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<document id="F5E321BB1174E47EA0F69070C40DBBC8" ID-DOI="10.11646/zootaxa.4789.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3884685" ID-ZooBank="5D5199B5-8A6A-45F6-A8CA-7B3DBB1AC591" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591602096676" checkinUser="plazi" docAuthor="Pachelle, Paulo P. G., Carvalho, Leina, Alves, Douglas F. R. &amp; Anker, Arthur" docDate="2020" docId="03BC87E6FF8FFF88C8D7FACBE537FCC8" docLanguage="en" docName="zootaxa.4789.1.2.pdf" docOrigin="Zootaxa 4789 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lysmata uncicornis Holthuis &amp; Maurin 1952" docType="treatment" docVersion="5" lastPageNumber="76" masterDocId="FF85FF9EFF9BFF9DC840FFF8E617FFF3" masterDocTitle="A revision of the Brazilian species of Lysmata Risso, 1816 (Decapoda: Caridea Lysmatidae), with discussion of the morphological characters used in their identification" masterLastPageNumber="90" masterPageNumber="55" pageNumber="75" updateTime="1698773191148" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="29F988D9B5B8AF386DC7E609DB12A3F2">A revision of the Brazilian species of Lysmata Risso, 1816 (Decapoda: Caridea Lysmatidae), with discussion of the morphological characters used in their identification</mods:title>
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<mods:namePart id="A214C05F4FBBABDB2FA8323199126086">Pachelle, Paulo P. G.</mods:namePart>
<mods:affiliation id="BDC2E911C429F72B92D5CA77253A7391">Laboratório de Carcinologia, Museu de Zoologia da Universidade de São Paulo (MZUSP), Av. Nazaré 481, Ipiranga, São Paulo, SP, 04263 - 000, Brazil</mods:affiliation>
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<mods:namePart id="5378ECFC2DBBAA9731A99C6E34029660">Carvalho, Leina</mods:namePart>
<mods:affiliation id="31150A59A416C18DEBD02C2EFA66206B">Laboratório de Carcinologia, Museu de Zoologia da Universidade de São Paulo (MZUSP), Av. Nazaré 481, Ipiranga, São Paulo, SP, 04263 - 000, Brazil</mods:affiliation>
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<mods:namePart id="74FAAA72B8A9370CC03AE3A363157F50">Alves, Douglas F. R.</mods:namePart>
<mods:affiliation id="1455E199435492E5B76A6BF05D325D00">Laboratório de Ecologia de Ecossistemas Aquáticos (LEEA), Universidade Federal de Uberlândia (UFU), Avenida Amazonas, s / n., Uberlândia, MG, 38400 - 902, Brazil</mods:affiliation>
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<mods:namePart id="4D2C140316875381FD105BB4786B21D1">Anker, Arthur</mods:namePart>
<mods:affiliation id="AA9B1ACF6FC89D948688B186F3EEA232">Instituto de Ciências Biológicas, ICB- 5, Universidade Federal de Goiás (UFG), Campus Samambaia, Av. Esperança s / n, Goiânia, GO, 74690 - 900, Brazil</mods:affiliation>
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<treatment id="03BC87E6FF8FFF88C8D7FACBE537FCC8" LSID="urn:lsid:plazi:treatment:03BC87E6FF8FFF88C8D7FACBE537FCC8" httpUri="http://treatment.plazi.org/id/03BC87E6FF8FFF88C8D7FACBE537FCC8" lastPageId="21" lastPageNumber="76" pageId="20" pageNumber="75">
<subSubSection id="C30F657BFF8FFF89C8D7FACBE4DEFABF" box="[151,713,1330,1357]" pageId="20" pageNumber="75" type="nomenclature">
<paragraph id="8BAA36F0FF8FFF89C8D7FACBE4DEFABF" blockId="20.[151,713,1330,1392]" box="[151,713,1330,1357]" pageId="20" pageNumber="75">
<heading id="D0E2819CFF8FFF89C8D7FACBE4DEFABF" bold="true" box="[151,713,1330,1357]" fontSize="11" level="1" pageId="20" pageNumber="75" reason="1">
<taxonomicName id="4C154D73FF8FFF89C8D7FACBE4DEFABF" authority="Holthuis &amp; Maurin, 1952" authorityName="Holthuis &amp; Maurin" authorityYear="1952" box="[151,713,1330,1357]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="uncicornis">
<emphasis id="B961EAE2FF8FFF89C8D7FACBE4DEFABF" bold="true" box="[151,713,1330,1357]" pageId="20" pageNumber="75">
<emphasis id="B961EAE2FF8FFF89C8D7FACBE792FABF" bold="true" box="[151,389,1330,1357]" italics="true" pageId="20" pageNumber="75">Lysmata uncicornis</emphasis>
<bibRefCitation id="EF844B01FF8FFF89C9CCFACAE4DEFABF" author="Holthuis, L. B. &amp; Maurin, C." box="[396,713,1330,1356]" pageId="20" pageNumber="75" pagination="197 - 202" refId="ref23240" refString="Holthuis, L. B. &amp; Maurin, C. (1952) Note sur Lysmata uncicornis nov. spec. et sur deux autres especes interessantes de crustaces decapodes macroures de la cote Atlantique du Maroc. Proceedings of the Koninklijke Nederlandse Akademie van Weten- schappen, 55 (2), 197 - 202." type="journal article" year="1952">Holthuis &amp; Maurin, 1952</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F657BFF8FFF89C8D7FAAEE701FA83" box="[151,278,1366,1392]" pageId="20" pageNumber="75" type="description">
<paragraph id="8BAA36F0FF8FFF89C8D7FAAEE701FA83" blockId="20.[151,713,1330,1392]" box="[151,278,1366,1392]" pageId="20" pageNumber="75">
(
<figureCitation id="132E2A75FF8FFF89C8DFFAAEE719FA83" box="[159,270,1366,1392]" captionStart="FIGURE 14" captionStartId="21.[151,250,1946,1971]" captionTargetBox="[151,1436,849,1921]" captionTargetId="figure@21.[151,1436,849,1921]" captionTargetPageId="21" captionText="FIGURE 14. Lysmata uncicornis Holthuis &amp; Maurin, 1952(?): non-ov. specimen from Caraguatatuba, São Paulo, in dorsal view. Voucher specimen not found. Photograph: L. Carvalho." figureDoi="http://doi.org/10.5281/zenodo.3884713" httpUri="https://zenodo.org/record/3884713/files/figure.png" pageId="20" pageNumber="75">Figure 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F657BFF8FFF89C8D7FA63E5E9FA03" pageId="20" pageNumber="75" type="reference_group">
<paragraph id="8BAA36F0FF8FFF89C8D7FA63E75AFA22" blockId="20.[151,1437,1434,1520]" pageId="20" pageNumber="75">
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<treatmentCitation id="0AB410E1FF8FFF89C8D7FA63E481FA40" author="Holthuis, L. B. &amp; Maurin, C." box="[151,662,1434,1459]" page="198" pageId="20" pageNumber="75" year="1952">
<taxonomicName id="4C154D73FF8FFF89C8D7FA63E481FA40" authority="Holthuis &amp; Maurin, 1952: 198" authorityName="Holthuis &amp; Maurin" authorityPageNumber="198" authorityYear="1952" box="[151,662,1434,1459]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="uncicornis">
<emphasis id="B961EAE2FF8FFF89C8D7FA63E74AFA41" box="[151,349,1435,1458]" italics="true" pageId="20" pageNumber="75">Lysmata uncicornis</emphasis>
<bibRefCitation id="EF844B01FF8FFF89C922FA62E481FA40" author="Holthuis, L. B. &amp; Maurin, C." box="[354,662,1434,1459]" pageId="20" pageNumber="75" pagination="197 - 202" refId="ref23240" refString="Holthuis, L. B. &amp; Maurin, C. (1952) Note sur Lysmata uncicornis nov. spec. et sur deux autres especes interessantes de crustaces decapodes macroures de la cote Atlantique du Maroc. Proceedings of the Koninklijke Nederlandse Akademie van Weten- schappen, 55 (2), 197 - 202." type="journal article" year="1952">Holthuis &amp; Maurin, 1952: 198</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 1, 2;
<treatmentCitation id="0AB410E1FF8FFF89CB43FA62E5DCFA40" author="Lagardere, J. P." box="[771,971,1434,1459]" page="99" pageId="20" pageNumber="75" year="1971">
<bibRefCitation id="EF844B01FF8FFF89CB43FA62E5DCFA40" author="Lagardere, J. P." box="[771,971,1434,1459]" pageId="20" pageNumber="75" pagination="1 - 140" refId="ref23611" refString="Lagardere, J. P. (1971) Les crevettes des cotes du Maroc. Travaux de l'Institut Scientifique Cherifien, Serie Zoologie, 36, 1 - 140." type="journal article" year="1971">Lagardère 1971: 99</bibRefCitation>
</treatmentCitation>
, figs. 232235;
<treatmentCitation id="0AB410E1FF8FFF89CC2BFA62E6F4FA21" author="Gonzalez-Ortegon, E. &amp; Garcia-Raso, J. E. &amp; Lopez de la Rosa, I. &amp; Guerrero, M. &amp; Cuesta, J. A." page="26" pageId="20" pageNumber="75" year="2020">
<bibRefCitation id="EF844B01FF8FFF89CC2BFA62E6F4FA21" author="Gonzalez-Ortegon, E. &amp; Garcia-Raso, J. E. &amp; Lopez de la Rosa, I. &amp; Guerrero, M. &amp; Cuesta, J. A." pageId="20" pageNumber="75" pagination="26" refId="ref22953" refString="Gonzalez-Ortegon, E., Garcia-Raso, J. E., Lopez de la Rosa, I., Guerrero, M. &amp; Cuesta, J. A. (2020) Atlantic expansion of the African caridean shrimp Lysmata uncicornis Holthuis &amp; Maurin, 1952 (Caridea: Lysmatidae). Marine Biodiversity, 50, 26. https: // doi. org / 10.1007 / s 12526 - 020 - 01056 - w" type="journal article" year="2020">
González-Ortegón
<emphasis id="B961EAE2FF8FFF89CD6BFA63E349FA40" box="[1323,1374,1435,1459]" italics="true" pageId="20" pageNumber="75">et al.</emphasis>
2020: 26
</bibRefCitation>
</treatmentCitation>
, figs. 14.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA36F0FF8FFF89C8D7FA21E5E9FA03" blockId="20.[151,1437,1434,1520]" box="[151,1022,1495,1520]" pageId="20" pageNumber="75">
<treatmentCitationGroup id="AB0511DEFF8FFF89C8D7FA21E5E9FA03" box="[151,1022,1495,1520]" pageId="20" pageNumber="75">
<treatmentCitation id="0AB410E1FF8FFF89C8D7FA21E583FA03" author="Giraldes, B. W. &amp; Macedo, T. P. &amp; Brandao, M. C. &amp; Baeza, J. A. &amp; Freire, A. S." box="[151,916,1495,1520]" page="5" pageId="20" pageNumber="75" year="2018">
<taxonomicName id="4C154D73FF8FFF89C8D7FA21E583FA03" authority="Giraldes, Macedo, Brandao, Baeza &amp; Freire, 2018: 5" authorityName="Giraldes, Macedo, Brandao, Baeza &amp; Freire" authorityPageNumber="5" authorityYear="2018" box="[151,916,1495,1520]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="arvoredensis">
<emphasis id="B961EAE2FF8FFF89C8D7FA21E762FA03" box="[151,373,1496,1520]" italics="true" pageId="20" pageNumber="75">Lysmata arvoredensis</emphasis>
<bibRefCitation id="EF844B01FF8FFF89C93BFA2FE583FA03" author="Giraldes, B. W. &amp; Macedo, T. P. &amp; Brandao, M. C. &amp; Baeza, J. A. &amp; Freire, A. S." box="[379,916,1495,1520]" pageId="20" pageNumber="75" pagination="1 - 21" refId="ref22858" refString="Giraldes, B. W., Macedo, T. P., Brandao, M. C., Baeza, J. A. &amp; Freire, A. S. (2018) Lysmata arvoredensis nov. sp. a new species of shrimp from the south coast of Brazil with a key to species of Lysmata (Caridea: Lysmatidae) recorded in the southwestern Atlantic. PeerJ, 6, e 5561, 1 - 21. https: // doi. org / 10.7717 / peerj. 5561" type="journal article" year="2018">Giraldes, Macedo, Brandão, Baeza &amp; Freire, 2018: 5</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 14.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30F657BFF8FFF89C8D7F9E5E24DF9AE" pageId="20" pageNumber="75" type="materials_examined">
<paragraph id="8BAA36F0FF8FFF89C8D7F9E5E7DBF9CA" blockId="20.[151,1437,1565,2025]" box="[151,460,1565,1593]" pageId="20" pageNumber="75">
<emphasis id="B961EAE2FF8FFF89C8D7F9E5E796F9CB" bold="true" box="[151,385,1565,1592]" pageId="20" pageNumber="75">Material examined.</emphasis>
None.
</paragraph>
<paragraph id="8BAA36F0FF8FFF89C887F9B9E24DF9AE" blockId="20.[151,1437,1565,2025]" box="[199,1114,1601,1629]" pageId="20" pageNumber="75">
<emphasis id="B961EAE2FF8FFF89C887F9B9E7CFF9AF" bold="true" box="[199,472,1601,1628]" pageId="20" pageNumber="75">
First record for
<collectingCountry id="F3027660FF8FFF89C9C8F9B9E7C4F9AF" box="[392,467,1601,1628]" name="Brazil" pageId="20" pageNumber="75">Brazil</collectingCountry>
.
</emphasis>
Santa Catarina (
<bibRefCitation id="EF844B01FF8FFF89CAD1F9BAE564F9AF" author="Giraldes, B. W. &amp; Macedo, T. P. &amp; Brandao, M. C. &amp; Baeza, J. A. &amp; Freire, A. S." box="[657,883,1602,1628]" pageId="20" pageNumber="75" pagination="1 - 21" refId="ref22858" refString="Giraldes, B. W., Macedo, T. P., Brandao, M. C., Baeza, J. A. &amp; Freire, A. S. (2018) Lysmata arvoredensis nov. sp. a new species of shrimp from the south coast of Brazil with a key to species of Lysmata (Caridea: Lysmatidae) recorded in the southwestern Atlantic. PeerJ, 6, e 5561, 1 - 21. https: // doi. org / 10.7717 / peerj. 5561" type="journal article" year="2018">
Giraldes
<emphasis id="B961EAE2FF8FFF89CAB9F9BBE526F9AF" box="[761,817,1602,1628]" italics="true" pageId="20" pageNumber="75">et al.</emphasis>
2018
</bibRefCitation>
, as
<taxonomicName id="4C154D73FF8FFF89CBDCF9BBE25DF9AF" authorityName="Giraldes, Macedo, Brandao, Baeza &amp; Freire" authorityYear="2018" box="[924,1098,1602,1628]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="arvoredensis">
<emphasis id="B961EAE2FF8FFF89CBDCF9BBE25DF9AF" box="[924,1098,1602,1628]" italics="true" pageId="20" pageNumber="75">L. arvoredensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F657BFF8FFF89C887F99DE5EAF93A" pageId="20" pageNumber="75" type="distribution">
<paragraph id="8BAA36F0FF8FFF89C887F99DE5EAF93A" blockId="20.[151,1437,1565,2025]" pageId="20" pageNumber="75">
<emphasis id="B961EAE2FF8FFF89C887F99DE776F973" bold="true" box="[199,353,1637,1664]" pageId="20" pageNumber="75">Distribution.</emphasis>
Eastern Atlantic: Gulf of Cadiz (
<collectingCountry id="F3027660FF8FFF89CA9EF99EE528F973" box="[734,831,1638,1664]" name="Portugal" pageId="20" pageNumber="75">Portugal</collectingCountry>
and
<collectingCountry id="F3027660FF8FFF89CB39F99EE5AAF972" box="[889,957,1638,1665]" name="Spain" pageId="20" pageNumber="75">Spain</collectingCountry>
), Africa (
<collectingCountry id="F3027660FF8FFF89CC68F99EE298F973" box="[1064,1167,1638,1664]" name="Morocco" pageId="20" pageNumber="75">Morocco</collectingCountry>
to Congo) (
<bibRefCitation id="EF844B01FF8FFF89CD5BF99EE73AF957" author="Holthuis, L. B. &amp; Maurin, C." pageId="20" pageNumber="75" pagination="197 - 202" refId="ref23240" refString="Holthuis, L. B. &amp; Maurin, C. (1952) Note sur Lysmata uncicornis nov. spec. et sur deux autres especes interessantes de crustaces decapodes macroures de la cote Atlantique du Maroc. Proceedings of the Koninklijke Nederlandse Akademie van Weten- schappen, 55 (2), 197 - 202." type="journal article" year="1952">Holthuis &amp; Maurin 1952</bibRefCitation>
;
<bibRefCitation id="EF844B01FF8FFF89C97BF972E7E7F957" author="Lagardere, J. P." box="[315,496,1674,1700]" pageId="20" pageNumber="75" pagination="1 - 140" refId="ref23611" refString="Lagardere, J. P. (1971) Les crevettes des cotes du Maroc. Travaux de l'Institut Scientifique Cherifien, Serie Zoologie, 36, 1 - 140." type="journal article" year="1971">Lagardère 1971</bibRefCitation>
;
<bibRefCitation id="EF844B01FF8FFF89C9BDF972E544F957" author="Gonzalez-Ortegon, E. &amp; Garcia-Raso, J. E. &amp; Lopez de la Rosa, I. &amp; Guerrero, M. &amp; Cuesta, J. A." box="[509,851,1674,1700]" pageId="20" pageNumber="75" pagination="26" refId="ref22953" refString="Gonzalez-Ortegon, E., Garcia-Raso, J. E., Lopez de la Rosa, I., Guerrero, M. &amp; Cuesta, J. A. (2020) Atlantic expansion of the African caridean shrimp Lysmata uncicornis Holthuis &amp; Maurin, 1952 (Caridea: Lysmatidae). Marine Biodiversity, 50, 26. https: // doi. org / 10.1007 / s 12526 - 020 - 01056 - w" type="journal article" year="2020">
González-Ortegón
<emphasis id="B961EAE2FF8FFF89CA94F973E519F957" box="[724,782,1674,1700]" italics="true" pageId="20" pageNumber="75">et al.</emphasis>
2020
</bibRefCitation>
). Western Atlantic:
<collectingCountry id="F3027660FF8FFF89CC73F972E260F957" box="[1075,1143,1674,1700]" name="Brazil" pageId="20" pageNumber="75">Brazil</collectingCountry>
(Santa Catarina, possibly
<collectingRegion id="49D1F812FF8FFF89C8D7F956E71DF93A" box="[151,266,1710,1737]" country="Brazil" name="Sao Paulo" pageId="20" pageNumber="75">São Paulo</collectingRegion>
) (
<bibRefCitation id="EF844B01FF8FFF89C960F956E41FF93B" author="Giraldes, B. W. &amp; Macedo, T. P. &amp; Brandao, M. C. &amp; Baeza, J. A. &amp; Freire, A. S." box="[288,520,1710,1737]" pageId="20" pageNumber="75" pagination="1 - 21" refId="ref22858" refString="Giraldes, B. W., Macedo, T. P., Brandao, M. C., Baeza, J. A. &amp; Freire, A. S. (2018) Lysmata arvoredensis nov. sp. a new species of shrimp from the south coast of Brazil with a key to species of Lysmata (Caridea: Lysmatidae) recorded in the southwestern Atlantic. PeerJ, 6, e 5561, 1 - 21. https: // doi. org / 10.7717 / peerj. 5561" type="journal article" year="2018">
Giraldes
<emphasis id="B961EAE2FF8FFF89C9C7F957E7D7F93B" box="[391,448,1710,1736]" italics="true" pageId="20" pageNumber="75">et al.</emphasis>
, 2018
</bibRefCitation>
,
<emphasis id="B961EAE2FF8FFF89CA53F957E4F6F93B" box="[531,737,1710,1736]" italics="true" pageId="20" pageNumber="75">
as
<taxonomicName id="4C154D73FF8FFF89CA73F957E4F6F93B" authorityName="Giraldes, Macedo, Brandao, Baeza &amp; Freire" authorityYear="2018" box="[563,737,1710,1736]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="arvoredensis">L. arvoredensis</taxonomicName>
</emphasis>
; present study) (
<figureCitation id="132E2A75FF8FFF89CBDCF956E5E7F93A" box="[924,1008,1710,1737]" captionStart="FIGURE 16" captionStartId="24.[151,250,1865,1890]" captionTargetBox="[151,1436,488,1816]" captionTargetId="figure@24.[151,1436,488,1840]" captionTargetPageId="24" captionText="FIGURE 16. Distribution map for 11 species of Lysmata Risso, 1816 currently known from the Brazilian coast and offshore islands. Abbreviations: Alagoas (AL), Amapá (AP), Atol das Rocas (AR), Bahia (BA), Ceará (CE), Espírito Santo (ES), Maranhão (MA), Pará (PA), Paraíba (PB), Paraná (PR), Pernambuco (PE), Piauí (PI), Rio Grande do Norte (RN), Rio Grande do Sul (RS), Rio de Janeiro (RJ), Santa Catarina (SC), São Paulo (SP), São Pedro e São Paulo (SPSP), Sergipe (SE), Trindade e Martim Vaz (TMV)." figureDoi="http://doi.org/10.5281/zenodo.3884717" httpUri="https://zenodo.org/record/3884717/files/figure.png" pageId="20" pageNumber="75">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F657BFF8FFF89C887F92AE795F8C7" pageId="20" pageNumber="75" type="biology_ecology">
<paragraph id="8BAA36F0FF8FFF89C887F92AE795F8C7" blockId="20.[151,1437,1565,2025]" pageId="20" pageNumber="75">
<emphasis id="B961EAE2FF8FFF89C887F92AE731F91F" bold="true" box="[199,294,1746,1772]" pageId="20" pageNumber="75">Ecology</emphasis>
.
<collectingCountry id="F3027660FF8FFF89C975F92AE769F91F" box="[309,382,1746,1772]" name="Brazil" pageId="20" pageNumber="75">Brazil</collectingCountry>
: rocky bottoms and possibly in mussel ropes, depth: around
<quantity id="4CED9B15FF8FFF89CC68F92AE275F91E" box="[1064,1122,1746,1773]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="75" unit="m" value="20.0">20 m</quantity>
(
<bibRefCitation id="EF844B01FF8FFF89CC30F92AE34CF91F" author="Giraldes, B. W. &amp; Macedo, T. P. &amp; Brandao, M. C. &amp; Baeza, J. A. &amp; Freire, A. S." box="[1136,1371,1746,1773]" pageId="20" pageNumber="75" pagination="1 - 21" refId="ref22858" refString="Giraldes, B. W., Macedo, T. P., Brandao, M. C., Baeza, J. A. &amp; Freire, A. S. (2018) Lysmata arvoredensis nov. sp. a new species of shrimp from the south coast of Brazil with a key to species of Lysmata (Caridea: Lysmatidae) recorded in the southwestern Atlantic. PeerJ, 6, e 5561, 1 - 21. https: // doi. org / 10.7717 / peerj. 5561" type="journal article" year="2018">
Giraldes
<emphasis id="B961EAE2FF8FFF89CC99F92BE305F91F" box="[1241,1298,1746,1772]" italics="true" pageId="20" pageNumber="75">et al.</emphasis>
, 2018
</bibRefCitation>
, as
<taxonomicName id="4C154D73FF8FFF89CDC6F92BE730F8E3" authorityName="Giraldes, Macedo, Brandao, Baeza &amp; Freire" authorityYear="2018" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="arvoredensis">
<emphasis id="B961EAE2FF8FFF89CDC6F92BE730F8E3" italics="true" pageId="20" pageNumber="75">L. arvoredensis</emphasis>
</taxonomicName>
); eastern Atlantic: subtidal rocky bottoms (
<bibRefCitation id="EF844B01FF8FFF89CB45F90EE5A0F8E3" author="Lagardere, J. P." box="[773,951,1782,1808]" pageId="20" pageNumber="75" pagination="1 - 140" refId="ref23611" refString="Lagardere, J. P. (1971) Les crevettes des cotes du Maroc. Travaux de l'Institut Scientifique Cherifien, Serie Zoologie, 36, 1 - 140." type="journal article" year="1971">Lagardère 1971</bibRefCitation>
) or “detritic bottom with rocks” (González- Ortegón
<emphasis id="B961EAE2FF8FFF89C8BAF8E3E725F8C7" box="[250,306,1818,1844]" italics="true" pageId="20" pageNumber="75">et al.</emphasis>
2020).
</paragraph>
</subSubSection>
<subSubSection id="C30F657BFF8FFF88C887F8C6E537FCC8" lastPageId="21" lastPageNumber="76" pageId="20" pageNumber="75" type="discussion">
<paragraph id="8BAA36F0FF8FFF88C887F8C6E264FEEC" blockId="20.[151,1437,1565,2025]" lastBlockId="21.[151,1437,153,827]" lastPageId="21" lastPageNumber="76" pageId="20" pageNumber="75">
<emphasis id="B961EAE2FF8FFF89C887F8C6E72CF8AB" bold="true" box="[199,315,1854,1880]" pageId="20" pageNumber="75">Remarks.</emphasis>
<taxonomicName id="4C154D73FF8FFF89C901F8C7E40BF8AB" authorityName="Holthuis &amp; Maurin" authorityYear="1952" box="[321,540,1855,1880]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="uncicornis">
<emphasis id="B961EAE2FF8FFF89C901F8C7E40BF8AB" box="[321,540,1855,1880]" italics="true" pageId="20" pageNumber="75">Lysmata uncicornis</emphasis>
</taxonomicName>
, despite having a distinctive colour pattern, can be easily confused with
<taxonomicName id="4C154D73FF8FFF89CD00F8C7E38BF8AB" authorityName="in Rhyne &amp; Lin" authorityYear="2006" box="[1344,1436,1854,1880]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="bahia">
<emphasis id="B961EAE2FF8FFF89CD00F8C7E38BF8AB" box="[1344,1436,1854,1880]" italics="true" pageId="20" pageNumber="75">L. bahia</emphasis>
</taxonomicName>
due to similarities in some important taxonomic characters, such as the rounded pterygostomian angle, stylocerite length, shape and armature of the rostrum, and relative length between scaphocerite and antennular peduncle (
<figureCitation id="132E2A75FF8FFF89CD24F87EE6D4F837" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="6.[151,250,1968,1993]" captionStartId-1="7.[151,250,1875,1900]" captionTargetBox-0="[151,1436,525,1943]" captionTargetBox-1="[199,1386,189,1841]" captionTargetId-0="figure@6.[151,1436,525,1943]" captionTargetId-1="figure@7.[194,1394,181,1850]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 3. Lysmata bahia Rhyne &amp; Lin, 2006: non-ov. specimen from Bahia (pocl 11.6 mm; MZUSP 36972) in dorsal (A) and lateral (B) views. Scale bars: 10 mm. Photographs: P. Pachelle." captionText-1="FIGURE 4. Lysmata bahia Rhyne &amp; Lin, 2006, non-ov. specimen from Sergipe (pocl 12.0 mm; MZUSP 37511): (A) frontal margin and cephalic appendages, lateral view; (B) infraorbital region, lateral view; (C) same, dorsolateral view; (D) left third maxilliped, antepenultimate and penultimate articles, ventromesial view; (E) left antennule, third article of antennular peduncle and flagella, mesial view; (F) same, bifurcation between upper flagellum and accessory ramus, mesial view; (G) right first pereopod, merus to propodus, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.3884691" figureDoi-1="http://doi.org/10.5281/zenodo.3884693" httpUri-0="https://zenodo.org/record/3884691/files/figure.png" httpUri-1="https://zenodo.org/record/3884693/files/figure.png" pageId="20" pageNumber="75">Figs. 34</figureCitation>
,
<figureCitation id="132E2A75FF8FFF89C890F852E6F8F837" box="[208,239,1962,1988]" captionStart="FIGURE 14" captionStartId="21.[151,250,1946,1971]" captionTargetBox="[151,1436,849,1921]" captionTargetId="figure@21.[151,1436,849,1921]" captionTargetPageId="21" captionText="FIGURE 14. Lysmata uncicornis Holthuis &amp; Maurin, 1952(?): non-ov. specimen from Caraguatatuba, São Paulo, in dorsal view. Voucher specimen not found. Photograph: L. Carvalho." figureDoi="http://doi.org/10.5281/zenodo.3884713" httpUri="https://zenodo.org/record/3884713/files/figure.png" pageId="20" pageNumber="75">14</figureCitation>
; see also
<bibRefCitation id="EF844B01FF8FFF89C91CF852E428F837" author="Giraldes, B. W. &amp; Macedo, T. P. &amp; Brandao, M. C. &amp; Baeza, J. A. &amp; Freire, A. S." box="[348,575,1962,1988]" pageId="20" pageNumber="75" pagination="1 - 21" refId="ref22858" refString="Giraldes, B. W., Macedo, T. P., Brandao, M. C., Baeza, J. A. &amp; Freire, A. S. (2018) Lysmata arvoredensis nov. sp. a new species of shrimp from the south coast of Brazil with a key to species of Lysmata (Caridea: Lysmatidae) recorded in the southwestern Atlantic. PeerJ, 6, e 5561, 1 - 21. https: // doi. org / 10.7717 / peerj. 5561" type="journal article" year="2018">
Giraldes
<emphasis id="B961EAE2FF8FFF89C982F853E7ECF837" box="[450,507,1962,1988]" italics="true" pageId="20" pageNumber="75">et al.</emphasis>
2018
</bibRefCitation>
: figs. 1, 2, as
<taxonomicName id="4C154D73FF8FFF89CA9CF853E59DF837" authorityName="Giraldes, Macedo, Brandao, Baeza &amp; Freire" authorityYear="2018" box="[732,906,1962,1988]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="arvoredensis">
<emphasis id="B961EAE2FF8FFF89CA9CF853E59DF837" box="[732,906,1962,1988]" italics="true" pageId="20" pageNumber="75">L. arvoredensis</emphasis>
</taxonomicName>
). However, these two species can be separated by the relative length of the first pereopod carpus (about three times as long as high in
<taxonomicName id="4C154D73FF8FFF89CC3DF837E375F81B" box="[1149,1378,1999,2024]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="75" phylum="Arthropoda" rank="subSpecies" species="uncicornis" subSpecies="versus">
<emphasis id="B961EAE2FF8FFF89CC3DF837E375F81B" box="[1149,1378,1999,2024]" italics="true" pageId="20" pageNumber="75">L. uncicornis versus</emphasis>
</taxonomicName>
over four times as long as high in
<taxonomicName id="4C154D73FF8EFF88C99EFF62E42BFF40" authorityName="in Rhyne &amp; Lin" authorityYear="2006" box="[478,572,153,179]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="bahia">
<emphasis id="B961EAE2FF8EFF88C99EFF62E42BFF40" box="[478,572,153,179]" italics="true" pageId="21" pageNumber="76">L. bahia</emphasis>
</taxonomicName>
) and the number of subdivisions in the second pereopod carpus (
<quantity id="4CED9B15FF8EFF88CD59FF61E369FF40" box="[1305,1406,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.096" metricValueMin="5.588" pageId="21" pageNumber="76" unit="in" value="23.0" valueMax="24.0" valueMin="22.0">2224 in</quantity>
<taxonomicName id="4C154D73FF8EFF88CDC6FF62E74AFF24" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="subSpecies" species="uncicornis" subSpecies="versus">
<emphasis id="B961EAE2FF8EFF88CDC6FF62E74AFF24" italics="true" pageId="21" pageNumber="76">L. uncicornis versus</emphasis>
</taxonomicName>
<quantity id="4CED9B15FF8EFF88C924FF45E7DFFF24" box="[356,456,189,215]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" metricValueMax="7.874" metricValueMin="7.366" pageId="21" pageNumber="76" unit="in" value="30.0" valueMax="31.0" valueMin="29.0">2931 in</quantity>
<taxonomicName id="4C154D73FF8EFF88C990FF46E439FF24" authorityName="in Rhyne &amp; Lin" authorityYear="2006" box="[464,558,189,215]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="bahia">
<emphasis id="B961EAE2FF8EFF88C990FF46E439FF24" box="[464,558,189,215]" italics="true" pageId="21" pageNumber="76">L. bahia</emphasis>
</taxonomicName>
). In addition, the free portion of the accessory ramus of the lateral antennular flagellum is short in
<taxonomicName id="4C154D73FF8EFF88C9C3FF1AE400FF08" authorityName="Holthuis &amp; Maurin" authorityYear="1952" box="[387,535,226,251]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="uncicornis">
<emphasis id="B961EAE2FF8EFF88C9C3FF1AE400FF08" box="[387,535,226,251]" italics="true" pageId="21" pageNumber="76">L. uncicornis</emphasis>
</taxonomicName>
and noticeably longer in
<taxonomicName id="4C154D73FF8EFF88CB79FF1AE58FFF08" authorityName="in Rhyne &amp; Lin" authorityYear="2006" box="[825,920,225,251]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="bahia">
<emphasis id="B961EAE2FF8EFF88CB79FF1AE58FFF08" box="[825,920,225,251]" italics="true" pageId="21" pageNumber="76">L. bahia</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A75FF8EFF88CBE8FF19E209FF08" box="[936,1054,225,251]" captionStart="FIGURE 4" captionStartId="7.[151,250,1875,1900]" captionTargetBox="[199,1386,189,1841]" captionTargetId="figure@7.[194,1394,181,1850]" captionTargetPageId="7" captionText="FIGURE 4. Lysmata bahia Rhyne &amp; Lin, 2006, non-ov. specimen from Sergipe (pocl 12.0 mm; MZUSP 37511): (A) frontal margin and cephalic appendages, lateral view; (B) infraorbital region, lateral view; (C) same, dorsolateral view; (D) left third maxilliped, antepenultimate and penultimate articles, ventromesial view; (E) left antennule, third article of antennular peduncle and flagella, mesial view; (F) same, bifurcation between upper flagellum and accessory ramus, mesial view; (G) right first pereopod, merus to propodus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3884693" httpUri="https://zenodo.org/record/3884693/files/figure.png" pageId="21" pageNumber="76">Fig. 4E, F</figureCitation>
;
<bibRefCitation id="EF844B01FF8EFF88CC6AFF19E319FF08" author="Giraldes, B. W. &amp; Macedo, T. P. &amp; Brandao, M. C. &amp; Baeza, J. A. &amp; Freire, A. S." box="[1066,1294,225,251]" pageId="21" pageNumber="76" pagination="1 - 21" refId="ref22858" refString="Giraldes, B. W., Macedo, T. P., Brandao, M. C., Baeza, J. A. &amp; Freire, A. S. (2018) Lysmata arvoredensis nov. sp. a new species of shrimp from the south coast of Brazil with a key to species of Lysmata (Caridea: Lysmatidae) recorded in the southwestern Atlantic. PeerJ, 6, e 5561, 1 - 21. https: // doi. org / 10.7717 / peerj. 5561" type="journal article" year="2018">
Giraldes
<emphasis id="B961EAE2FF8EFF88CCD1FF1AE2DDFF08" box="[1169,1226,225,251]" italics="true" pageId="21" pageNumber="76">et al.</emphasis>
2018
</bibRefCitation>
: fig. 2E), although the validity of character needs to be confirmed by examination of more material.
</paragraph>
<paragraph id="8BAA36F0FF8EFF88C887FED2E209FD74" blockId="21.[151,1437,153,827]" pageId="21" pageNumber="76">
<taxonomicName id="4C154D73FF8EFF88C887FED2E7B5FEB0" authorityName="Holthuis &amp; Maurin" authorityYear="1952" box="[199,418,298,323]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="uncicornis">
<emphasis id="B961EAE2FF8EFF88C887FED2E7B5FEB0" box="[199,418,298,323]" italics="true" pageId="21" pageNumber="76">Lysmata uncicornis</emphasis>
</taxonomicName>
is also morphologically close to
<taxonomicName id="4C154D73FF8EFF88CB50FED2E277FEB0" authority="Baeza &amp; Anker, 2008" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[784,1120,297,323]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B961EAE2FF8EFF88CB50FED2E57DFEB0" box="[784,874,297,323]" italics="true" pageId="21" pageNumber="76">L. hochi</emphasis>
<bibRefCitation id="EF844B01FF8EFF88CB31FED1E277FEB0" author="Baeza, J. A. &amp; Anker, A." box="[881,1120,297,323]" pageId="21" pageNumber="76" pagination="148 - 155" refId="ref21069" refString="Baeza, J. A. &amp; Anker, A. (2008) Lysmata hochi n. sp., a new hermaphroditic shrimp from the southwestern Caribbean Sea (Caridea: Hippolytidae). Journal of Crustacean Biology, 28 (1), 148 - 155. https: // doi. org / 10.1651 / 07 - 2839 R. 1" type="journal article" year="2008">Baeza &amp; Anker, 2008</bibRefCitation>
</taxonomicName>
from the Caribbean Sea and Florida (
<bibRefCitation id="EF844B01FF8EFF88C8B6FEB5E7F2FE94" author="Baeza, J. A. &amp; Anker, A." box="[246,485,333,359]" pageId="21" pageNumber="76" pagination="148 - 155" refId="ref21069" refString="Baeza, J. A. &amp; Anker, A. (2008) Lysmata hochi n. sp., a new hermaphroditic shrimp from the southwestern Caribbean Sea (Caridea: Hippolytidae). Journal of Crustacean Biology, 28 (1), 148 - 155. https: // doi. org / 10.1651 / 07 - 2839 R. 1" type="journal article" year="2008">Baeza &amp; Anker 2008</bibRefCitation>
; Baeza 2009), although phylogenetically these two species are not closely related (cf.
<bibRefCitation id="EF844B01FF8EFF88C8D7FE89E763FE78" author="Giraldes, B. W. &amp; Macedo, T. P. &amp; Brandao, M. C. &amp; Baeza, J. A. &amp; Freire, A. S." box="[151,372,369,395]" pageId="21" pageNumber="76" pagination="1 - 21" refId="ref22858" refString="Giraldes, B. W., Macedo, T. P., Brandao, M. C., Baeza, J. A. &amp; Freire, A. S. (2018) Lysmata arvoredensis nov. sp. a new species of shrimp from the south coast of Brazil with a key to species of Lysmata (Caridea: Lysmatidae) recorded in the southwestern Atlantic. PeerJ, 6, e 5561, 1 - 21. https: // doi. org / 10.7717 / peerj. 5561" type="journal article" year="2018">
Giraldes
<emphasis id="B961EAE2FF8EFF88C8BCFE8AE724FE78" box="[252,307,369,395]" italics="true" pageId="21" pageNumber="76">et al.</emphasis>
2018
</bibRefCitation>
: fig. 5). As pointed out by
<bibRefCitation id="EF844B01FF8EFF88CAD7FE89E5E2FE78" author="Gonzalez-Ortegon, E. &amp; Garcia-Raso, J. E. &amp; Lopez de la Rosa, I. &amp; Guerrero, M. &amp; Cuesta, J. A." box="[663,1013,369,395]" pageId="21" pageNumber="76" pagination="26" refId="ref22953" refString="Gonzalez-Ortegon, E., Garcia-Raso, J. E., Lopez de la Rosa, I., Guerrero, M. &amp; Cuesta, J. A. (2020) Atlantic expansion of the African caridean shrimp Lysmata uncicornis Holthuis &amp; Maurin, 1952 (Caridea: Lysmatidae). Marine Biodiversity, 50, 26. https: // doi. org / 10.1007 / s 12526 - 020 - 01056 - w" type="journal article" year="2020">
González-Ortegón
<emphasis id="B961EAE2FF8EFF88CB2BFE8AE5B4FE78" box="[875,931,369,395]" italics="true" pageId="21" pageNumber="76">et al.</emphasis>
(2020)
</bibRefCitation>
, some of the characters used by
<bibRefCitation id="EF844B01FF8EFF88CD18FE89E75AFE5C" author="Baeza, J. A. &amp; Anker, A." pageId="21" pageNumber="76" pagination="148 - 155" refId="ref21069" refString="Baeza, J. A. &amp; Anker, A. (2008) Lysmata hochi n. sp., a new hermaphroditic shrimp from the southwestern Caribbean Sea (Caridea: Hippolytidae). Journal of Crustacean Biology, 28 (1), 148 - 155. https: // doi. org / 10.1651 / 07 - 2839 R. 1" type="journal article" year="2008">Baeza &amp; Anker (2008)</bibRefCitation>
to separate
<taxonomicName id="4C154D73FF8EFF88C998FE6EE422FE5C" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[472,565,405,431]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B961EAE2FF8EFF88C998FE6EE422FE5C" box="[472,565,405,431]" italics="true" pageId="21" pageNumber="76">L. hochi</emphasis>
</taxonomicName>
from
<taxonomicName id="4C154D73FF8EFF88CA3DFE6EE505FE5C" authorityName="Holthuis &amp; Maurin" authorityYear="1952" box="[637,786,406,431]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="uncicornis">
<emphasis id="B961EAE2FF8EFF88CA3DFE6EE505FE5C" box="[637,786,406,431]" italics="true" pageId="21" pageNumber="76">L. uncicornis</emphasis>
</taxonomicName>
may not be valid due to the intraspecific variation in the latter species. However, the two species can still be separated by some details in their colour patterns, for instance, by the width of the transverse bands on the pleon, the disposition and shape of streaks and bands on the carapace and the presence/absence of white patches on the pleonites (cf.
<bibRefCitation id="EF844B01FF8EFF88CB20FDF9E247FDE8" author="Baeza, J. A." box="[864,1104,513,539]" pageId="21" pageNumber="76" pagination="269 - 310" refId="ref20970" refString="Baeza, J. A. (2018) Sexual systems in shrimps (infraorder Caridea Dana, 1852), with special reference to the historical origin and adaptive value of protandric simultaneous hermaphroditism. In: Leonard, J. L. (Ed.), Transitions between sexual systems. Understanding the mechanisms of, and pathways between, dioecy, hermaphroditism and other sexual systems. Springer, Cham, 269 - 310. https: // doi. org / 10.1007 / 978 - 3 - 319 - 94139 - 4 _ 10" type="book chapter" year="2018">Baeza &amp; Anker 2018</bibRefCitation>
: fig. 2 and
<bibRefCitation id="EF844B01FF8EFF88CC8DFDF9E704FDCC" author="Gonzalez-Ortegon, E. &amp; Garcia-Raso, J. E. &amp; Lopez de la Rosa, I. &amp; Guerrero, M. &amp; Cuesta, J. A." pageId="21" pageNumber="76" pagination="26" refId="ref22953" refString="Gonzalez-Ortegon, E., Garcia-Raso, J. E., Lopez de la Rosa, I., Guerrero, M. &amp; Cuesta, J. A. (2020) Atlantic expansion of the African caridean shrimp Lysmata uncicornis Holthuis &amp; Maurin, 1952 (Caridea: Lysmatidae). Marine Biodiversity, 50, 26. https: // doi. org / 10.1007 / s 12526 - 020 - 01056 - w" type="journal article" year="2020">
González-Ortegón
<emphasis id="B961EAE2FF8EFF88C8D7FDDEE6C7FDCC" box="[151,208,549,575]" italics="true" pageId="21" pageNumber="76">et al.</emphasis>
2020
</bibRefCitation>
: fig. 3a, b). Noteworthy, the recent record of
<taxonomicName id="4C154D73FF8EFF88CB50FDDEE57BFDCC" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[784,876,549,575]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B961EAE2FF8EFF88CB50FDDEE57BFDCC" box="[784,876,549,575]" italics="true" pageId="21" pageNumber="76">L. hochi</emphasis>
</taxonomicName>
from the Indian Ocean (
<bibRefCitation id="EF844B01FF8EFF88CCC1FDDDE323FDCC" author="Jose, S. &amp; Purushothaman, P. &amp; M. Madhavan &amp; Akash, S. &amp; Bharathi, S. &amp; Dhina-Karan, A. &amp; Kumar, T. T. A. &amp; Lal, K. K." box="[1153,1332,549,575]" pageId="21" pageNumber="76" pagination="353 - 364" refId="ref23340" refString="Jose, S., Purushothaman, P., M. Madhavan, Akash, S., Bharathi, S., Dhina-Karan, A., Kumar, T. T. A. &amp; Lal, K. K. (2020) Two new records of hippolytoid shrimps, Lysmata hochi Baeza &amp; Anker, 2008 (Decapoda: Lysmatidae) and Lysmata amboinensis (de Man, 1888) from Lakshadweep Islands, India with taxonomic notes. Zootaxa, 4755 (2), 353 - 364. https: // doi. org / 10.11646 / zootaxa. 4755.2.9" type="journal article" year="2020">
Jose
<emphasis id="B961EAE2FF8EFF88CCFAFDDEE2E5FDCC" box="[1210,1266,549,575]" italics="true" pageId="21" pageNumber="76">et al.</emphasis>
2020
</bibRefCitation>
) is likely erroneous since the colour pattern of the
<collectingCountry id="F3027660FF8EFF88CA1EFDB1E4EFFD90" box="[606,760,585,611]" name="India" pageId="21" pageNumber="76">Lakshadweep</collectingCountry>
specimens does not match that of
<taxonomicName id="4C154D73FF8EFF88CC37FDB2E2C5FD90" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[1143,1234,585,611]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B961EAE2FF8EFF88CC37FDB2E2C5FD90" box="[1143,1234,585,611]" italics="true" pageId="21" pageNumber="76">L. hochi</emphasis>
</taxonomicName>
; this record could refer to the closely related
<taxonomicName id="4C154D73FF8EFF88C982FD96E50BFD74" authority="(De Man, 1902)" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[450,796,621,647]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="kuekenthali">
<emphasis id="B961EAE2FF8EFF88C982FD96E476FD74" box="[450,609,621,647]" italics="true" pageId="21" pageNumber="76">L. kuekenthali</emphasis>
(
<bibRefCitation id="EF844B01FF8EFF88CA30FD95E504FD74" author="De Man, J. G." box="[624,787,621,647]" pageId="21" pageNumber="76" pagination="466 - 929" refId="ref22525" refString="De Man, J. G. (1902) Die von Herrn Professor Kukenthal im Indischen Archipel gesammelten Dekapoden und Stomatopoden. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 25, 466 - 929. https: // doi. org / 10.5962 / bhl. title. 10409" type="journal article" year="1902">De Man, 1902</bibRefCitation>
)
</taxonomicName>
or yet another species.
</paragraph>
<paragraph id="8BAA36F0FF8EFF88C887FD69E537FCC8" blockId="21.[151,1437,153,827]" pageId="21" pageNumber="76">
A photograph of a shrimp extracted from a mussel rope in Caraguatatuba (
<figureCitation id="132E2A75FF8EFF88CC6AFD69E297FD58" box="[1066,1152,657,683]" captionStart="FIGURE 14" captionStartId="21.[151,250,1946,1971]" captionTargetBox="[151,1436,849,1921]" captionTargetId="figure@21.[151,1436,849,1921]" captionTargetPageId="21" captionText="FIGURE 14. Lysmata uncicornis Holthuis &amp; Maurin, 1952(?): non-ov. specimen from Caraguatatuba, São Paulo, in dorsal view. Voucher specimen not found. Photograph: L. Carvalho." figureDoi="http://doi.org/10.5281/zenodo.3884713" httpUri="https://zenodo.org/record/3884713/files/figure.png" pageId="21" pageNumber="76">Fig. 14</figureCitation>
), taken by one of us (L. Carvalho) in 2012, suggests the presence of
<taxonomicName id="4C154D73FF8EFF88CAD2FD4EE530FD3C" authorityName="Holthuis &amp; Maurin" authorityYear="1952" box="[658,807,694,719]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="uncicornis">
<emphasis id="B961EAE2FF8EFF88CAD2FD4EE530FD3C" box="[658,807,694,719]" italics="true" pageId="21" pageNumber="76">L. uncicornis</emphasis>
</taxonomicName>
also in the state of
<collectingRegion id="49D1F812FF8EFF88CC4BFD4DE297FD3C" box="[1035,1152,693,719]" country="Brazil" name="Sao Paulo" pageId="21" pageNumber="76">São Paulo</collectingRegion>
. This individual presents a colour pattern that in all aspects is very similar to that of
<taxonomicName id="4C154D73FF8EFF88CB77FD22E2C8FD00" authority="in Giraldes et al. (2018" authorityName="in Giraldes" authorityYear="2018" box="[823,1247,729,755]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="uncicornis">
<emphasis id="B961EAE2FF8EFF88CB77FD22E5DBFD00" box="[823,972,730,755]" italics="true" pageId="21" pageNumber="76">L. uncicornis</emphasis>
in
<bibRefCitation id="EF844B01FF8EFF88CBB2FD21E2C8FD00" author="Giraldes, B. W. &amp; Macedo, T. P. &amp; Brandao, M. C. &amp; Baeza, J. A. &amp; Freire, A. S." box="[1010,1247,729,755]" pageId="21" pageNumber="76" pagination="1 - 21" refId="ref22858" refString="Giraldes, B. W., Macedo, T. P., Brandao, M. C., Baeza, J. A. &amp; Freire, A. S. (2018) Lysmata arvoredensis nov. sp. a new species of shrimp from the south coast of Brazil with a key to species of Lysmata (Caridea: Lysmatidae) recorded in the southwestern Atlantic. PeerJ, 6, e 5561, 1 - 21. https: // doi. org / 10.7717 / peerj. 5561" type="journal article" year="2018">
Giraldes
<emphasis id="B961EAE2FF8EFF88CC19FD22E285FD00" box="[1113,1170,729,755]" italics="true" pageId="21" pageNumber="76">et al.</emphasis>
(2018
</bibRefCitation>
</taxonomicName>
: figs. 1, 4F as
<taxonomicName id="4C154D73FF8EFF88CDC6FD22E730FCE4" authorityName="Giraldes, Macedo, Brandao, Baeza &amp; Freire" authorityYear="2018" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="arvoredensis">
<emphasis id="B961EAE2FF8EFF88CDC6FD22E730FCE4" italics="true" pageId="21" pageNumber="76">L. arvoredensis</emphasis>
</taxonomicName>
) and
<bibRefCitation id="EF844B01FF8EFF88C928FD05E4D2FCE4" author="Gonzalez-Ortegon, E. &amp; Garcia-Raso, J. E. &amp; Lopez de la Rosa, I. &amp; Guerrero, M. &amp; Cuesta, J. A." box="[360,709,765,791]" pageId="21" pageNumber="76" pagination="26" refId="ref22953" refString="Gonzalez-Ortegon, E., Garcia-Raso, J. E., Lopez de la Rosa, I., Guerrero, M. &amp; Cuesta, J. A. (2020) Atlantic expansion of the African caridean shrimp Lysmata uncicornis Holthuis &amp; Maurin, 1952 (Caridea: Lysmatidae). Marine Biodiversity, 50, 26. https: // doi. org / 10.1007 / s 12526 - 020 - 01056 - w" type="journal article" year="2020">
González-Ortegón
<emphasis id="B961EAE2FF8EFF88CA7EFD06E460FCE4" box="[574,631,765,791]" italics="true" pageId="21" pageNumber="76">et al.</emphasis>
(2020
</bibRefCitation>
: fig, 3a, b). Unfortunately, the photographed specimen could not be traced and therefore its true identity remains unknown.
</paragraph>
</subSubSection>
</treatment>
</document>