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<document id="BC629995B8260E148C5CAC4C7D24A885" ID-CLB-Dataset="8531" ID-DOI="10.11646/zootaxa.4851.2.5" ID-GBIF-Dataset="7beefd33-c83a-4e05-82d1-7522b65c112c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4407593" ID-ZooBank="C0027037-9243-40E5-AAE1-22A9A272BD14" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609433360249" checkinUser="plazi" docAuthor="Park, Kyu-Tek &amp; Bae, Yang-Seop" docDate="2020" docId="03D987EDD941E04D4BFEF9097B66FBE9" docLanguage="en" docName="zootaxa.4851.2.5.pdf" docOrigin="Zootaxa 4851 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torodora barathrona Park 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="308" masterDocId="FFE0FF95D940E04E4B69FFB37810FFDC" masterDocTitle="Genus Torodora Meyrick (Lepidoptera, Lecithoceridae, Torodorinae) in Laos with descriptions of five new species and two newly recorded species" masterLastPageNumber="318" masterPageNumber="305" pageNumber="306" updateTime="1698850697182" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A175CAAEBF2782B8D53D61989138A07A">Genus Torodora Meyrick (Lepidoptera, Lecithoceridae, Torodorinae) in Laos with descriptions of five new species and two newly recorded species</mods:title>
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<mods:namePart id="897821B8D330D5830603253382F476C6">Park, Kyu-Tek</mods:namePart>
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<mods:affiliation id="366F1771160ADB531E543705B5389F73">BioResource and Environmental Center, Division of Life Sciences, Incheon National University, Incheon, 22012 Korea ktpark 02 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9933 - 4497</mods:affiliation>
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<mods:namePart id="8733A0DDEB62B16F896D623561E994B2">Bae, Yang-Seop</mods:namePart>
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<mods:affiliation id="35E2EC8DC06A264900E6737E4E1C2E37">BioResource and Environmental Center, Division of Life Sciences, Incheon National University, Incheon, 22012 Korea ktpark 02 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9933 - 4497 &amp; Division of Life Sciences, Incheon National University, Incheon, 22012 Korea</mods:affiliation>
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<treatment id="03D987EDD941E04D4BFEF9097B66FBE9" ID-DOI="http://doi.org/10.5281/zenodo.4407953" ID-GBIF-Taxon="177203859" ID-Zenodo-Dep="4407953" LSID="urn:lsid:plazi:treatment:03D987EDD941E04D4BFEF9097B66FBE9" httpUri="http://treatment.plazi.org/id/03D987EDD941E04D4BFEF9097B66FBE9" lastPageId="3" lastPageNumber="308" pageId="1" pageNumber="306">
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<paragraph id="8BCF36FBD941E04F4BFEF9097A58F909" blockId="1.[151,584,1722,1785]" box="[151,584,1722,1749]" pageId="1" pageNumber="306">
<heading id="D0878197D941E04F4BFEF9097A58F909" bold="true" box="[151,584,1722,1749]" fontSize="11" level="1" pageId="1" pageNumber="306" reason="1">
<emphasis id="B904EAE9D941E04F4BFEF9097A58F909" bold="true" box="[151,584,1722,1749]" pageId="1" pageNumber="306">
<taxonomicName id="4C704D78D941E04F4BFEF90979CEF909" ID-CoL="9FPX3" authority="Park" authorityName="Park" authorityYear="2020" box="[151,478,1722,1749]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="306" phylum="Arthropoda" rank="species" species="barathrona" status="sp. nov.">
<emphasis id="B904EAE9D941E04F4BFEF9097987F909" bold="true" box="[151,407,1722,1749]" italics="true" pageId="1" pageNumber="306">Torodora barathrona</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="A2375792D941E04F4A82F9087A58F909" box="[491,584,1723,1749]" pageId="1" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C36A6570D941E04F4BFEF96C7934F925" box="[151,292,1759,1785]" pageId="1" pageNumber="306" type="description">
<paragraph id="8BCF36FBD941E04F4BFEF96C7934F925" blockId="1.[151,584,1722,1785]" box="[151,292,1759,1785]" pageId="1" pageNumber="306">
(
<figureCitation id="134B2A7ED941E04F4BF6F96C78FCF925" box="[159,236,1759,1785]" captionStart="FIGURE 2" captionStartId="2.[151,250,1876,1900]" captionTargetBox="[165,1424,321,1834]" captionTargetId="figure-76@2.[151,1436,309,1853]" captionTargetPageId="2" captionText="FIGURE 2. Torodora barathrona Park, sp. nov.: A, holotype; B, paratype in lateral view; C, ditto, head with antenna and labial palpus; D, male genitalia, gen. slide no. CIS-8103, holotype; D, ditto, aedeagus; E; ditto, abdomen. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407600" httpUri="https://zenodo.org/record/4407600/files/figure.png" pageId="1" pageNumber="306">Figs. 2</figureCitation>
A-F)
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D941E04F4BFEF89479B0F855" pageId="1" pageNumber="306" type="materials_examined">
<paragraph id="8BCF36FBD941E04F4BFEF89479B0F855" blockId="1.[151,1436,1830,2038]" pageId="1" pageNumber="306">
<emphasis id="B904EAE9D941E04F4BFEF894795EF89C" bold="true" box="[151,334,1830,1857]" pageId="1" pageNumber="306">Type material.</emphasis>
<materialsCitation id="3B183CA6D941E04F4A34F89479B0F855" ID-GBIF-Occurrence="3028414309" collectingDate="2019-08-02" collectorName="YS Bae &amp; Bae, MJ &amp; Qi, DJ &amp; Lee, TG &amp; Lee, YB &amp; Cha, JH &amp; CM Jang" country="Laos" elevation="470" latitude="18.45639" location="Phou Khao Khouay National Protected Area" longLatPrecision="21" longitude="103.05139" pageId="1" pageNumber="306" specimenCount="1" stateProvince="Bolikhamxai" typeStatus="holotype">
<typeStatus id="54CB8859D941E04F4A34F89479DAF89D" box="[349,458,1831,1857]" pageId="1" pageNumber="306" type="holotype">Holotype</typeStatus>
: male,
<collectingCountry id="F367766BD941E04F4943F8947A67F89D" box="[554,631,1831,1857]" name="Laos" pageId="1" pageNumber="306">LAOS</collectingCountry>
:
<collectingRegion id="49B4F819D941E04F49E2F8947B75F89D" box="[651,869,1831,1857]" country="Laos" name="Bolikhamxai" pageId="1" pageNumber="306">Bolikhamsai Prov.</collectingRegion>
,
<location id="8EAF6020D941E04F4810F8947D88F89D" LSID="urn:lsid:plazi:treatment:03D987EDD941E04D4BFEF9097B66FBE9:8EAF6020D941E04F4810F8947D88F89D" box="[889,1432,1831,1857]" country="Laos" latitude="18.45639" longLatPrecision="21" longitude="103.05139" name="Phou Khao Khouay National Protected Area" pageId="1" pageNumber="306" stateProvince="Bolikhamxai">Phou Khao Khouay National Protected Area</location>
,
<geoCoordinate id="EE44503CD941E04F4BFEF8F87935F8B9" box="[151,293,1867,1893]" degrees="18" direction="north" minutes="27" orientation="latitude" pageId="1" pageNumber="306" precision="15" seconds="23" value="18.45639">18°2723N</geoCoordinate>
<geoCoordinate id="EE44503CD941E04F4A42F8F879DBF8BA" box="[299,459,1867,1894]" degrees="103" direction="east" minutes="03" orientation="longitude" pageId="1" pageNumber="306" precision="15" seconds="05" value="103.05139">103° 0305E</geoCoordinate>
,
<quantity id="4C889B1ED941E04F4ABEF8F87A30F8B9" box="[471,544,1867,1893]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="306" unit="m" value="470.0">
<elevation id="005DD1C8D941E04F4ABEF8F87A30F8B9" box="[471,544,1867,1893]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="306" unit="m" value="470.0">470 m</elevation>
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,
<date id="FFCE103BD941E04F4942F8F87AB6F8B9" box="[555,678,1867,1893]" pageId="1" pageNumber="306" value="2019-08-02">
<collectingDate id="EF8AE9D3D941E04F4942F8F87AB6F8B9" box="[555,678,1867,1893]" pageId="1" pageNumber="306" value="2019-08-02">2 viii 2019</collectingDate>
</date>
, leg.
<collectorName id="2685532DD941E04F49B6F8F87B28F8B9" box="[735,824,1867,1893]" pageId="1" pageNumber="306">YS Bae</collectorName>
, MJ
<collectorName id="2685532DD941E04F4804F8F87CCCF8B9" box="[877,1244,1867,1893]" pageId="1" pageNumber="306">Qi, DJ Lee, TG Lee, YB Cha, JH</collectorName>
Ko, &amp;
<collectorName id="2685532DD941E04F4E46F8F87D88F8B9" box="[1327,1432,1867,1893]" pageId="1" pageNumber="306">CM Jang</collectorName>
; gen. side no. CIS-8103.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D941E04C4BAEF8217C4CFF25" lastPageId="2" lastPageNumber="307" pageId="1" pageNumber="306" type="diagnosis">
<paragraph id="8BCF36FBD941E04C4BAEF8217C4CFF25" blockId="1.[151,1436,1830,2038]" lastBlockId="2.[151,1436,151,249]" lastPageId="2" lastPageNumber="307" pageId="1" pageNumber="306">
<emphasis id="B904EAE9D941E04F4BAEF8217952F870" bold="true" box="[199,322,1938,1964]" pageId="1" pageNumber="306">Diagnosis.</emphasis>
This new species is one of the smallest species of the genus
<taxonomicName id="4C704D78D941E04F4891F8207CDAF871" authority="Meyrick" authorityName="Meyrick" authorityYear="1894" box="[1016,1226,1939,1965]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B904EAE9D941E04F4891F8207C70F871" box="[1016,1120,1939,1965]" italics="true" pageId="1" pageNumber="306">Torodora</emphasis>
Meyrick
</taxonomicName>
, and similar to the following new species,
<taxonomicName id="4C704D78D941E04F4AC9F80B7A1DF80D" authority="Park &amp; Bae, 2020" authorityName="Park &amp; Bae" authorityYear="2020" box="[416,525,1975,2001]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="306" phylum="Arthropoda" rank="species" species="canalis" status="sp. nov.">
<emphasis id="B904EAE9D941E04F4AC9F80B7A1DF80D" box="[416,525,1975,2001]" italics="true" pageId="1" pageNumber="306">T. canalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2375792D941E04F497DF8047A7DF80D" box="[532,621,1975,2001]" pageId="1" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
, but it can be distinguished from the latter and its allies by the following diagnostic characters: i) the antenna with distinct dark-brown annulations on flagellum; ii) the forewing with a large, blackish fascia before middle and an elongated trapezoidal patch arising from 2/3 of costa extended to the lower margin of discal cell; iii) the forewing termen deeply concave beyond the apex; and iv) the valva of male genitalia having a short process medially followed by deep emargination on the ventral margin.
</paragraph>
</subSubSection>
<caption id="DF0F6673D942E04C4BFEF8E77946F868" ID-DOI="http://doi.org/10.5281/zenodo.4407600" ID-Zenodo-Dep="4407600" httpUri="https://zenodo.org/record/4407600/files/figure.png" pageId="2" pageNumber="307" startId="2.[151,250,1876,1900]" targetBox="[165,1424,321,1834]" targetPageId="2">
<paragraph id="8BCF36FBD942E04C4BFEF8E77946F868" blockId="2.[151,1437,1876,1973]" pageId="2" pageNumber="307">
<emphasis id="B904EAE9D942E04C4BFEF8E77902F8B0" bold="true" box="[151,274,1876,1900]" pageId="2" pageNumber="307">FIGURE 2.</emphasis>
<taxonomicName id="4C704D78D942E04C4A71F8E77A35F8B0" authority="Park" authorityName="Park" authorityYear="2020" box="[280,549,1876,1900]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="307" phylum="Arthropoda" rank="species" species="barathrona" status="sp. nov.">
<emphasis id="B904EAE9D942E04C4A71F8E779FEF8B0" box="[280,494,1876,1900]" italics="true" pageId="2" pageNumber="307">Torodora barathrona</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="A2375792D942E04C4947F8E67A91F8B0" box="[558,641,1877,1901]" pageId="2" pageNumber="307" rank="species">
<emphasis id="B904EAE9D942E04C4947F8E67A69F8B1" bold="true" box="[558,633,1877,1901]" pageId="2" pageNumber="307">sp. nov</emphasis>
.
</taxonomicNameLabel>
: A, holotype; B, paratype in lateral view; C, ditto, head with antenna and labial palpus; D, male genitalia, gen. slide no. CIS-8103, holotype; D, ditto, aedeagus; E; ditto, abdomen. Scale bar for genitalia and aedeagus: 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C36A6570D943E04D4BAEFF257981FC15" pageId="3" pageNumber="308" type="description">
<paragraph id="8BCF36FBD943E04D4BAEFF257C55FDC5" blockId="3.[151,1437,150,1077]" pageId="3" pageNumber="308">
<emphasis id="B904EAE9D943E04D4BAEFF257941FF6C" bold="true" box="[199,337,150,176]" pageId="3" pageNumber="308">Description</emphasis>
. Male (
<figureCitation id="134B2A7ED943E04D4AC1FF2479FBFF6D" box="[424,491,151,177]" captionStart="FIGURE 2" captionStartId="2.[151,250,1876,1900]" captionTargetBox="[165,1424,321,1834]" captionTargetId="figure-76@2.[151,1436,309,1853]" captionTargetPageId="2" captionText="FIGURE 2. Torodora barathrona Park, sp. nov.: A, holotype; B, paratype in lateral view; C, ditto, head with antenna and labial palpus; D, male genitalia, gen. slide no. CIS-8103, holotype; D, ditto, aedeagus; E; ditto, abdomen. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407600" httpUri="https://zenodo.org/record/4407600/files/figure.png" pageId="3" pageNumber="308">Fig. 2</figureCitation>
A-C). Wingspan 11.0 mm.
<emphasis id="B904EAE9D943E04D4873FF247B47FF6D" box="[794,855,151,177]" italics="true" pageId="3" pageNumber="308">Head</emphasis>
: Covered with shiny, orange-gray or brownish-gray scales dorsally. Basal segment of antenna elongated, dark brown on dorsal and ventral surfaces, creamy-white on anterior and posterior surfaces; flagellum creamy white, with distinct, dark-brown annulations (
<figureCitation id="134B2A7ED943E04D4FB3FF6C7D24FF25" box="[1242,1332,223,249]" captionStart="FIGURE 2" captionStartId="2.[151,250,1876,1900]" captionTargetBox="[165,1424,321,1834]" captionTargetId="figure-76@2.[151,1436,309,1853]" captionTargetPageId="2" captionText="FIGURE 2. Torodora barathrona Park, sp. nov.: A, holotype; B, paratype in lateral view; C, ditto, head with antenna and labial palpus; D, male genitalia, gen. slide no. CIS-8103, holotype; D, ditto, aedeagus; E; ditto, abdomen. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407600" httpUri="https://zenodo.org/record/4407600/files/figure.png" pageId="3" pageNumber="308">Fig. 2C</figureCitation>
). Second segment of labial palpus thickened, brownish gray on outer surface, with white ring at apex; 3rd segment strongly upturned, as long as 2nd segment, blackish ventrally.
<emphasis id="B904EAE9D943E04D4998FE947B52FE9D" box="[753,834,295,321]" italics="true" pageId="3" pageNumber="308">Thorax</emphasis>
: Tegula and thorax brownish gray dorsally. Forewing ground color brownish gray, scattered with dark-brown scales; a large, irregular-shaped, blackish fascia well-developed before middle, triangularly emarginated medially on outer margin and an elongated trapezoidal patch arising from 2/3 of costa extended to the lower margin of discal cell, surrounded by white line; postmedian white line more or less zigzagged, arising from 4/5 of costa; apex sharply produced; termen strongly concave beyond apex, with black scales along margin; fringe brownish orange. Hind wing pale brownish gray,
<emphasis id="B904EAE9D943E04D4F31FE687CD2FE29" box="[1112,1218,475,501]" italics="true" pageId="3" pageNumber="308">Abdomen</emphasis>
(
<figureCitation id="134B2A7ED943E04D4FBAFE687D3CFE29" box="[1235,1324,475,501]" captionStart="FIGURE 2" captionStartId="2.[151,250,1876,1900]" captionTargetBox="[165,1424,321,1834]" captionTargetId="figure-76@2.[151,1436,309,1853]" captionTargetPageId="2" captionText="FIGURE 2. Torodora barathrona Park, sp. nov.: A, holotype; B, paratype in lateral view; C, ditto, head with antenna and labial palpus; D, male genitalia, gen. slide no. CIS-8103, holotype; D, ditto, aedeagus; E; ditto, abdomen. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407600" httpUri="https://zenodo.org/record/4407600/files/figure.png" pageId="3" pageNumber="308">Fig. 2F</figureCitation>
): Spinous zones broadly developed; sternite VIII broad, with slightly sinuate posterior margin.
</paragraph>
<paragraph id="8BCF36FBD943E04D4BAEFD907D16FC79" blockId="3.[151,1437,150,1077]" pageId="3" pageNumber="308">
<emphasis id="B904EAE9D943E04D4BAEFD90797DFDE1" box="[199,365,547,573]" italics="true" pageId="3" pageNumber="308">Male genitalia</emphasis>
(
<figureCitation id="134B2A7ED943E04D4A15FD907A13FDE1" box="[380,515,547,573]" captionStart="FIGURE 2" captionStartId="2.[151,250,1876,1900]" captionTargetBox="[165,1424,321,1834]" captionTargetId="figure-76@2.[151,1436,309,1853]" captionTargetPageId="2" captionText="FIGURE 2. Torodora barathrona Park, sp. nov.: A, holotype; B, paratype in lateral view; C, ditto, head with antenna and labial palpus; D, male genitalia, gen. slide no. CIS-8103, holotype; D, ditto, aedeagus; E; ditto, abdomen. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407600" httpUri="https://zenodo.org/record/4407600/files/figure.png" pageId="3" pageNumber="308">Figs. 2D, E</figureCitation>
): Uncus rather short, with rounded apex. Median process of gnathos rather small, strongly bent beyond basal 2/3. Tegumen weakly sclerotized, deeply concave on anterior margin. Valva broad at base, basal part nearly quadrate; costa sclerotized, nearly straight in basal 1/5, then abruptly oblique, nearly with right angle, deeply concave before middle; ventral margin with short process beyond middle followed by deep emargination; cucullus thumb-like, with rounded apex, densely setose; sacculus narrow, extended to lower corner of cucullus. Juxta rectangular, concave on caudal margin, with a small, triangular median plate; lateral margin slightly concave medially. Vinculum narrow, banded; saccal zone rounded. Aedeagus stout, broader than basal part of valva, shorter than valva, slightly narrowed distally, bifurcated apically: dorsal process broadened distally with truncate caudal margin; ventral process bar-shaped, with two small triangular spines medially; cornuti consisting of a long heavily sclerotized stripe extended from near basal 1/4 to near apex, with a bird-shaped heavily sclerotized distal plate, and with a long, large sac from near base to near 2/3, bearing numerous conic spines internally.
</paragraph>
<paragraph id="8BCF36FBD943E04D4BAEFC1C7981FC15" blockId="3.[151,1437,150,1077]" box="[199,401,943,969]" pageId="3" pageNumber="308">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C36A6570D943E04D4BAEFC617A5DFC32" box="[199,589,978,1006]" pageId="3" pageNumber="308" type="distribution">
<paragraph id="8BCF36FBD943E04D4BAEFC617A5DFC32" blockId="3.[151,1437,150,1077]" box="[199,589,978,1006]" pageId="3" pageNumber="308">
<emphasis id="B904EAE9D943E04D4BAEFC617971FC30" bold="true" box="[199,353,978,1004]" pageId="3" pageNumber="308">Distribution.</emphasis>
<collectingCountry id="F367766BD943E04D4A0EFC60798DFC31" box="[359,413,979,1005]" name="Laos" pageId="3" pageNumber="308">Laos</collectingCountry>
(
<collectingRegion id="49B4F819D943E04D4AC4FC607A51FC32" box="[429,577,979,1006]" country="Laos" name="Bolikhamxai" pageId="3" pageNumber="308">Bolikhamsai</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D943E04D4BAEFC447B66FBE9" pageId="3" pageNumber="308" type="etymology">
<paragraph id="8BCF36FBD943E04D4BAEFC447B66FBE9" blockId="3.[151,1437,150,1077]" pageId="3" pageNumber="308">
<emphasis id="B904EAE9D943E04D4BAEFC44795EFBCD" bold="true" box="[199,334,1015,1041]" pageId="3" pageNumber="308">Etymology.</emphasis>
The species name is derived from the Latin,
<emphasis id="B904EAE9D943E04D4808FC447BC9FBCD" box="[865,985,1015,1041]" italics="true" pageId="3" pageNumber="308">barathrum</emphasis>
(= gulf, pit), referring to the hollowed emargination on the ventral margin of valva in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>