BAEFF257949FF6C" bold="true" box="[199,345,150,176]" pageId="8" pageNumber="313">Description.</emphasis>
Female (
<figureCitation id="134B2A7ED948E0464AAEFF257A06FF6D" box="[455,534,150,177]" captionStart="FIGURE 5" captionStartId="7.[151,250,1883,1907]" captionTargetBox="[160,1426,186,1842]" captionTargetId="figure-19@7.[151,1436,181,1859]" captionTargetPageId="7" captionText="FIGURE 5. Torodora cordisigna Park, sp. nov.: A, holotype; B, labial palpus in lateral view; C, hind tibia; D, forewing venation; E, female genitalia, gen. slide no. CIS-8077, holotype, with close-up signum. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407608" httpUri="https://zenodo.org/record/4407608/files/figure.png" pageId="8" pageNumber="313">Figs. 5</figureCitation>
A-C). Wingspan
<quantity id="4C889B1ED948E04649B2FF257B55FF6D" box="[731,837,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="8" pageNumber="313" unit="mm" value="13.5">13.5 mm</quantity>
.
<emphasis id="B904EAE9D948E046483BFF257B9FFF6C" box="[850,911,150,176]" italics="true" pageId="8" pageNumber="313">Head</emphasis>
: Pale yellowish brown dorsally, with yellowish-white scales laterally. Antenna slightly longer than forewing; basal segment dilated distally, yellowish brown on dorsal surface, yellowish white on anterior and posterior surface; flagellum yellowish white, with dark-brown annulations. Second segment of labial palpus thickened, arched, yellowish brown in basal 3/4 and yellowish white in distal 1/4 with narrow yellowish-brown band on outer surface; yellowish white, rarely speckled with brownish scales medially on inner surface; 3
<superScript id="7C059BB3D948E0464938FEF97A4FFE84" attach="left" box="[593,607,330,344]" fontSize="6" pageId="8" pageNumber="313">rd</superScript>
segment, slender, longer than 2
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segment, yellowish white, dark brown ventrally.
<emphasis id="B904EAE9D948E0464A6EFEDD7971FE54" box="[263,353,366,392]" italics="true" pageId="8" pageNumber="313">Thorax:</emphasis>
Tegula and thorax yellowish white dorsally. Forewing ground color yellowish brown. Hind tibia with yellowish-white, rough scales above and creamy-white scales at apex. Forewing ground color yellowish brown evenly, with a small, yellowish-white costal patch at 3/4 of costa; apex obtuse; termen concave beyond apex; fringe concolorous with ground color, with narrow yellowish-white basal line and yellowish white apically; venation (
<figureCitation id="134B2A7ED948E0464E1CFE6978AAFDC5" captionStart="FIGURE 5" captionStartId="7.[151,250,1883,1907]" captionTargetBox="[160,1426,186,1842]" captionTargetId="figure-19@7.[151,1436,181,1859]" captionTargetPageId="7" captionText="FIGURE 5. Torodora cordisigna Park, sp. nov.: A, holotype; B, labial palpus in lateral view; C, hind tibia; D, forewing venation; E, female genitalia, gen. slide no. CIS-8077, holotype, with close-up signum. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407608" httpUri="https://zenodo.org/record/4407608/files/figure.png" pageId="8" pageNumber="313">Fig 5D</figureCitation>
) with R
<subScript id="17F434BED948E0464A7DFDA2790CFDC3" attach="left" box="[276,284,529,543]" fontSize="6" pageId="8" pageNumber="313">1</subScript>
arising from before middle; distance between origins of R
<subScript id="17F434BED948E04648CEFDA27BBFFDC3" attach="left" box="[935,943,529,543]" fontSize="6" pageId="8" pageNumber="313">2</subScript>
and R
<subScript id="17F434BED948E046489EFDA27BEFFDC3" attach="left" box="[1015,1023,529,543]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
about 1/2 the length of R
<subScript id="17F434BED948E0464E72FDA27D33FDC3" attach="left" box="[1307,1315,529,543]" fontSize="6" pageId="8" pageNumber="313">1</subScript>
and R
<subScript id="17F434BED948E0464E02FDA27D63FDC3" attach="left" box="[1387,1395,529,543]" fontSize="6" pageId="8" pageNumber="313">2</subScript>
; R
<subScript id="17F434BED948E0464EFDFDA27D8CFDC3" attach="left" box="[1428,1436,529,543]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
and R
<subScript id="17F434BED948E0464BB0FD8678F1FD9F" attach="left" box="[217,225,565,579]" fontSize="6" pageId="8" pageNumber="313">4</subScript>
stalked for basal 2/3; R
<subScript id="17F434BED948E0464A8EFD8779FFFD9E" attach="left" box="[487,495,564,578]" fontSize="6" pageId="8" pageNumber="313">5</subScript>
separated from just beyond stalk of R
<subScript id="17F434BED948E04648FEFD867B8FFD9F" attach="left" box="[919,927,565,579]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
and R
<subScript id="17F434BED948E046488EFD867BFFFD9F" attach="left" box="[999,1007,565,579]" fontSize="6" pageId="8" pageNumber="313">4</subScript>
; R
<subScript id="17F434BED948E0464F79FD877C08FD9E" attach="left" box="[1040,1048,564,578]" fontSize="6" pageId="8" pageNumber="313">5</subScript>
to termen; M
<subScript id="17F434BED948E0464FD9FD867CA8FD9F" attach="left" box="[1200,1208,565,579]" fontSize="6" pageId="8" pageNumber="313">2</subScript>
much nearer to M
<subScript id="17F434BED948E0464EECFD867D9DFD9F" attach="left" box="[1413,1421,565,579]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
;, nearly parallel to M
<subScript id="17F434BED948E0464A1BFDEA796AFDBB" attach="left" box="[370,378,601,615]" fontSize="6" pageId="8" pageNumber="313">1</subScript>
; CuA
<subScript id="17F434BED948E0464AD4FDEA79D5FDBB" attach="both" box="[445,453,601,615]" fontSize="6" pageId="8" pageNumber="313">1</subScript>
and CuA
<subScript id="17F434BED948E0464941FDEA7A20FDBB" attach="left" box="[552,560,601,615]" fontSize="6" pageId="8" pageNumber="313">2</subScript>
long-stalked, for basal 2/5; cell weakly closed. Hind wing paler than forewing, as wide as forewing; venation with M
<subScript id="17F434BED948E0464920FDCE7A41FD57" attach="left" box="[585,593,637,651]" fontSize="6" pageId="8" pageNumber="313">2</subScript>
close to M
<subScript id="17F434BED948E04649BDFDCE7ACCFD57" attach="left" box="[724,732,637,651]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
at base; M
<subScript id="17F434BED948E0464834FDCE7B75FD57" attach="left" box="[861,869,637,651]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
and CuA
<subScript id="17F434BED948E04648BCFDCE7BCDFD57" attach="left" box="[981,989,637,651]" fontSize="6" pageId="8" pageNumber="313">1</subScript>
stalked for basal 1/3; cell closed; apex acute; termen sinuate; fringe concolorous with ground color.
</paragraph>
<paragraph id="8BCF36FBD948E0464BAEFD0179C6FC80" blockId="8.[151,1437,150,1004]" pageId="8" pageNumber="313">
<emphasis id="B904EAE9D948E0464BAEFD017999FD10" box="[199,393,690,716]" italics="true" pageId="8" pageNumber="313">Female genitalia</emphasis>
(
<figureCitation id="134B2A7ED948E0464AF3FD0179E4FD11" box="[410,500,690,717]" captionStart="FIGURE 5" captionStartId="7.[151,250,1883,1907]" captionTargetBox="[160,1426,186,1842]" captionTargetId="figure-19@7.[151,1436,181,1859]" captionTargetPageId="7" captionText="FIGURE 5. Torodora cordisigna Park, sp. nov.: A, holotype; B, labial palpus in lateral view; C, hind tibia; D, forewing venation; E, female genitalia, gen. slide no. CIS-8077, holotype, with close-up signum. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407608" httpUri="https://zenodo.org/record/4407608/files/figure.png" pageId="8" pageNumber="313">Fig. 5E</figureCitation>
): Apophyses anteriores about 2/5 the length of apophyses posteriores. Abdominal sternite VIII concave on posterior margin medially, with long setae. Antrum membranous, funnel-shaped. Ductus bursae long, about 1.5x longer than corpus bursae; ductus seminalis narrow, arising from near posterior end of ductus bursae. Corpus bursae ovate; signum large, semiovate, with strong, conic spines in posterior 1/3 and minute ones anteriorly and laterally.
</paragraph>
<paragraph id="8BCF36FBD948E0464BAEFCD57968FC5C" blockId="8.[151,1437,150,1004]" box="[199,376,870,896]" pageId="8" pageNumber="313">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="C36A6570D948E0464BAEFC397A5DFC79" box="[199,589,906,933]" pageId="8" pageNumber="313" type="distribution">
<paragraph id="8BCF36FBD948E0464BAEFC397A5DFC79" blockId="8.[151,1437,150,1004]" box="[199,589,906,933]" pageId="8" pageNumber="313">
<emphasis id="B904EAE9D948E0464BAEFC397971FC78" bold="true" box="[199,353,906,932]" pageId="8" pageNumber="313">Distribution.</emphasis>
<collectingCountry id="F367766BD948E0464A0EFC38798DFC79" box="[359,413,907,933]" name="Laos" pageId="8" pageNumber="313">Laos</collectingCountry>
(
<collectingRegion id="49B4F819D948E0464AC4FC397A51FC79" box="[429,577,906,933]" country="Laos" name="Bolikhamxai" pageId="8" pageNumber="313">Bolikhamsai</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D948E0464BAEFC1D798AFC30" pageId="8" pageNumber="313" type="etymology">
<paragraph id="8BCF36FBD948E0464BAEFC1D798AFC30" blockId="8.[151,1437,150,1004]" pageId="8" pageNumber="313">
<emphasis id="B904EAE9D948E0464BAEFC1D795EFC14" bold="true" box="[199,334,942,968]" pageId="8" pageNumber="313">Etymology.</emphasis>
The species name is derived from the Latin,
<emphasis id="B904EAE9D948E0464827FC1D7B98FC14" box="[846,904,942,968]" italics="true" pageId="8" pageNumber="313">cordi</emphasis>
(= heart), referring to the heart-shaped signum of the female genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>