treatments-xml/data/03/C8/6C/03C86C16EB285754FF6EDE1CFD47F9B7.xml
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<document id="B247F5E4B1056C7C42854420CD72EF56" ID-CLB-Dataset="21673" ID-DOI="10.11646/zootaxa.4789.1.1" ID-GBIF-Dataset="c12aaed8-264c-459c-b1aa-09eaeb686670" ID-ISSN="1175-5326" ID-Zenodo-Dep="3884610" ID-ZooBank="311A27A2-A053-4D20-A43E-EF57DBE55166" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591601966137" checkinUser="plazi" docAuthor="Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit &amp; Katoh, Toru" docDate="2020" docId="03C86C16EB285754FF6EDE1CFD47F9B7" docLanguage="en" docName="zootaxa.4789.1.1.pdf" docOrigin="Zootaxa 4789 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phortica (Ashima) biclavata Chen in Chen &amp; Maca 2012" docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="FFF1146EEB26575AFFF9DD08FFEBFFAA" masterDocTitle="Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="15" updateTime="1698842174822" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1F0E50F1222D296439CC615D29E5445A">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:namePart id="2282340346F2C805005CA477625287FD">Toda, Masanori J.</mods:namePart>
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<mods:namePart id="224815D131351347E7AD18B10B193341">Bänziger, Hans</mods:namePart>
<mods:affiliation id="F79A94AB3B4D654A54DBF04F41B621D4">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
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<mods:namePart id="27E1D8B1F024548A7E1F236588360E30">Sati, Pradeep C.</mods:namePart>
<mods:affiliation id="A68810346E5E208CC72EB1154179C055">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575</mods:affiliation>
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<mods:namePart id="5FB4E758714E9878F22D9DAE6AAA2B96">Fartyal, Rajendra S.</mods:namePart>
<mods:affiliation id="73F78C167DF1B087043215722AC45DF1">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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<mods:affiliation id="2713DDD727A3AF261C2635966FA03D43">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com</mods:affiliation>
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<heading id="D0966A6CEB285754FF6EDE1CFD74FC84" bold="true" box="[151,671,788,815]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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5)
<taxonomicName id="4C61A683EB285754FF4EDE1CFD74FC84" ID-CoL="9HXD2" authority="Chen, 2012" authorityName="Chen in Chen &amp; Maca" authorityYear="2012" box="[183,671,788,815]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="biclavata" subGenus="Ashima">
<emphasis id="B9150112EB285754FF4EDE1CFEF4FC85" bold="true" box="[183,287,788,815]" italics="true" pageId="14" pageNumber="15">Phortica</emphasis>
(
<emphasis id="B9150112EB285754FEC9DE1CFE66FC85" bold="true" box="[304,397,788,815]" italics="true" pageId="14" pageNumber="15">Ashima</emphasis>
)
<emphasis id="B9150112EB285754FE67DE1CFDE0FC85" bold="true" box="[414,523,788,815]" italics="true" pageId="14" pageNumber="15">biclavata</emphasis>
<bibRefCitation id="EFF0A0F1EB285754FDEADE1CFD74FC84" author="Chen" box="[531,671,788,814]" firstAuthor="Chen" pageId="14" pageNumber="15" pagination="493 - 509" refId="ref33941" refString="Chen, H. W. &amp; Maca, J. (2012) Ten new species of the genus Phortica from the Afrotropical and Oriental regions (Diptera: Drosophilidae). Zootaxa, 3478 (1), 493 - 509. https: // doi. org / 10.11646 / zootaxa. 3478.1.42" type="journal article" year="2012">Chen, 2012</bibRefCitation>
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<taxonomicName id="4C61A683EB285754FF6EDE50FCF3FCDA" ID-CoL="9HXD2" authority="Chen in Chen &amp; Maca, 2012: 499" authorityName="Chen in Chen &amp; Maca" authorityPageNumber="499" authorityYear="2012" box="[151,792,856,881]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="biclavata" subGenus="Ashima">
<emphasis id="B9150112EB285754FF6EDE50FF1BFCDA" box="[151,240,856,880]" italics="true" pageId="14" pageNumber="15">Phortica</emphasis>
(
<emphasis id="B9150112EB285754FF06DE50FEA0FCDA" box="[255,331,856,880]" italics="true" pageId="14" pageNumber="15">Ashima</emphasis>
)
<emphasis id="B9150112EB285754FEA0DE50FE5CFCDA" box="[345,439,856,880]" italics="true" pageId="14" pageNumber="15">biclavata</emphasis>
Chen in
<bibRefCitation id="EFF0A0F1EB285754FDEDDE50FCF3FCDA" author="Chen &amp; Maca" box="[532,792,856,881]" firstAuthor="Chen" pageId="14" pageNumber="15" pagination="493 - 509" refId="ref33941" refString="Chen, H. W. &amp; Maca, J. (2012) Ten new species of the genus Phortica from the Afrotropical and Oriental regions (Diptera: Drosophilidae). Zootaxa, 3478 (1), 493 - 509. https: // doi. org / 10.11646 / zootaxa. 3478.1.42" type="journal article" year="2012">Chen &amp; Máca, 2012: 499</bibRefCitation>
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.
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<emphasis id="B9150112EB285754FF6EDE96FEF9FC13" bold="true" box="[151,274,926,953]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
Aedeagal sheath with submedial and anterior, sclerotized, marginally serrated postgonites on left lateral lobe (“Fig. 18” in
<bibRefCitation id="EFF0A0F1EB285754FE9DDECBFDAEFC77" author="Chen &amp; Maca" box="[356,581,963,989]" firstAuthor="Chen" pageId="14" pageNumber="15" pagination="493 - 509" refId="ref33941" refString="Chen, H. W. &amp; Maca, J. (2012) Ten new species of the genus Phortica from the Afrotropical and Oriental regions (Diptera: Drosophilidae). Zootaxa, 3478 (1), 493 - 509. https: // doi. org / 10.11646 / zootaxa. 3478.1.42" type="journal article" year="2012">Chen &amp; Máca 2012</bibRefCitation>
).
</paragraph>
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<emphasis id="B9150112EB285754FF3EDEEEFDF8FBAB" bold="true" box="[199,531,998,1025]" pageId="14" pageNumber="15">Supplementary description.</emphasis>
Supracervical setae 1517. Dorsomedial, tentorial apodeme 1/2 as long as basal, parallel portion of dorsolateral, tentorial apodeme. Some interfrontal setulae as long as lowest setulae on ocellar triangle (ch.3-0). Arista with neither dorsal nor ventral branches (ch.5,6-2); apical, leaf-like expansion narrower than long, apically tapering, slightly serrated on apical margin. Cibarial, medial sensilla 79 per side; posterior sensilla 46 per side. Setulae just before prescutellar acrostichal setae somewhat longer than other acrostichal setulae (ch.15-1). Fore and mid tibiae entirely yellow (ch.17,18-0); all tarsi with gray tarsomeres IV and V. Abdominal sternite III wider than long (ch.22-1), widest among sternites (ch.23-1), with dense setulae on medial surface (ch.26-1); IV and V wider than long (ch.24,25-1). Surstylus without expansion at caudoventral corner (ch.35-0). Pregonite not concave on dorsosubapical margin of apical knob (ch.48-0). Sclerotized postgonites on lateral lobes of aedeagal sheath medially separated from each other (ch.66-0).
</paragraph>
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<emphasis id="B9150112EB285754FF3ED846FE2CFAC3" bold="true" box="[199,455,1358,1385]" pageId="14" pageNumber="15">Specimens examined.</emphasis>
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:
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,
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,
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<collectingDate id="EF9B0228EB285754FCC3D847FC4DFAC3" box="[826,934,1359,1385]" pageId="14" pageNumber="15" value="1982-01-14">14.i.1982</collectingDate>
</date>
,
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leg. (
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)
</materialsCitation>
;
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<specimenCount id="9D671689EB285754FB3CD847FB03FAC3" box="[1221,1256,1359,1385]" pageId="14" pageNumber="15" type="male">2♂</specimenCount>
<typeStatus id="54DA63A2EB285754FB16D847FAB6FAC3" box="[1263,1373,1359,1385]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
, ditto (
<collectionCode id="ED7045C5EB285754FF66D87BFF05FA27" box="[159,238,1395,1421]" pageId="14" pageNumber="15">SEHU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B09D75DEB285754FEF9D87BFD14FA27" ID-GBIF-Occurrence="2636268313" box="[256,767,1395,1421]" collectingDate="1982-01-10" collectionCode="SEHU" collectorName="M. J. Toda" country="Myanmar" location="Yangon" pageId="14" pageNumber="15" specimenCount="2" specimenCount-male="2" stateProvince="Yangon" typeStatus="paratype">
<specimenCount id="9D671689EB285754FEF9D87BFEC8FA27" box="[256,291,1395,1421]" pageId="14" pageNumber="15" type="male">2♂</specimenCount>
<typeStatus id="54DA63A2EB285754FED0D87BFE7CFA27" box="[297,407,1395,1421]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
, ditto, except
<date id="FFDFFBC0EB285754FDC9D87BFD71FA27" box="[560,666,1395,1421]" pageId="14" pageNumber="15" value="1982-01-10">
<collectingDate id="EF9B0228EB285754FDC9D87BFD71FA27" box="[560,666,1395,1421]" pageId="14" pageNumber="15" value="1982-01-10">10.i.1982</collectingDate>
</date>
(
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)
</materialsCitation>
;
<materialsCitation id="3B09D75DEB285754FCF0D87BFADBFA27" ID-GBIF-Occurrence="2636268303" box="[777,1328,1394,1421]" collectingDate="1982-01-11" collectingDateMax="1982-01-15" collectingDateMin="1982-01-11" collectionCode="SEHU" collectorName="M. J. Toda" country="Myanmar" location="Yangon" pageId="14" pageNumber="15" specimenCount="4" specimenCount-male="4" stateProvince="Yangon" typeStatus="paratype">
<specimenCount id="9D671689EB285754FCF0D87BFCC7FA27" box="[777,812,1395,1421]" pageId="14" pageNumber="15" type="male">4♂</specimenCount>
<typeStatus id="54DA63A2EB285754FCC8D87BFC74FA27" box="[817,927,1395,1421]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
, ditto, except
<date id="FFDFFBC0EB285754FBC1D87AFB20FA27" box="[1080,1227,1394,1421]" pageId="14" pageNumber="15" value="1982-01-11" valueMax="1982-01-15" valueMin="1982-01-11">
<collectingDate id="EF9B0228EB285754FBC1D87AFB20FA27" box="[1080,1227,1394,1421]" pageId="14" pageNumber="15" value="1982-01-11" valueMax="1982-01-15" valueMin="1982-01-11">1115.i.1982</collectingDate>
</date>
(
<collectionCode id="ED7045C5EB285754FB20D87BFAC3FA27" box="[1241,1320,1395,1421]" pageId="14" pageNumber="15">SEHU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B09D75DEB285754FAC3D87BFCB2FA1B" ID-GBIF-Occurrence="2636268355" collectingDate="1981-12-26" collectingDateMax="1982-01-04" collectingDateMin="1981-12-26" collectionCode="SEHU" collectorName="M. J. Toda" country="Myanmar" location="Mandalay" pageId="14" pageNumber="15" specimenCount="4" specimenCount-male="4" stateProvince="Mandalay" typeStatus="paratype">
<specimenCount id="9D671689EB285754FAC3D87BFAB6FA27" box="[1338,1373,1395,1421]" pageId="14" pageNumber="15" type="male">4♂</specimenCount>
para- types,
<collectingRegion id="49A513E2EB285754FF19D89FFEBAFA1B" box="[224,337,1431,1457]" country="Myanmar" name="Mandalay" pageId="14" pageNumber="15">Mandalay</collectingRegion>
,
<date id="FFDFFBC0EB285754FEA5D89FFE36FA1B" box="[348,477,1431,1457]" pageId="14" pageNumber="15" value="1981-12-26">
<collectingDate id="EF9B0228EB285754FEA5D89FFE36FA1B" box="[348,477,1431,1457]" pageId="14" pageNumber="15" value="1981-12-26">26.xii.1981</collectingDate>
</date>
<date id="FFDFFBC0EB285754FE14D89FFDA3FA1B" box="[493,584,1431,1457]" pageId="14" pageNumber="15" value="1982-01-04">
<collectingDate id="EF9B0228EB285754FE14D89FFDA3FA1B" box="[493,584,1431,1457]" pageId="14" pageNumber="15" value="1982-01-04">4.i.1982</collectingDate>
</date>
,
<collectorName id="2694B8D6EB285754FDAAD89FFD28FA1B" box="[595,707,1431,1457]" pageId="14" pageNumber="15">M.J. Toda</collectorName>
leg. (
<collectionCode id="ED7045C5EB285754FCFBD89FFCBAFA1B" box="[770,849,1431,1457]" pageId="14" pageNumber="15">SEHU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B09D75DEB285754FC9AD89FFCB9FA7F" ID-GBIF-Occurrence="2636268332" collectingDate="2008-02-22" collectionCode="SEHU" collectorName="H. Banziger" country="Thailand" elevation="700" location="Chiang Dao" municipality="Tonluong Temple" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
<collectingCountry id="F3769D90EB285754FC9AD89FFC20FA1B" box="[867,971,1431,1457]" name="Thailand" pageId="14" pageNumber="15">Thailand</collectingCountry>
:
<specimenCount id="9D671689EB285754FC2CD89FFC12FA1B" box="[981,1017,1431,1457]" pageId="14" pageNumber="15" type="male">1♂</specimenCount>
, near
<collectingMunicipality id="6BBA477AEB285754FBC5D89FFAEFFA1B" box="[1084,1284,1431,1457]" pageId="14" pageNumber="15">Tonluong Temple</collectingMunicipality>
,
<location id="8EBE8BDBEB285754FAF6D89FFA73FA1B" LSID="urn:lsid:plazi:treatment:03C86C16EB285754FF6EDE1CFD47F9B7:8EBE8BDBEB285754FAF6D89FFA73FA1B" box="[1295,1432,1431,1457]" country="Thailand" municipality="Tonluong Temple" name="Chiang Dao" pageId="14" pageNumber="15" stateProvince="Chiang Mai">Chiang Dao</location>
,
<collectingRegion id="49A513E2EB285754FF6ED8B3FEF5FA7F" box="[151,286,1467,1493]" country="Thailand" name="Chiang Mai" pageId="14" pageNumber="15">Chiang Mai</collectingRegion>
,
<elevation id="004C3A33EB285754FED0D8B3FE40FA7F" box="[297,427,1467,1493]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="15" unit="m" value="700.0">
<quantity id="4C9970E5EB285754FED0D8B3FE84FA7F" box="[297,367,1467,1493]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="15" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFBC0EB285754FE4ED8B3FDC0FA7F" box="[439,555,1467,1493]" pageId="14" pageNumber="15" value="2008-02-22">
<collectingDate id="EF9B0228EB285754FE4ED8B3FDC0FA7F" box="[439,555,1467,1493]" pageId="14" pageNumber="15" value="2008-02-22">22.ii.2008</collectingDate>
</date>
,
<collectorName id="2694B8D6EB285754FDCFD8B3FD56FA7F" box="[566,701,1467,1493]" pageId="14" pageNumber="15">H. Bänziger</collectorName>
leg. (
<collectionCode id="ED7045C5EB285754FD02D8B3FCA1FA7F" box="[763,842,1467,1493]" pageId="14" pageNumber="15">SEHU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B09D75DEB285754FCA4D8B3FB06FA7F" ID-GBIF-Occurrence="2636268395" box="[861,1261,1467,1493]" collectingDate="2008-11-06" collectionCode="SEHU" collectorName="H. Banziger" country="Thailand" elevation="700" location="Chiang Dao" municipality="Tonluong Temple" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
<specimenCount id="9D671689EB285754FCA4D8B3FC6AFA7F" box="[861,897,1467,1493]" pageId="14" pageNumber="15" type="male">1♂</specimenCount>
, ditto, except
<date id="FFDFFBC0EB285754FBE7D8B3FB6CFA7F" box="[1054,1159,1467,1493]" pageId="14" pageNumber="15" value="2008-11-06">
<collectingDate id="EF9B0228EB285754FBE7D8B3FB6CFA7F" box="[1054,1159,1467,1493]" pageId="14" pageNumber="15" value="2008-11-06">6.xi.2008</collectingDate>
</date>
(
<collectionCode id="ED7045C5EB285754FB6FD8B3FB0EFA7F" box="[1174,1253,1467,1493]" pageId="14" pageNumber="15">SEHU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B09D75DEB285754FB0ED8B3FC99FA53" ID-GBIF-Occurrence="2636268335" collectingDate="2009-07-26" collectionCode="SEHU" collectorName="H. Banziger" country="Laos" location="Phousi" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Louangphabang" typeStatus="paratype">
<collectingCountry id="F3769D90EB285754FB0ED8B3FAD9FA7F" box="[1271,1330,1467,1493]" name="Laos" pageId="14" pageNumber="15">Laos</collectingCountry>
:
<specimenCount id="9D671689EB285754FAC5D8B3FA8BFA7F" box="[1340,1376,1467,1493]" pageId="14" pageNumber="15" type="male">1♂</specimenCount>
, path to
<location id="8EBE8BDBEB285754FF4DD8D7FEE8FA53" LSID="urn:lsid:plazi:treatment:03C86C16EB285754FF6EDE1CFD47F9B7:8EBE8BDBEB285754FF4DD8D7FEE8FA53" box="[180,259,1503,1529]" country="Laos" name="Phousi" pageId="14" pageNumber="15" stateProvince="Louangphabang">Phousi</location>
,
<collectingRegion id="49A513E2EB285754FEF7D8D7FE57FA53" box="[270,444,1503,1529]" country="Laos" name="Louangphabang" pageId="14" pageNumber="15">Luang Prabang</collectingRegion>
,
<date id="FFDFFBC0EB285754FE3ED8D7FDA3FA53" box="[455,584,1503,1529]" pageId="14" pageNumber="15" value="2009-07-26">
<collectingDate id="EF9B0228EB285754FE3ED8D7FDA3FA53" box="[455,584,1503,1529]" pageId="14" pageNumber="15" value="2009-07-26">26.vii.2009</collectingDate>
</date>
,
<collectorName id="2694B8D6EB285754FDADD8D7FD30FA53" box="[596,731,1503,1529]" pageId="14" pageNumber="15">H. Bänziger</collectorName>
leg. (
<collectionCode id="ED7045C5EB285754FCE2D8D7FC81FA53" box="[795,874,1503,1529]" pageId="14" pageNumber="15">SEHU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8E8BEB285754FF3EDB0AFD47F9B7" box="[199,684,1538,1565]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BDEDD00EB285754FF3EDB0AFD47F9B7" blockId="14.[151,1437,926,1565]" box="[199,684,1538,1565]" pageId="14" pageNumber="15">
<emphasis id="B9150112EB285754FF3EDB0AFE8AF9B7" bold="true" box="[199,353,1538,1565]" pageId="14" pageNumber="15">Distribution.</emphasis>
<collectingCountry id="F3769D90EB285754FE9EDB0BFE3DF9B7" box="[359,470,1539,1565]" name="Myanmar" pageId="14" pageNumber="15">Myanmar</collectingCountry>
,
<collectingCountry id="F3769D90EB285754FE18DB0BFDA1F9B7" box="[481,586,1539,1565]" name="Thailand" pageId="14" pageNumber="15">Thailand</collectingCountry>
*,
<collectingCountry id="F3769D90EB285754FD99DB0BFD77F9B7" box="[608,668,1539,1565]" name="Laos" pageId="14" pageNumber="15">Laos</collectingCountry>
*.
</paragraph>
</subSubSection>
</treatment>
</document>