treatments-xml/data/03/D9/87/03D9879BE84C177E7EFDFC4D6949A567.xml
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<document id="C3516D29AFDD2F85734B776639B44CF7" ID-CLB-Dataset="99517" ID-DOI="10.11646/zootaxa.5165.4.1" ID-GBIF-Dataset="cd84b880-d77f-401a-bf4c-a98021a5e166" ID-ISSN="1175-5326" ID-Zenodo-Dep="6853942" ID-ZooBank="DBE812FF-7B25-4522-97FE-45A86ECAEFD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658127043518" checkinUser="plazi" docAuthor="Yang, Qiao-Zhi &amp; Pathomwattananurak, Wuttipon" docDate="2022" docId="03D9879BE84C177E7EFDFC4D6949A567" docLanguage="en" docName="zootaxa.5165.4.1.pdf" docOrigin="Zootaxa 5165 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eophileurus minor Yang &amp; Pathomwattananurak 2022, new species" docType="treatment" docVersion="7" lastPageNumber="469" masterDocId="FFE0FFE3E85D176C7E6AFFEE6B79A22A" masterDocTitle="Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species" masterLastPageNumber="485" masterPageNumber="451" pageNumber="468" updateTime="1699360166553" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A7CE0F6600B11FADE9C8F09DA707819C">Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species</mods:title>
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<mods:namePart id="5FC4E95199E18B53AAA02C33912B74F0">Yang, Qiao-Zhi</mods:namePart>
<mods:affiliation id="AB2168A68FA5B6F9C8AA9173738D5B98">3 B, Pingguoshequ, Baiziwan Road, Chaoyang Dist., Beijing, 100022, P. R. China.</mods:affiliation>
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<mods:namePart id="64E796C4EC263CF057414BF161EF3BFA">Pathomwattananurak, Wuttipon</mods:namePart>
<mods:affiliation id="AB50203C46A1DE7ECD22F2C19107A484">1051 / 38 Wiang, Mueang Dist., Chiang Rai Province, 57000, Thailand.</mods:affiliation>
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<mods:date id="C188113AB0F1AAC72798A19989B171BA">2022</mods:date>
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<taxonomicName id="4C704D0EE84C177D7EFDFC4D6878A194" authority="Yang &amp; Pathomwattananurak" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" box="[151,769,931,958]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="B904EA9FE84C177D7EFDFC4D6A04A194" bold="true" box="[151,381,931,958]" italics="true" pageId="17" pageNumber="468">Eophileurus minor</emphasis>
Yang &amp; Pathomwattananurak
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(
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,
<figureCitation id="134B2A08E84C177D7F59FC266AEFA1C8" box="[307,406,968,994]" captionStart="FIGURES109116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109116.HabitusofEophileurusArrow,1908species(obliqueview).109110,E.minorYang&amp;Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111112, E. prelli Yang &amp; Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113114, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115116, E. sidereus Yang &amp; Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="17" pageNumber="468">109110</figureCitation>
,
<figureCitation id="134B2A08E84C177D7FC8FC26697EA1C8" box="[418,519,968,994]" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="17" pageNumber="468">169172</figureCitation>
,
<figureCitation id="134B2A08E84C177D7C78FC26690CA1C9" box="[530,629,968,995]" captionStart="FIGURES 185196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185196. Parameres of Eophileurus Arrow, 1908. 185192, E. minor Yang &amp; Pathomwattananurak, new species, 185187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193196, E. sidereus Yang &amp; Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="17" pageNumber="468">185192</figureCitation>
)
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leg.”
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;
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<collectingMunicipality id="6BABACF7E84C177D7A3AFB2B6FD5A6F5" box="[1104,1196,1221,1247]" pageId="17" pageNumber="468">Mueang</collectingMunicipality>
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<collectingRegion id="49B4F86FE84C177D7EB7FB066970A729" box="[221,521,1256,1283]" country="Thailand" name="Nakhon Ratchasima" pageId="17" pageNumber="468">Nakhon Ratchasima Prov.</collectingRegion>
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<collectingCountry id="F367761DE84C177D7C7CFB066905A728" box="[534,636,1256,1282]" name="Thailand" pageId="17" pageNumber="468">Thailand</collectingCountry>
, 2020,
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leg.”
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;
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<specimenCount id="9D76FD04E84C177D7A66FB066F56A728" box="[1036,1071,1256,1282]" pageId="17" pageNumber="468" type="male">1♂</specimenCount>
(
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), “
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<collectingRegion id="49B4F86FE84C177D7EFDFAE26A3FA70D" box="[151,326,1292,1319]" country="Thailand" name="Chon Buri" pageId="17" pageNumber="468">Chonburi Prov.</collectingRegion>
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<collectingCountry id="F367761DE84C177D7F38FAE26AC1A70C" box="[338,440,1292,1318]" name="Thailand" pageId="17" pageNumber="468">Thailand</collectingCountry>
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<collectorName id="2685535BE84C177D7FAEFAE269E6A70C" box="[452,671,1292,1318]" pageId="17" pageNumber="468">Pichaphop Junaime</collectorName>
leg., adult emerged in
<date id="FFCE104DE84C177D7DCFFAE26F69A70C" box="[933,1040,1292,1318]" pageId="17" pageNumber="468" value="2021-07">
<collectingDate id="EF8AE9A5E84C177D7DCFFAE26F69A70C" box="[933,1040,1292,1318]" pageId="17" pageNumber="468" value="2021-07">VII.2021</collectingDate>
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(
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), “
<location id="8EAF6056E84C177D7AD1FAE26EE1A70C" LSID="urn:lsid:plazi:treatment:03D9879BE84C177E7EFDFC4D6949A567:8EAF6056E84C177D7AD1FAE26EE1A70C" box="[1211,1432,1292,1318]" country="Thailand" municipality="Nong Khae Dist." name="Nong Nak Subdist." pageId="17" pageNumber="468" stateProvince="Sara Buri">Nong Nak Subdist.</location>
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<collectingRegion id="49B4F86FE84C177D7F0BFADE697DA761" box="[353,516,1328,1355]" country="Thailand" name="Sara Buri" pageId="17" pageNumber="468">Saraburi Prov.</collectingRegion>
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,
<date id="FFCE104DE84C177D7C15FADE6872A760" box="[639,779,1328,1354]" pageId="17" pageNumber="468" value="2021-07-22">
<collectingDate id="EF8AE9A5E84C177D7C15FADE6872A760" box="[639,779,1328,1354]" pageId="17" pageNumber="468" value="2021-07-22">22.VII.2021</collectingDate>
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,
<collectorName id="2685535BE84C177D7D7CFADE68BEA760" box="[790,967,1328,1354]" pageId="17" pageNumber="468">Bannatad Benja</collectorName>
leg.”
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;
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<specimenCount id="9D76FD04E84C177D7A78FADE6F34A760" box="[1042,1101,1328,1354]" pageId="17" pageNumber="468" type="male">3♂♂</specimenCount>
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(
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), “
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generation of female from
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<collectingCountry id="F367761DE84C177D7DC2FABA6F77A744" box="[936,1038,1364,1390]" name="Thailand" pageId="17" pageNumber="468">Thailand</collectingCountry>
,
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<collectingDate id="EF8AE9A5E84C177D7A70FABA6FDFA744" box="[1050,1190,1364,1390]" pageId="17" pageNumber="468" value="2021-07-22">22.VII.2021</collectingDate>
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,
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leg., adult emerged in
<date id="FFCE104DE84C177D7F34FA966AB9A7B8" box="[350,448,1400,1426]" pageId="17" pageNumber="468" value="2021-11">
<collectingDate id="EF8AE9A5E84C177D7F34FA966AB9A7B8" box="[350,448,1400,1426]" pageId="17" pageNumber="468" value="2021-11">XI.2021</collectingDate>
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(
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generation of female from
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<collectingMunicipality id="6BABACF7E84C177D7B70FA976BA9A79C" pageId="17" pageNumber="468">Nong Khae Dist.</collectingMunicipality>
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<collectingRegion id="49B4F86FE84C177D7EB1FA726A06A79D" box="[219,383,1436,1463]" country="Thailand" name="Sara Buri" pageId="17" pageNumber="468">Saraburi Prov.</collectingRegion>
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<collectingCountry id="F367761DE84C177D7FE3FA726A96A79C" box="[393,495,1436,1462]" name="Thailand" pageId="17" pageNumber="468">Thailand</collectingCountry>
,
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<collectingDate id="EF8AE9A5E84C177D7F91FA7269FEA79C" box="[507,647,1436,1462]" pageId="17" pageNumber="468" value="2021-07-22">22.VII.2021</collectingDate>
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,
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leg., adult emerged in
<date id="FFCE104DE84C177D7A22FA726FD3A79C" box="[1096,1194,1436,1462]" pageId="17" pageNumber="468" value="2021-11">
<collectingDate id="EF8AE9A5E84C177D7A22FA726FD3A79C" box="[1096,1194,1436,1462]" pageId="17" pageNumber="468" value="2021-11">XI.2021</collectingDate>
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(
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,
<collectingMunicipality id="6BABACF7E84C177D7F08FA2E6969A7F0" box="[354,528,1472,1498]" pageId="17" pageNumber="468">Selaphum Dist.</collectingMunicipality>
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<collectingCountry id="F367761DE84C177D7CC1FA2E6868A7F0" box="[683,785,1472,1498]" name="Thailand" pageId="17" pageNumber="468">Thailand</collectingCountry>
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<collectingDate id="EF8AE9A5E84C177D7D71FA2E68E7A7F0" box="[795,926,1472,1498]" pageId="17" pageNumber="468" value="2021-09-18">18.IX.2021</collectingDate>
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leg.”
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;
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(
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,
<collectingMunicipality id="6BABACF7E84C177D7FF5FA0B6916A7D4" box="[415,623,1508,1535]" pageId="17" pageNumber="468">Na Kluea Subdist.</collectingMunicipality>
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<collectingCountry id="F367761DE84C177D7A43FA0A6FF6A7D4" box="[1065,1167,1508,1534]" name="Thailand" pageId="17" pageNumber="468">Thailand</collectingCountry>
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</date>
,
<collectorName id="2685535BE84C177D7B18FA0B6B96A408" pageId="17" pageNumber="468">BoNing Li</collectorName>
leg.”
</materialsCitation>
;
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<specimenCount id="9D76FD04E84C177D7F53F9E66A25A408" box="[313,348,1544,1570]" pageId="17" pageNumber="468" type="male">1♂</specimenCount>
(
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,
<collectingMunicipality id="6BABACF7E84C177D7CB6F9E768B2A408" box="[732,971,1544,1571]" pageId="17" pageNumber="468">Ham Thuan Bac dist.</collectingMunicipality>
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<collectingCountry id="F367761DE84C177D7AC2F9E66E73A408" box="[1192,1290,1544,1570]" name="Vietnam" pageId="17" pageNumber="468">Vietnam</collectingCountry>
,
<date id="FFCE104DE84C177D7B7EF9E66E1EA408" box="[1300,1383,1544,1570]" pageId="17" pageNumber="468" value="2020-05">
<collectingDate id="EF8AE9A5E84C177D7B7EF9E66E1EA408" box="[1300,1383,1544,1570]" pageId="17" pageNumber="468" value="2020-05">
<collectionCode id="ED61AE48E84C177D7B7EF9E66E51A408" box="[1300,1320,1544,1570]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="17" pageNumber="468" type="Museum">V</collectionCode>
.2020
</collectingDate>
</date>
,
<collectorName id="2685535BE84C177D7B1BF9E76BADA46D" pageId="17" pageNumber="468">Van Dang</collectorName>
leg.”
</materialsCitation>
</paragraph>
<paragraph id="8BCF368DE84C177E7EFDF9BE6F60A017" blockId="17.[151,1437,1040,2039]" lastBlockId="18.[151,1437,150,1869]" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468">
<emphasis id="B904EA9FE84C177D7EFDF9BE6A7DA440" bold="true" box="[151,260,1616,1642]" pageId="17" pageNumber="468">
<typeStatus id="54CB882FE84C177D7EFDF9BE6A7DA440" box="[151,260,1616,1642]" pageId="17" pageNumber="468" type="holotype">Holotype</typeStatus>
</emphasis>
(male).
<emphasis id="B904EA9FE84C177D7F09F9BE6ABAA440" bold="true" box="[355,451,1616,1642]" pageId="17" pageNumber="468">General</emphasis>
(
<figureCitation id="134B2A08E84C177D7FBEF9BE6925A440" box="[468,604,1616,1642]" captionStart="FIGURES 8792" captionStartId="19.[151,264,1518,1542]" captionTargetBox="[155,1430,185,1490]" captionTargetId="figure-18@19.[151,1435,181,1494]" captionTargetPageId="19" captionText="FIGURES 8792. Habitus of Eophileurus minorYang &amp; Pathomwattananurak, new species. 8788, holotype; 89, male paratype (Mueang Nakhon Ratchasima); 9091, female paratype (Bua Yai); 92, male paratype (Dong Tien, image ©Ming-Zhi Zhao); 87, 89, 90, 92 dorsal view; 88, 91, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853970" httpUri="https://zenodo.org/record/6853970/files/figure.png" pageId="17" pageNumber="468">Figs. 8788</figureCitation>
,
<figureCitation id="134B2A08E84C177D7C02F9BE69EAA440" box="[616,659,1616,1642]" captionStart="FIGURES109116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109116.HabitusofEophileurusArrow,1908species(obliqueview).109110,E.minorYang&amp;Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111112, E. prelli Yang &amp; Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113114, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115116, E. sidereus Yang &amp; Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="17" pageNumber="468">109</figureCitation>
): Body black with very slight reddish color, rather oblong, flattened, surface shiny; ventral surface clads with very dense, long reddish yellow setae.
<emphasis id="B904EA9FE84C177D7A42F99A6F08A4A4" bold="true" box="[1064,1137,1652,1678]" pageId="17" pageNumber="468">Head:</emphasis>
Clypeus densely punctate with rugopunctures, with a very short, slightly backward, conical horn at middle, an indistinct ridge from horn to apex and a more distinct ridge behind the horn; clypeal apex rounded. Frons slightly depressed, disk with dense, small punctures, becoming denser and larger near sides. Mandible slightly sinuate at outer margin.
<emphasis id="B904EA9FE84C177D7B70F90E6EE5A4D0" bold="true" box="[1306,1436,1760,1786]" pageId="17" pageNumber="468">Pronotum:</emphasis>
Surface with dense, small punctures, becoming denser and larger near all margins. Anterior region with a small, very shallow, round fovea; punctures in fovea dense and large. All margins moderately beaded except before fovea, more strongly beaded at sides. Anterior angle acute, apex slightly rounded, posterior angle obtusely angulate.
<emphasis id="B904EA9FE84C177D7B71F8A26EE5A54C" bold="true" box="[1307,1436,1868,1894]" pageId="17" pageNumber="468">Scutellum:</emphasis>
Surface almost impunctate, with only a few small punctures at center.
<emphasis id="B904EA9FE84C177D7DC7F89E6F7AA5A0" bold="true" box="[941,1027,1904,1930]" pageId="17" pageNumber="468">Elytra:</emphasis>
Surface with dense micropunctures, each with scale-liked seta, and dense and large, sub-U-shape or round punctures. Interstice 1 with punctures in two to three irregular rows combined at middle to one irregular row, interstice 2 with punctures in one irregular row. Primary costa A and primary costa B indistinct. Subapical umbones slightly prominent. Margins of elytra parallel, strongly and horizontally dilated from base to posterior two-fifth.
<emphasis id="B904EA9FE84F177E7DEAFF786885A29A" bold="true" box="[896,1020,150,176]" pageId="18" pageNumber="469">Pygidium:</emphasis>
convex, with dense, small punctures throughout, punctures denser and smaller near apical margin.
<emphasis id="B904EA9FE84F177E7D0AFF556F75A2FF" bold="true" box="[864,1036,187,213]" pageId="18" pageNumber="469">Metasternum:</emphasis>
Surface mainly dark reddish, with darker color at posterior region; region near sides with dense and large punctures; central portion with only sparse and small punctures; setae reddish yellow, long, near all margins except posterior margin.
<emphasis id="B904EA9FE84F177E7AF6FEEC6EE5A336" bold="true" box="[1180,1436,258,284]" pageId="18" pageNumber="469">Abdominal ventrites:</emphasis>
Surfaces with sparse micropunctures throughout and larger punctures at each side except on ventrite 6; ventrite 6 with large punctures throughout and a distinct, horizontally oval depression at center.
<emphasis id="B904EA9FE84F177E7A17FEA56FC7A34F" bold="true" box="[1149,1214,331,357]" pageId="18" pageNumber="469">Legs:</emphasis>
Protibia tridentate, protarsi strongly thickened, inner protibial claw distinctly enlarged and elongated. Inner metatibial spur very long, strongly curved outward near apex; outer metatibial spur shorter, slightly curved.
<emphasis id="B904EA9FE84F177E7A3BFE7D6FABA387" bold="true" box="[1105,1234,403,429]" pageId="18" pageNumber="469">Parameres</emphasis>
(
<figureCitation id="134B2A08E84F177E7A8FFE7D6EF2A387" box="[1253,1419,403,429]" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="18" pageNumber="469">Figs. 169170</figureCitation>
): In frontal view (
<figureCitation id="134B2A08E84F177E7F3FFE596AC0A3FB" box="[341,441,439,465]" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="18" pageNumber="469">Fig. 169</figureCitation>
), inner margins overlapping from middle to near apex; outer sides strongly prominent outward at basal and middle regions; with a pair of wide processes close to middle extending almost to apex, apex of each process extending upward and curving inward, processes at asymmetric positions; apex extending outward; in lateral view (
<figureCitation id="134B2A08E84F177E7F2DFDCD6AD1A017" box="[327,424,547,573]" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="18" pageNumber="469">Fig. 170</figureCitation>
), processes near middle trapezoid; apex bent backward.
</paragraph>
<paragraph id="8BCF368DE84F177E7EADFDA9688EA18F" blockId="18.[151,1437,150,1869]" pageId="18" pageNumber="469">
<emphasis id="B904EA9FE84F177E7EADFDA96A46A04B" bold="true" box="[199,319,583,609]" pageId="18" pageNumber="469">
<typeStatus id="54CB882FE84F177E7EADFDA96A46A04B" box="[199,319,583,609]" pageId="18" pageNumber="469" type="paratype">Paratypes</typeStatus>
</emphasis>
(male). Characters mainly stable, for
<typeStatus id="54CB882FE84F177E7C9CFDA96818A04B" box="[758,865,583,609]" pageId="18" pageNumber="469" type="paratype">paratypes</typeStatus>
from
<collectingCountry id="F367761DE84F177E7DC0FDA96F77A04B" box="[938,1038,583,609]" name="Thailand" pageId="18" pageNumber="469">Thailand</collectingCountry>
(
<figureCitation id="134B2A08E84F177E7A75FDA96F0FA04B" box="[1055,1142,583,609]" captionStart="FIGURES 8792" captionStartId="19.[151,264,1518,1542]" captionTargetBox="[155,1430,185,1490]" captionTargetId="figure-18@19.[151,1435,181,1494]" captionTargetPageId="19" captionText="FIGURES 8792. Habitus of Eophileurus minorYang &amp; Pathomwattananurak, new species. 8788, holotype; 89, male paratype (Mueang Nakhon Ratchasima); 9091, female paratype (Bua Yai); 92, male paratype (Dong Tien, image ©Ming-Zhi Zhao); 87, 89, 90, 92 dorsal view; 88, 91, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853970" httpUri="https://zenodo.org/record/6853970/files/figure.png" pageId="18" pageNumber="469">Fig. 89</figureCitation>
): in general, compared to the
<typeStatus id="54CB882FE84F177E7EA9FD856A51A0AF" box="[195,296,619,645]" pageId="18" pageNumber="469" type="holotype">holotype</typeStatus>
, elytral margin very rarely much less dilated; parameres sometimes more slender, overlapping area very rarely much smaller than the
<typeStatus id="54CB882FE84F177E7C4EFD6169F0A083" box="[548,649,655,681]" pageId="18" pageNumber="469" type="holotype">holotype</typeStatus>
, processes sometimes thinner, rarely wider, very rarely at symmetric positions; for individuals more well-developed than the
<typeStatus id="54CB882FE84F177E7D6EFD5D6810A0E7" box="[772,873,691,717]" pageId="18" pageNumber="469" type="holotype">holotype</typeStatus>
, clypeus and frons more sparsely punctate; clypeal horn longer; all margins of pronotum beaded; fovea slightly larger and deeper, with a very weak protuberance at each side at posterior part, punctures within fovea much sparser; for individuals more underdeveloped than the
<typeStatus id="54CB882FE84F177E7EFDFCF16B85A113" box="[151,252,799,825]" pageId="18" pageNumber="469" type="holotype">holotype</typeStatus>
, clypeus and frons more densely rugopunctate; clypeal horn shorter; fovea absent (
<figureCitation id="134B2A08E84F177E7ADBFCF16BBCA177" captionStart="FIGURES 185196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185196. Parameres of Eophileurus Arrow, 1908. 185192, E. minor Yang &amp; Pathomwattananurak, new species, 185187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193196, E. sidereus Yang &amp; Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="18" pageNumber="469">Figs. 185187, 190 192</figureCitation>
); for the
<typeStatus id="54CB882FE84F177E7F42FCAD6AF1A177" box="[296,392,835,861]" pageId="18" pageNumber="469" type="paratype">paratype</typeStatus>
from
<collectingCountry id="F367761DE84F177E7FA1FCAD6953A177" box="[459,554,835,861]" name="Vietnam" pageId="18" pageNumber="469">Vietnam</collectingCountry>
(
<figureCitation id="134B2A08E84F177E7C53FCAD69F4A177" box="[569,653,835,861]" captionStart="FIGURES 8792" captionStartId="19.[151,264,1518,1542]" captionTargetBox="[155,1430,185,1490]" captionTargetId="figure-18@19.[151,1435,181,1494]" captionTargetPageId="19" captionText="FIGURES 8792. Habitus of Eophileurus minorYang &amp; Pathomwattananurak, new species. 8788, holotype; 89, male paratype (Mueang Nakhon Ratchasima); 9091, female paratype (Bua Yai); 92, male paratype (Dong Tien, image ©Ming-Zhi Zhao); 87, 89, 90, 92 dorsal view; 88, 91, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853970" httpUri="https://zenodo.org/record/6853970/files/figure.png" pageId="18" pageNumber="469">Fig. 92</figureCitation>
): slightly more well-developed than the
<typeStatus id="54CB882FE84F177E7A27FCAD6FCBA177" box="[1101,1202,835,861]" pageId="18" pageNumber="469" type="holotype">holotype</typeStatus>
, characters generally identical to well-developed males from
<collectingCountry id="F367761DE84F177E7C3BFC8969CEA1AB" box="[593,695,871,897]" name="Thailand" pageId="18" pageNumber="469">Thailand</collectingCountry>
, except elytral margin slightly less dilated; processes of parameres much thinner, not extending to apex, at symmetric positions (
<figureCitation id="134B2A08E84F177E7D2FFC656892A18F" box="[837,1003,907,933]" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="18" pageNumber="469">Figs. 171172</figureCitation>
).
</paragraph>
<paragraph id="8BCF368DE84F177E7EADFC416A15A673" blockId="18.[151,1437,150,1869]" pageId="18" pageNumber="469">
<emphasis id="B904EA9FE84F177E7EADFC416A46A1E3" bold="true" box="[199,319,943,969]" pageId="18" pageNumber="469">
<typeStatus id="54CB882FE84F177E7EADFC416A46A1E3" box="[199,319,943,969]" pageId="18" pageNumber="469" type="paratype">Paratypes</typeStatus>
</emphasis>
(female). Similar to male, with clypeus and frons much more densely rugopunctate throughout; clypeal horn much shorter; fovea absent; all margins of pronotum beaded; elytral margin slightly more strongly enlarged and extended further posteriorly; depression absent on last ventrite; protarsi not thickened, inner claw not enlarged and elongated; Inner metatibial relatively shorter, spur less strongly curved outward, sometimes straight (
<figureCitation id="134B2A08E84F177E7EF5FBD16A5EA673" box="[159,295,1087,1113]" captionStart="FIGURES 8792" captionStartId="19.[151,264,1518,1542]" captionTargetBox="[155,1430,185,1490]" captionTargetId="figure-18@19.[151,1435,181,1494]" captionTargetPageId="19" captionText="FIGURES 8792. Habitus of Eophileurus minorYang &amp; Pathomwattananurak, new species. 8788, holotype; 89, male paratype (Mueang Nakhon Ratchasima); 9091, female paratype (Bua Yai); 92, male paratype (Dong Tien, image ©Ming-Zhi Zhao); 87, 89, 90, 92 dorsal view; 88, 91, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853970" httpUri="https://zenodo.org/record/6853970/files/figure.png" pageId="18" pageNumber="469">Figs. 9091</figureCitation>
,
<figureCitation id="134B2A08E84F177E7F59FBD16A19A673" box="[307,352,1087,1113]" captionStart="FIGURES109116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109116.HabitusofEophileurusArrow,1908species(obliqueview).109110,E.minorYang&amp;Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111112, E. prelli Yang &amp; Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113114, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115116, E. sidereus Yang &amp; Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="18" pageNumber="469">110</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84F177E7EADFB8D6844A68B" pageId="18" pageNumber="469" type="description">
<paragraph id="8BCF368DE84F177E7EADFB8D6844A68B" blockId="18.[151,1437,150,1869]" pageId="18" pageNumber="469">
<emphasis id="B904EA9FE84F177E7EADFB8D6A07A657" bold="true" box="[199,382,1123,1149]" pageId="18" pageNumber="469">Measurements.</emphasis>
Body length: male
<quantity id="4C889B68E84F177E7C0EFB8C6873A657" box="[612,778,1122,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.25" metricValueMin="1.73" pageId="18" pageNumber="469" unit="mm" value="19.9" valueMax="22.5" valueMin="17.3">17.322.5 mm</quantity>
(
<typeStatus id="54CB882FE84F177E7D70FB8D6807A657" box="[794,894,1123,1149]" pageId="18" pageNumber="469" type="holotype">holotype</typeStatus>
<quantity id="4C889B68E84F177E7DEDFB8D688BA657" box="[903,1010,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="18" pageNumber="469" unit="mm" value="19.3">19.3 mm</quantity>
), female 18.122.0 mm; body width: male
<quantity id="4C889B68E84F177E7EBFFB696A10A68B" box="[213,361,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.0050000000000001" metricValueMax="1.1300000000000001" metricValueMin="0.8800000000000001" pageId="18" pageNumber="469" unit="mm" value="10.05" valueMax="11.3" valueMin="8.8">8.811.3 mm</quantity>
(
<typeStatus id="54CB882FE84F177E7F1DFB696AA2A68B" box="[375,475,1159,1185]" pageId="18" pageNumber="469" type="holotype">holotype</typeStatus>
<quantity id="4C889B68E84F177E7F8BFB696942A68B" box="[481,571,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="18" pageNumber="469" unit="mm" value="9.6">9.6 mm</quantity>
), female
<quantity id="4C889B68E84F177E7CC8FB696841A68B" box="[674,824,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" metricValueMax="1.1199999999999999" metricValueMin="0.9199999999999999" pageId="18" pageNumber="469" unit="mm" value="10.2" valueMax="11.2" valueMin="9.2">9.211.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84F177E7EADFB446A03A71B" pageId="18" pageNumber="469" type="diagnosis">
<paragraph id="8BCF368DE84F177E7EADFB446A03A71B" blockId="18.[151,1437,150,1869]" pageId="18" pageNumber="469">
<emphasis id="B904EA9FE84F177E7EADFB446A3BA6EE" bold="true" box="[199,322,1194,1220]" pageId="18" pageNumber="469">Diagnosis.</emphasis>
This species can be distinguished from other
<taxonomicName id="4C704D0EE84F177E7D1EFB456884A6EF" authorityName=", Endrodi" authorityYear="1977" box="[884,1021,1195,1221]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B904EA9FE84F177E7D1EFB456884A6EF" box="[884,1021,1195,1221]" italics="true" pageId="18" pageNumber="469">Eophileurus</emphasis>
</taxonomicName>
by its overall small size, strongly horizontally enlarged elytral margin which extends beyond the level of midpoint of elytron (
<figureCitation id="134B2A08E84F177E7AC5FB216EF6A6C3" box="[1199,1423,1231,1257]" captionStart="FIGURES 8792" captionStartId="19.[151,264,1518,1542]" captionTargetBox="[155,1430,185,1490]" captionTargetId="figure-18@19.[151,1435,181,1494]" captionTargetPageId="19" captionText="FIGURES 8792. Habitus of Eophileurus minorYang &amp; Pathomwattananurak, new species. 8788, holotype; 89, male paratype (Mueang Nakhon Ratchasima); 9091, female paratype (Bua Yai); 92, male paratype (Dong Tien, image ©Ming-Zhi Zhao); 87, 89, 90, 92 dorsal view; 88, 91, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853970" httpUri="https://zenodo.org/record/6853970/files/figure.png" pageId="18" pageNumber="469">Figs. 87, 8990, 92</figureCitation>
), significantly long and curved inner metatibial spur (
<figureCitation id="134B2A08E84F177E7C9EFB1D6807A727" box="[756,894,1267,1293]" captionStart="FIGURES 8792" captionStartId="19.[151,264,1518,1542]" captionTargetBox="[155,1430,185,1490]" captionTargetId="figure-18@19.[151,1435,181,1494]" captionTargetPageId="19" captionText="FIGURES 8792. Habitus of Eophileurus minorYang &amp; Pathomwattananurak, new species. 8788, holotype; 89, male paratype (Mueang Nakhon Ratchasima); 9091, female paratype (Bua Yai); 92, male paratype (Dong Tien, image ©Ming-Zhi Zhao); 87, 89, 90, 92 dorsal view; 88, 91, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853970" httpUri="https://zenodo.org/record/6853970/files/figure.png" pageId="18" pageNumber="469">Figs. 8792</figureCitation>
) and the unique shape of the parameres (
<figureCitation id="134B2A08E84F177E7B0EFB1D6B85A71B" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="18" pageNumber="469">Figs. 169172</figureCitation>
,
<figureCitation id="134B2A08E84F177E7F6DFAF86A17A71B" box="[263,366,1302,1329]" captionStart="FIGURES 185196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185196. Parameres of Eophileurus Arrow, 1908. 185192, E. minor Yang &amp; Pathomwattananurak, new species, 185187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193196, E. sidereus Yang &amp; Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="18" pageNumber="469">185192</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84F177E7EADFAD56ACCA753" pageId="18" pageNumber="469" type="etymology">
<paragraph id="8BCF368DE84F177E7EADFAD56ACCA753" blockId="18.[151,1437,150,1869]" pageId="18" pageNumber="469">
<emphasis id="B904EA9FE84F177E7EADFAD56A37A77F" bold="true" box="[199,334,1339,1365]" pageId="18" pageNumber="469">Etymology.</emphasis>
The name refers to its uniquely small size compared to similarly developed individuals of all other Indochinese
<taxonomicName id="4C704D0EE84F177E7F4FFAB16AD7A753" authorityName=", Endrodi" authorityYear="1977" box="[293,430,1375,1401]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B904EA9FE84F177E7F4FFAB16AD7A753" box="[293,430,1375,1401]" italics="true" pageId="18" pageNumber="469">Eophileurus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84F177E7EADFA6C6FA0A47B" pageId="18" pageNumber="469" type="distribution">
<paragraph id="8BCF368DE84F177E7EADFA6C6946A7B7" blockId="18.[151,1437,150,1869]" box="[199,575,1410,1437]" pageId="18" pageNumber="469">
<emphasis id="B904EA9FE84F177E7EADFA6C6A18A7B6" bold="true" box="[199,353,1410,1436]" pageId="18" pageNumber="469">Distribution.</emphasis>
<collectingCountry id="F367761DE84F177E7F0DFA6D6AB4A7B7" box="[359,461,1411,1437]" name="Thailand" pageId="18" pageNumber="469">Thailand</collectingCountry>
,
<collectingCountry id="F367761DE84F177E7FB2FA6D6942A7B7" box="[472,571,1411,1437]" name="Vietnam" pageId="18" pageNumber="469">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8BCF368DE84F177E7EADFA486FA0A47B" blockId="18.[151,1437,150,1869]" pageId="18" pageNumber="469">
<emphasis id="B904EA9FE84F177E7EADFA486A33A7EA" bold="true" box="[199,330,1446,1472]" pageId="18" pageNumber="469">Variations.</emphasis>
The phenomenon of unstable morphology of the parameres is also seen in this species. Indicated in
<figureCitation id="134B2A08E84F177E7EFDFA256A42A7CF" box="[151,315,1482,1509]" captionStart="FIGURES 185196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185196. Parameres of Eophileurus Arrow, 1908. 185192, E. minor Yang &amp; Pathomwattananurak, new species, 185187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193196, E. sidereus Yang &amp; Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="18" pageNumber="469">Figs. 185187</figureCitation>
, are parameres of males raised from eggs laid by the same wild female, in which variations such as the lateral processes are slightly differently shaped, and a pair of very small teeth can be seen on the lateral margin in the first two individuals but absent in the third. Distinct geographical variations can also be seen in
<figureCitation id="134B2A08E84F177E7A9CF9FD6EE1A407" box="[1270,1432,1554,1581]" captionStart="FIGURES 185196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185196. Parameres of Eophileurus Arrow, 1908. 185192, E. minor Yang &amp; Pathomwattananurak, new species, 185187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193196, E. sidereus Yang &amp; Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="18" pageNumber="469">Figs. 185192</figureCitation>
, with slight variations on certain parts which are mentioned above in the
<typeStatus id="54CB882FE84F177E7DAAF9D96F52A47B" box="[960,1067,1591,1617]" pageId="18" pageNumber="469" type="paratype">paratypes</typeStatus>
(male) section.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84F177E7EADF9B56949A567" pageId="18" pageNumber="469" type="discussion">
<paragraph id="8BCF368DE84F177E7EADF9B56949A567" blockId="18.[151,1437,150,1869]" pageId="18" pageNumber="469">
<emphasis id="B904EA9FE84F177E7EADF9B56A42A45F" bold="true" box="[199,315,1627,1653]" pageId="18" pageNumber="469">Remarks.</emphasis>
This species is likely closely related to some Indian species due to the following two aspects. Firstly, the morphology of its parameres (
<figureCitation id="134B2A08E84F177E7C70F99169B9A4B3" box="[538,704,1663,1689]" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="18" pageNumber="469">Figs. 169172</figureCitation>
,
<figureCitation id="134B2A08E84F177E7CA6F9906856A4B3" box="[716,815,1662,1689]" captionStart="FIGURES 185196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185196. Parameres of Eophileurus Arrow, 1908. 185192, E. minor Yang &amp; Pathomwattananurak, new species, 185187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193196, E. sidereus Yang &amp; Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="18" pageNumber="469">185192</figureCitation>
) has an overlapping region close to the center in front view which is shared in many Indian species such as
<taxonomicName id="4C704D0EE84F177E7C91F94D6883A497" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[763,1018,1699,1725]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE84F177E7C91F94D680BA497" box="[763,882,1699,1725]" italics="true" pageId="18" pageNumber="469">E. felschei</emphasis>
<bibRefCitation id="EFE14B7CE84F177E7D10F94D6883A497" author="Prell, H." box="[890,1018,1699,1725]" pageId="18" pageNumber="469" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
, (
<figureCitation id="134B2A08E84F177E7A64F94D6FE1A497" box="[1038,1176,1698,1725]" captionStart="FIGURES 3848" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[253,1339,186,1912]" captionTargetId="figure-18@10.[241,1346,181,1920]" captionTargetPageId="10" captionText="FIGURES 3848. Type specimens of Eophileurus felschei Prell, 1913. 3831, lectotype, female, 38, habitus, dorsal; 39, habitus, lateral; 40, habitus, ventral; 41, labels; 4248, lectoparatype, male (images ©MTD/Olaf Jäger), 42, habitus, dorsal; 43, habitus, lateral; 44, habitus, ventral, 4546, parameres, apical; 47, parameres, lateral; 48, labels." figureDoi="http://doi.org/10.5281/zenodo.6853956" httpUri="https://zenodo.org/record/6853956/files/figure.png" pageId="18" pageNumber="469">Figs. 4547</figureCitation>
,
<figureCitation id="134B2A08E84F177E7ACFF94C6F96A497" box="[1189,1263,1698,1725]" captionStart="FIGURES 4953" captionStartId="11.[151,264,1333,1357]" captionTargetBox="[156,1433,418,1309]" captionTargetId="figure-149@11.[151,1435,417,1309]" captionTargetPageId="11" captionText="FIGURES 4953. Holotype of Eophileurus grossepunctatus Dupuis, 2014 (images ©MNHN/A. Mantilleri). 49, habitus, dorsal; 50, habitus, lateral; 51, parameres, apical; 52, parameres, lateral; 53, labels." figureDoi="http://doi.org/10.5281/zenodo.6853958" httpUri="https://zenodo.org/record/6853958/files/figure.png" pageId="18" pageNumber="469">5152</figureCitation>
). Secondly, the small size of this species is unique amongst Indochinese species, whereas it is more commonly seen in Indian species. All specimens of this species examined are from low altitude regions. Additionally, the gap in its known distribution between eastern
<collectingCountry id="F367761DE84F177E7FB7F8E16938A503" box="[477,577,1807,1833]" name="Thailand" pageId="18" pageNumber="469">Thailand</collectingCountry>
and southern
<collectingCountry id="F367761DE84F177E7C8BF8E16839A503" box="[737,832,1807,1833]" name="Vietnam" pageId="18" pageNumber="469">Vietnam</collectingCountry>
suggests probable occurrence in regions with similar environment in
<collectingCountry id="F367761DE84F177E7F23F8DD6AC5A567" box="[329,444,1843,1869]" name="Cambodia" pageId="18" pageNumber="469">Cambodia</collectingCountry>
and
<collectingCountry id="F367761DE84F177E7F98F8DD6955A567" box="[498,556,1843,1869]" name="Laos" pageId="18" pageNumber="469">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>