169 lines
19 KiB
XML
169 lines
19 KiB
XML
<document id="394378194F0FB7CDF81E82662586ED26" ID-DOI="10.5281/zenodo.276689" ID-GBIF-Dataset="b6781430-d288-4cb4-b178-db6192a61dde" ID-ISSN="1175-5326" ID-Zenodo-Dep="276689" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460230557710" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2011" docId="03BC87EAFFB7317BFF22F8DA0456FC69" docLanguage="en" docName="zt02753p041.pdf" docOrigin="Zootaxa 2753" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Parabolus calceus Enghoff, 2011, n. sp." docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FF85FF92FFAD3160FFB5FFF00761FFF9" masterDocTitle="East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="27" updateTime="1698238933756" updateUser="plazi">
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<mods:titleInfo id="5F84454585C6509A240F27B197C6C199">
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<mods:title id="B0B59E272AB1C841D657A02A688DFAF6">East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:name id="30F701C8BDBAB7A8ABDFBECEB8AD4667" type="personal">
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<mods:roleTerm id="239ED48C614DCD00D2075194BD08FB12">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="60F6F46F42305BD9D3757B0356B0131E">Enghoff, Henrik</mods:namePart>
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<mods:typeOfResource id="7360729E504D58C06F81FAE28F484776">text</mods:typeOfResource>
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<mods:title id="3167D1832D6B495EB3C8439CD4C6AD2A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="85A86E640339837935FDCC8B211F34A4">
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<mods:date id="E6B8DC6857B93F15797EDD088A8462E9">2011</mods:date>
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<mods:detail id="C87EA177912DBEC5C0F41151D08EF1EB" type="volume">
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<mods:number id="143ECF33C280CB743700E22B3395CCB1">2753</mods:number>
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<mods:start id="113DA5F9A0A84FF9A84F0A093AADB9F7">1</mods:start>
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<mods:end id="BDE194F7F72B3F9ECC14EC8DA49E3519">41</mods:end>
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<mods:identifier id="3F58B57BA81BBBF8C172BB33E14E7879" type="DOI">10.5281/zenodo.276689</mods:identifier>
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<mods:identifier id="C6AE0E8AD348249B1667C43DFE6C080A" type="GBIF-Dataset">b6781430-d288-4cb4-b178-db6192a61dde</mods:identifier>
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<mods:identifier id="834082B740366DB6C1DFA8632F67EA30" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="300FD5BDD29914203E48939AB992D8FB" type="Zenodo-Dep">276689</mods:identifier>
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<treatment id="03BC87EAFFB7317BFF22F8DA0456FC69" ID-DOI="http://doi.org/10.5281/zenodo.5612664" ID-GBIF-Taxon="119387228" ID-Zenodo-Dep="5612664" LSID="urn:lsid:plazi:treatment:03BC87EAFFB7317BFF22F8DA0456FC69" httpUri="http://treatment.plazi.org/id/03BC87EAFFB7317BFF22F8DA0456FC69" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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<subSubSection id="C30F6577FFB7317AFF22F8DA065FF89D" pageId="26" pageNumber="27" type="nomenclature">
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<paragraph id="8BAA36FCFFB7317AFF22F8DA06DCF8BA" blockId="26.[151,445,1833,1892]" box="[151,445,1833,1860]" pageId="26" pageNumber="27">
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<heading id="D0E28190FFB7317AFF22F8DA06DCF8BA" bold="true" box="[151,445,1833,1860]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
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<emphasis id="B961EAEEFFB7317AFF22F8DA06DCF8BA" bold="true" box="[151,445,1833,1860]" pageId="26" pageNumber="27">
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<taxonomicName id="4C154D7FFFB7317AFF22F8DA0615F8BD" box="[151,372,1834,1860]" class="Diplopoda" family="Pachybolidae" genus="Parabolus" kingdom="Animalia" order="Spirobolida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="calceus" status="sp. nov.">
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<emphasis id="B961EAEEFFB7317AFF22F8DA0615F8BD" bold="true" box="[151,372,1834,1860]" italics="true" pageId="26" pageNumber="27">Parabolus calceus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2525795FFB7317AFECEF8D906DCF8BA" box="[379,445,1833,1859]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAA36FCFFB7317AFF22F8BC065FF89D" blockId="26.[151,445,1833,1892]" box="[151,318,1868,1892]" pageId="26" pageNumber="27">
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<figureCitation id="132E2A79FFB7317AFF22F8BC0677F89D" box="[151,278,1868,1892]" captionStart="FIGURES 54 – 57" captionStartId="28.[151,264,1850,1873]" captionTargetBox="[189,1398,201,1826]" captionTargetId="figure@28.[183,1403,195,1829]" captionTargetPageId="28" captionText="FIGURES 54 – 57. Parabolus calceus, paratypes. 54: anterior gonopods, without right telopodite, anterior view, 55: do., posterior view, 56: left posterior gonopod, anterior view, 57: do, posterior view. Scale for all: 2 cm. cx: coxa, mr: mesobasal ridge, st: sternum, tlp: telopodite, tt: triangular tooth." httpUri="https://zenodo.org/record/276709/files/figure.png" pageId="26" pageNumber="27">Figs 54–57</figureCitation>
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,
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<figureCitation id="132E2A79FFB7317AFE97F8BC065FF89D" box="[290,318,1868,1892]" captionStart="FIGURE 84" captionStartId="36.[151,249,1479,1502]" captionTargetBox="[456,1120,410,1444]" captionTargetId="figure@36.[456,1122,407,1445]" captionTargetPageId="36" captionText="FIGURE 84. Known distribution of East African Pachybolini, except Hadrobolus crassicollis and Epibolus pulchripes. dots (●): Crurifarcimen vagans, filled diamonds (♦): Hyperbolus apicomplexus, open diamonds (♢): Hyperbolus morogoroensis, filled triangle (˔): Parabolus dimorphus, inverted open triangle (): Parabolus calceus, inverted filled triangle (ˑ): Pachybolus tectus,, filled square (․): Pachybolini indet. Based on Yahoo maps." httpUri="https://zenodo.org/record/276715/files/figure.png" pageId="26" pageNumber="27">84</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F6577FFB7317AFF22F87F0547F816" pageId="26" pageNumber="27" type="materials_examined">
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<paragraph id="8BAA36FCFFB7317AFF22F87F0547F816" blockId="26.[151,1436,1935,2032]" pageId="26" pageNumber="27">
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<typeStatus id="54AE885EFFB7317AFF22F87F064AF85E" box="[151,299,1935,1959]" pageId="26" pageNumber="27" type="holotype">HOLOTYPE</typeStatus>
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3
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<collectingCountry id="F302766CFFB7317AFEF8F86006CEF851" box="[333,431,1936,1960]" name="Kenya" pageId="26" pageNumber="27">KENYA</collectingCountry>
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, Gede,
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<geoCoordinate id="EE21503BFFB7317AFDB6F87F0532F85E" box="[515,595,1935,1959]" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="-3.4666667">3°28’S</geoCoordinate>
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, 39°18E,
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<date id="FFAB103CFFB7317AFD74F87F045BF85E" box="[705,826,1935,1959]" pageId="26" pageNumber="27" value="1981-04-29">29.iv.1981</date>
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, CIE A13123 (
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<collectionCode id="ED04AE39FFB7317AFC54F87F035BF85E" box="[993,1082,1935,1959]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="26" pageNumber="27">NHML</collectionCode>
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). –
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<typeStatus id="54AE885EFFB7317AFBD4F87F0261F85E" box="[1121,1280,1935,1959]" pageId="26" pageNumber="27" type="paratype">PARATYPES</typeStatus>
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: 2 3, 3 ƤƤ, strongly fragmented,
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<collectingCountry id="F302766CFFB7317AFE3CF844068AF835" box="[393,491,1972,1996]" name="Kenya" pageId="26" pageNumber="27">KENYA</collectingCountry>
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, Changamwe
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<quantity id="4CED9B19FFB7317AFD39F84305BEF832" box="[652,735,1971,1995]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="26" pageNumber="27" unit="mi" value="4.0">4 miles</quantity>
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W of
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<collectingCountry id="F302766CFFB7317AFC9CF84304FAF832" box="[809,923,1971,1995]" name="Kenya" pageId="26" pageNumber="27">Mombasa</collectingCountry>
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,
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<geoCoordinate id="EE21503BFFB7317AFC12F8430345F832" box="[935,1060,1971,1995]" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="-4.025">4°01’30’’S</geoCoordinate>
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,
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<geoCoordinate id="EE21503BFFB7317AFB84F84303DCF832" box="[1073,1213,1971,1995]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="39.629166">39°37’45’’E</geoCoordinate>
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,
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<date id="FFAB103CFFB7317AFB7FF8430258F832" box="[1226,1337,1971,1995]" pageId="26" pageNumber="27" value="1921-03-30">21–30.iii.</date>
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(?) 1909, E.A. Mearns leg. (
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<collectionCode id="ED04AE39FFB7317AFED1F82706D8F816" LSID="urn:lsid:biocol.org:col:34871" box="[356,441,2007,2031]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
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,
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<collectionCode id="ED04AE39FFB7317AFE71F827057BF816" box="[452,538,2007,2031]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="26" pageNumber="27">ZMUC</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F6577FFB6317BFF72FF6804F1FF2D" pageId="27" pageNumber="28" type="etymology">
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<paragraph id="8BAA36FCFFB6317BFF72FF6804F1FF2D" blockId="27.[151,1437,151,912]" pageId="27" pageNumber="28">
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<emphasis id="B961EAEEFFB6317BFF72FF68062FFF49" bold="true" box="[199,334,152,176]" pageId="27" pageNumber="28">Etymology.</emphasis>
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The specific epithet is a Latin noun in apposition meaning “shoe” and refers to the tip of the posterior gonopod which somewhat resembles a (rather distorted) shoe.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F6577FFB6317BFF72FF1104AFFDBD" pageId="27" pageNumber="28" type="diagnosis">
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<paragraph id="8BAA36FCFFB6317BFF72FF110559FEBA" blockId="27.[151,1437,151,912]" pageId="27" pageNumber="28">
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<emphasis id="B961EAEEFFB6317BFF72FF110620FF03" bold="true" box="[199,321,225,250]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
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A species of
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<taxonomicName id="4C154D7FFFB6317BFE6CFF11052FFF00" box="[473,590,225,249]" class="Diplopoda" family="Pachybolidae" genus="Parabolus" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B961EAEEFFB6317BFE6CFF11052FFF00" box="[473,590,225,249]" italics="true" pageId="27" pageNumber="28">Parabolus</emphasis>
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</taxonomicName>
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in which the anterior gonopod coxa has a distinct distolateral shoulder, and the terminal segment of the posterior gonopod has a faintly serrate mesobasal ridge, a strongly concave distolateral profile and no slender apical process.
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</paragraph>
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<paragraph id="8BAA36FCFFB6317BFF72FEBF03D4FE9E" blockId="27.[151,1437,151,912]" box="[199,1205,335,360]" pageId="27" pageNumber="28">
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<emphasis id="B961EAEEFFB6317BFF72FEBF06F2FE91" bold="true" box="[199,403,335,360]" pageId="27" pageNumber="28">Descriptive notes</emphasis>
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(mainly based on
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<typeStatus id="54AE885EFFB6317BFDD7FEBF05ABFE9E" box="[610,714,335,359]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
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): As
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<taxonomicName id="4C154D7FFFB6317BFCB7FEA0049BFE91" box="[770,1018,336,360]" class="Diplopoda" family="Pachybolidae" genus="Crurifarcimen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vagans">
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<emphasis id="B961EAEEFFB6317BFCB7FEA0049BFE91" box="[770,1018,336,360]" italics="true" pageId="27" pageNumber="28">Crurifarcimen vagans</emphasis>
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</taxonomicName>
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except as noted:
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</paragraph>
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<paragraph id="8BAA36FCFFB6317BFF72FE8404AFFDBD" blockId="27.[151,1437,151,912]" pageId="27" pageNumber="28">
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57 podous rings, no apodous rings. L
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<quantity id="4CED9B19FFB6317BFDC1FE8405DCFE75" box="[628,701,372,396]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="28" unit="cm" value="14.0">14 cm</quantity>
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, B 13.4 mm. 10–11 rows of ocelli in six horizontal series, 41–44 ocelli per eye. 2+2 labral teeth, labral setae not visible. Lamellae linguales each with 5–6 distal setae. Collum smooth, no striae, with marginal furrow at anterior margin. Body rings: tergo-pleural suture visible on pro-, meso- and metazonite. Terga smooth, except for fine oblique striae ventrally on mesozona and fine, dense longitudinal striae on metazona. Ozopores 1 diameter in front of suture. Legs: length 51% of body width (markedly shorter than in
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<taxonomicName id="4C154D7FFFB6317BFF01FDDE0626FDBC" box="[180,327,557,581]" class="Diplopoda" family="Pachybolidae" genus="Parabolus" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="dimorphus">
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<emphasis id="B961EAEEFFB6317BFF01FDDE0626FDBC" box="[180,327,557,581]" italics="true" pageId="27" pageNumber="28">P. dimorphus</emphasis>
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</taxonomicName>
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). Telson as in
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<taxonomicName id="4C154D7FFFB6317BFE5EFDDD05AEFDBD" box="[491,719,556,581]" class="Diplopoda" family="Pachybolidae" genus="Crurifarcimen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="variety" variety="but">
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<emphasis id="B961EAEEFFB6317BFE5EFDDD05C5FDBC" box="[491,676,557,581]" italics="true" pageId="27" pageNumber="28">Crurifarcimen v.</emphasis>
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but
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</taxonomicName>
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dorsal profile straight.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F6577FFB6317BFF72FDA205B3FCBF" pageId="27" pageNumber="28" type="description">
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<paragraph id="8BAA36FCFFB6317BFF72FDA2040BFCD8" blockId="27.[151,1437,151,912]" pageId="27" pageNumber="28">
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<emphasis id="B961EAEEFFB6317BFF72FDA2068BFD93" bold="true" box="[199,490,594,618]" pageId="27" pageNumber="28">Male sexual characters.</emphasis>
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Anterior gonopods (
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<figureCitation id="132E2A79FFB6317BFD55FDA10405FD90" box="[736,868,593,617]" captionStart="FIGURES 54 – 57" captionStartId="28.[151,264,1850,1873]" captionTargetBox="[189,1398,201,1826]" captionTargetId="figure@28.[183,1403,195,1829]" captionTargetPageId="28" captionText="FIGURES 54 – 57. Parabolus calceus, paratypes. 54: anterior gonopods, without right telopodite, anterior view, 55: do., posterior view, 56: left posterior gonopod, anterior view, 57: do, posterior view. Scale for all: 2 cm. cx: coxa, mr: mesobasal ridge, st: sternum, tlp: telopodite, tt: triangular tooth." httpUri="https://zenodo.org/record/276709/files/figure.png" pageId="27" pageNumber="28">Figs 54–55</figureCitation>
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) as in
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<taxonomicName id="4C154D7FFFB6317BFC09FDA20335FD90" box="[956,1108,593,617]" class="Diplopoda" family="Pachybolidae" genus="Parabolus" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="dimorphus">
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<emphasis id="B961EAEEFFB6317BFC09FDA20335FD90" box="[956,1108,593,617]" italics="true" pageId="27" pageNumber="28">P. dimorphus</emphasis>
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</taxonomicName>
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, except that the coxa has a marked distolateral ’shoulder’, is apically truncate and lacks the internal ridge at the base of the mesal process Posterior gonopods (
|
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<figureCitation id="132E2A79FFB6317BFECAFD6B069EFD4A" box="[383,511,667,691]" captionStart="FIGURES 54 – 57" captionStartId="28.[151,264,1850,1873]" captionTargetBox="[189,1398,201,1826]" captionTargetId="figure@28.[183,1403,195,1829]" captionTargetPageId="28" captionText="FIGURES 54 – 57. Parabolus calceus, paratypes. 54: anterior gonopods, without right telopodite, anterior view, 55: do., posterior view, 56: left posterior gonopod, anterior view, 57: do, posterior view. Scale for all: 2 cm. cx: coxa, mr: mesobasal ridge, st: sternum, tlp: telopodite, tt: triangular tooth." httpUri="https://zenodo.org/record/276709/files/figure.png" pageId="27" pageNumber="28">Figs 56–57</figureCitation>
|
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) as in
|
||
<taxonomicName id="4C154D7FFFB6317BFDFCFD6C05BCFD4A" box="[585,733,667,691]" class="Diplopoda" family="Pachybolidae" genus="Parabolus" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="dimorphus">
|
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<emphasis id="B961EAEEFFB6317BFDFCFD6C05BCFD4A" box="[585,733,667,691]" italics="true" pageId="27" pageNumber="28">P. dimorphus</emphasis>
|
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</taxonomicName>
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, except for shape of terminal segment which has a low, faintly serrate mesobasal ridge (instead of the coarsely serrate lobe of
|
||
<emphasis id="B961EAEEFFB6317BFCD4FD310490FD21" box="[865,1009,704,728]" italics="true" pageId="27" pageNumber="28">P. d i m o r p h u s</emphasis>
|
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), a strongly concave distolateral profile, concavity basally delimited by strong triangular tooth; distolateral surface deeply hollowed, forming a long, curved trough; oral wall of trough with a low subtriangular lobe.
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</paragraph>
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<paragraph id="8BAA36FCFFB6317BFF72FCDE05B3FCBF" blockId="27.[151,1437,151,912]" box="[199,722,814,838]" pageId="27" pageNumber="28">
|
||
<emphasis id="B961EAEEFFB6317BFF72FCDE0695FCBF" bold="true" box="[199,500,814,838]" pageId="27" pageNumber="28">Female sexual characters</emphasis>
|
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as in
|
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<emphasis id="B961EAEEFFB6317BFD83FCDF05B3FCBF" box="[566,722,814,838]" italics="true" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C154D7FFFB6317BFD83FCDF05AFFCBF" box="[566,718,814,838]" class="Diplopoda" family="Pachybolidae" genus="Parabolus" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="dimorphus">P. dimorphus</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F6577FFB6317BFF72FCA20456FC69" pageId="27" pageNumber="28" type="distribution">
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||
<paragraph id="8BAA36FCFFB6317BFF72FCA20456FC69" blockId="27.[151,1437,151,912]" pageId="27" pageNumber="28">
|
||
<emphasis id="B961EAEEFFB6317BFF72FCA20601FC92" bold="true" box="[199,352,850,875]" pageId="27" pageNumber="28">Distribution.</emphasis>
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Only known from two sites in SE
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<collectingCountry id="F302766CFFB6317BFD6AFCA4044DFC95" box="[735,812,852,876]" name="Kenya" pageId="27" pageNumber="28">Kenya</collectingCountry>
|
||
. (
|
||
<figureCitation id="132E2A79FFB6317BFC8BFCA304EEFC92" box="[830,911,851,875]" captionStart="FIGURE 84" captionStartId="36.[151,249,1479,1502]" captionTargetBox="[456,1120,410,1444]" captionTargetId="figure@36.[456,1122,407,1445]" captionTargetPageId="36" captionText="FIGURE 84. Known distribution of East African Pachybolini, except Hadrobolus crassicollis and Epibolus pulchripes. dots (●): Crurifarcimen vagans, filled diamonds (♦): Hyperbolus apicomplexus, open diamonds (♢): Hyperbolus morogoroensis, filled triangle (˔): Parabolus dimorphus, inverted open triangle (): Parabolus calceus, inverted filled triangle (ˑ): Pachybolus tectus,, filled square (․): Pachybolini indet. Based on Yahoo maps." httpUri="https://zenodo.org/record/276715/files/figure.png" pageId="27" pageNumber="28">Fig. 84</figureCitation>
|
||
) This is the only species of Pachybolini known from
|
||
<collectingCountry id="F302766CFFB6317BFF61FC880641FC69" box="[212,288,888,912]" name="Kenya" pageId="27" pageNumber="28">Kenya</collectingCountry>
|
||
, excepting the widespread
|
||
<emphasis id="B961EAEEFFB6317BFDFAFC880456FC69" box="[591,823,888,912]" italics="true" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C154D7FFFB6317BFDFAFC880452FC69" box="[591,819,888,912]" class="Actinopterygii" family="Labridae" genus="Epibolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="pulchripes">Epibolus pulchripes</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |