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<document id="F5B0F629F68D28FE6F0D8AF11CFECC29" ID-DOI="10.11646/zootaxa.2780.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5290601" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629713954357" checkinUser="felipe" docAuthor="Reboleira, A. S. P. S., Gonçalves, F. &amp; Oromí, P." docDate="2011" docId="03BC87DD4333FF83FF6FA6A0FD4AB994" docLanguage="en" docName="zt02780p056.pdf" docOrigin="Zootaxa 2780 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2780.1.5" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Domene (Lathromene) lusitanica Reboleira &amp; Gonçalves &amp; Oromí 2011, new species" docType="treatment" docUuid="B7E260ED-670A-4DC6-AF02-8FD7D7A7ADD7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="52" masterDocId="FF85FFA54331FF87FFF8A42FFF8ABF61" masterDocTitle="On the Iberian endemic subgenus Lathromene Koch (Coleoptera: Staphylinidae Paederinae): description of the first hypogean Domene Fauvel, 1872 from Portugal" masterLastPageNumber="56" masterPageNumber="48" pageNumber="50" updateTime="1699100473795" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="C0AFB488486A8D1260EE17667ACAF9AC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D3941D89C22AB5B03CBF9843CD7712C9">
<mods:title id="254111E70E023B281B384659A43B8687">On the Iberian endemic subgenus Lathromene Koch (Coleoptera: Staphylinidae Paederinae): description of the first hypogean Domene Fauvel, 1872 from Portugal</mods:title>
</mods:titleInfo>
<mods:name id="5B2B3B5903FCD113B387A11A06669AFB" type="personal">
<mods:role id="2EDBFE7595ACE3E471B462C228B1151D">
<mods:roleTerm id="CEE2088B42A7C7D043BD422C52CDF634">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2CAADA2CEB8EDFFC1BD58ED7F3B30F4D">Reboleira, A. S. P. S.</mods:namePart>
<mods:affiliation id="D09DAA11614EF4879004BD0D41E83874">CESAM - Centro de Estudos do Ambiente e do Mar &amp; Departamento de Biologia, Universidade de Aveiro, Campus Universitário de Santiago, 3810 - Aveiro, Portugal &amp; Departamento de Biología Animal, Universidad de La Laguna, 38206 La Laguna, Spain. E-mail: poromi @ ull. es</mods:affiliation>
<mods:nameIdentifier id="B577F26E67656E501A8D124F72333DA1" type="email">sreboleira@ua.pt</mods:nameIdentifier>
</mods:name>
<mods:name id="3AC378E2215A7775C1C0C459A47A1F83" type="personal">
<mods:role id="75DAA7A20CA00FEB591A077123C91AA9">
<mods:roleTerm id="07E09C80B9A8FDBC20FBBAD67BB003AB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="269868EE6FD520C145359C7C3282EB55">Gonçalves, F.</mods:namePart>
<mods:affiliation id="B5069C28C98619E39E372F6EAD1EA9F7">CESAM - Centro de Estudos do Ambiente e do Mar &amp; Departamento de Biologia, Universidade de Aveiro, Campus Universitário de Santiago, 3810 - Aveiro, Portugal</mods:affiliation>
</mods:name>
<mods:name id="203E57161E87663A23BFCA7BDE48E557" type="personal">
<mods:role id="33DAD7A24481069FB32C9714E8CB60F2">
<mods:roleTerm id="40E89F82C6438C7826E6A276C8D6EC82">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C15A18BC340AECE03F88858A6E5510F1">Oromí, P.</mods:namePart>
<mods:affiliation id="1940E066980BCB6A8AB092116F43B037">Departamento de Biología Animal, Universidad de La Laguna, 38206 La Laguna, Spain. E-mail: poromi @ ull. es</mods:affiliation>
</mods:name>
<mods:typeOfResource id="3655AE5E6BF10624469CE60F9E2328D5">text</mods:typeOfResource>
<mods:relatedItem id="95B1270AA51B2C9C3D0940845E2631CC" type="host">
<mods:titleInfo id="A698C24D8EB23407B2F7F4EA83B39060">
<mods:title id="98CC594332012ACED63AE500ED47FCFE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="63CDDA88FE93E61F8504FAE0C5F40905">
<mods:date id="3BE61AFB3D05A44FE6DA376F8FE69B64">2011</mods:date>
<mods:detail id="B90FB75398996718118E0B8AAABCA639" type="pubDate">
<mods:number id="AAE180F5B4EA591AE0335CA2069A0BE6">2011-03-01</mods:number>
</mods:detail>
<mods:detail id="F435FEDF5656EA13353C47911A484652" type="volume">
<mods:number id="D03121A01D006DBD1C166EDDFFBD473B">2780</mods:number>
</mods:detail>
<mods:detail id="91CB6799337EF14CA20863CACFDE34E1" type="issue">
<mods:number id="4612551C9B8BABF8C92883C7F3721A65">1</mods:number>
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<mods:start id="41F5B55DAF7E7A30C0A150BD483FE035">48</mods:start>
<mods:end id="B2E003B9AA2001A637BE4A0A206FC6A7">56</mods:end>
</mods:extent>
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<mods:location id="6F1A2F710A87C2988A81CE900D32E2D7">
<mods:url id="EEC0F2D511DECC45B77CBB34C18611B1">https://biotaxa.org/Zootaxa/article/view/zootaxa.2780.1.5</mods:url>
</mods:location>
<mods:classification id="ED6336F14C78432315337D9ED6AA6E58">journal article</mods:classification>
<mods:identifier id="18B6D5A12B96EC434E8DDB52A19F7230" type="DOI">10.11646/zootaxa.2780.1.5</mods:identifier>
<mods:identifier id="E95DEC8A9CF9FF41E5AF2B558AA1DD39" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A74564B0A5E9FD548D7AE01F6BD33C6B" type="Zenodo-Dep">5290601</mods:identifier>
<mods:identifier id="827848B96091CC51C5035A18299E2801" type="ZooBank">18EE66C7-3E0D-4032-9EF1-41E987F3E490</mods:identifier>
</mods:mods>
<treatment id="03BC87DD4333FF83FF6FA6A0FD4AB994" ID-DOI="http://doi.org/10.5281/zenodo.5294778" ID-Zenodo-Dep="5294778" LSID="urn:lsid:zoobank.org:act:B7E260ED-670A-4DC6-AF02-8FD7D7A7ADD7" httpUri="http://treatment.plazi.org/id/03BC87DD4333FF83FF6FA6A0FD4AB994" lastPageId="4" lastPageNumber="52" pageId="2" pageNumber="50">
<subSubSection id="C30F65404333FF85FF6FA6A0FC30BDC8" box="[151,954,655,681]" pageId="2" pageNumber="50" type="nomenclature">
<paragraph id="8BAA36CB4333FF85FF6FA6A0FC30BDC8" blockId="2.[151,954,655,714]" box="[151,954,655,681]" pageId="2" pageNumber="50">
<heading id="D0E281A74333FF85FF6FA6A0FC30BDC8" bold="true" box="[151,954,655,681]" fontSize="11" level="1" pageId="2" pageNumber="50" reason="1">
<emphasis id="B961EAD94333FF85FF6FA6A0FC30BDC8" bold="true" box="[151,954,655,681]" pageId="2" pageNumber="50">
<taxonomicName id="4C154D484333FF85FF6FA6A0FDA2BDC8" ID-CoL="3796K" authority="Reboleira &amp; Gonçalves &amp; Oromí, 2011" authorityName="Reboleira &amp; Gonçalves &amp; Oromí" authorityYear="2011" box="[151,552,655,681]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="species" species="lusitanica" status="sp. nov." subGenus="Lathromene">
<emphasis id="B961EAD94333FF85FF6FA6A0FDA2BDC8" bold="true" box="[151,552,655,681]" italics="true" pageId="2" pageNumber="50">Domene (Lathromene) lusitanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257A24333FF85FDC8A6A0FD4ABDC8" box="[560,704,655,681]" pageId="2" pageNumber="50" rank="species">new species</taxonomicNameLabel>
Reboleira &amp; Oromí
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F65404333FF85FF6FA69DFED1BDAB" box="[151,347,690,714]" pageId="2" pageNumber="50" type="description">
<paragraph id="8BAA36CB4333FF85FF6FA69DFED1BDAB" blockId="2.[151,954,655,714]" box="[151,347,690,714]" pageId="2" pageNumber="50">
(
<figureCitation id="132E2A4E4333FF85FF67A69DFE8BBDAB" box="[159,257,690,714]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="2.[151,250,1567,1590]" captionStartId-1="3.[151,250,1320,1343]" captionStartId-2="3.[151,250,1993,2016]" captionStartId-3="4.[151,250,1085,1108]" captionStartId-4="4.[151,250,1510,1533]" captionTargetBox-0="[189,1412,1037,1535]" captionTargetBox-1="[159,1426,294,1292]" captionTargetBox-2="[475,1101,1432,1958]" captionTargetBox-4="[566,1019,1159,1485]" captionTargetId-0="figure-473@2.[167,1463,1020,1565]" captionTargetId-1="figure-122@3.[151,1436,277,1299]" captionTargetId-2="figure-125@3.[473,1113,1423,1972]" captionTargetId-4="figure-330@4.[558,1029,1156,1489]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-4="4" captionText-0="FIGURE 2. Habitus of Domene lusitanica n. sp. Reboleira &amp; Oromí. Scale bar: 1 mm." captionText-1="FIGURE 3. Head of Domene lusitanica n. sp. Reboleira &amp; Oromí. a) detail of the reduced eye and chaetotaxy in lateral view; b) detail of mouthparts and gular suture in ventral view. Scale: 0.25 mm." captionText-2="FIGURE 4. Male sternite VIII in ventral view, with detailed microsculpture of the glabrous anterior part. Scale: 0.2 mm." captionText-3="FIGURE 5. Genital segment in ventral view. a) male; b) female." captionText-4="FIGURE 6. Aedeagus of Domene lusitanica n. sp. a) lateral view; b) dorsal view; c) ventral view. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5290607" figureDoi-1="http://doi.org/10.5281/zenodo.5290609" figureDoi-2="http://doi.org/10.5281/zenodo.5290613" figureDoi-4="http://doi.org/10.5281/zenodo.5290615" httpUri-0="https://zenodo.org/record/5290607/files/figure.png" httpUri-1="https://zenodo.org/record/5290609/files/figure.png" httpUri-2="https://zenodo.org/record/5290613/files/figure.png" httpUri-4="https://zenodo.org/record/5290615/files/figure.png" pageId="2" pageNumber="50">Figs 2–6</figureCitation>
and
<figureCitation id="132E2A4E4333FF85FEC0A69DFED9BDAB" box="[312,339,690,714]" captionStart="FIGURE 7" captionStartId="6.[151,250,1108,1131]" captionTargetBox="[158,1407,309,1065]" captionTargetId="figure-287@6.[151,1436,301,1072]" captionTargetPageId="6" captionText="FIGURE 7. Aedeagus outlines of the subgenus Lathromene, in lateral and ventral views. a) D. bergidi; b) D. cantabrica; c) D. gridelliana; d) D. gallaeciana; e) D. lusitanica n. sp.; f) D. caurelensis; g) D. subiasi; and h) D. scopaeella. (Adapted from Coiffait 1982; Feldmann &amp; Hernando 2005 and Outerelo 2000)." figureDoi="http://doi.org/10.5281/zenodo.5290619" httpUri="https://zenodo.org/record/5290619/files/figure.png" pageId="2" pageNumber="50">7e</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F65404333FF85FF6FA6D4FEF4BC1F" pageId="2" pageNumber="50" type="materials_examined">
<paragraph id="8BAA36CB4333FF85FF6FA6D4FEF4BC1F" blockId="2.[151,1436,762,966]" pageId="2" pageNumber="50">
<materialsCitation id="3B7D3C964333FF85FF6FA6D4FAC6BC73" box="[151,1356,762,787]" collectingDate="2009-11-28" collectionCode="PO" collectorName="da Ceramica &amp; S. Reboleira" country="Portugal" location="Serra de Sico" municipality="Gruta da Ceramica" pageId="2" pageNumber="50" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B961EAD94333FF85FF6FA6D4FE87BC72" bold="true" box="[151,269,763,787]" pageId="2" pageNumber="50">
<typeStatus id="54AE88694333FF85FF6FA6D4FE82BC72" box="[151,264,763,787]" pageId="2" pageNumber="50" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D13FD424333FF85FEEEA6D4FEA1BC73" box="[278,299,763,786]" pageId="2" pageNumber="50" type="male">♂</specimenCount>
,
<collectingCountry id="F302765B4333FF85FEC3A6D5FE16BC73" box="[315,412,762,786]" name="Portugal" pageId="2" pageNumber="50">Portugal</collectingCountry>
,
<location id="8ECA60104333FF85FE52A6D5FDCFBC73" LSID="urn:lsid:plazi:treatment:03BC87DD4333FF83FF6FA6A0FD4AB994:8ECA60104333FF85FE52A6D5FDCFBC73" box="[426,581,762,786]" country="Portugal" municipality="Gruta da Ceramica" name="Serra de Sico" pageId="2" pageNumber="50">Serra de Sicó</location>
,
<collectingMunicipality id="6BCEACB14333FF85FDABA6D5FCA4BC73" box="[595,814,762,786]" pageId="2" pageNumber="50">
Gruta
<collectorName id="26E0531D4333FF85FD64A6D5FCA4BC73" box="[668,814,762,786]" pageId="2" pageNumber="50">da Cerâmica</collectorName>
</collectingMunicipality>
,
<date id="FFAB100B4333FF85FCC3A6D5FC37BC73" box="[827,957,762,786]" pageId="2" pageNumber="50" value="2009-11-28">
<collectingDate id="EFEFE9E34333FF85FCC3A6D5FC37BC73" box="[827,957,762,786]" pageId="2" pageNumber="50" value="2009-11-28">28.XI.2009</collectingDate>
</date>
,
<collectorName id="26E0531D4333FF85FC33A6D5FBDDBC73" box="[971,1111,762,786]" pageId="2" pageNumber="50">S. Reboleira</collectorName>
leg, deposited at
<collectionCode id="ED04AE0E4333FF85FADDA6D5FAC6BC73" box="[1317,1356,762,786]" country="Russia" name="Collection of the Zoological Institute of the Russian Academy of Sciences" pageId="2" pageNumber="50">PO</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7D3C964333FF85FAA1A6D4FCB0BC57" collectingDate="2009-11-28" collectionCode="PO" collectorName="da Ceramica &amp; S. Reboleira" country="Portugal" location="Serra de Sico" municipality="Gruta da Ceramica" pageId="2" pageNumber="50" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B961EAD94333FF85FAA1A6D4FF54BC56" bold="true" pageId="2" pageNumber="50">
<typeStatus id="54AE88694333FF85FAA1A6D4FF53BC56" pageId="2" pageNumber="50" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D13FD424333FF85FF13A731FE91BC57" box="[235,283,798,822]" pageId="2" pageNumber="50" type="male">1 ♂</specimenCount>
, same data as the holotype, deposited at
<collectionCode id="ED04AE0E4333FF85FCEBA731FCB0BC57" box="[787,826,798,822]" country="Russia" name="Collection of the Zoological Institute of the Russian Academy of Sciences" pageId="2" pageNumber="50">PO</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B7D3C964333FF85FCB3A731FD2CBC3B" collectingDate="2010-03-21" collectorName="da Ceramica &amp; S. Reboleira" country="Portugal" location="Serra de Sico" municipality="Gruta da Ceramica" pageId="2" pageNumber="50" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D13FD424333FF85FCB3A731FCF1BC57" box="[843,891,798,822]" pageId="2" pageNumber="50" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D13FD424333FF85FC45A731FC67BC57" box="[957,1005,798,822]" pageId="2" pageNumber="50" type="female">1 ♀</specimenCount>
,
<location id="8ECA60104333FF85FBF9A731FB2EBC57" LSID="urn:lsid:plazi:treatment:03BC87DD4333FF83FF6FA6A0FD4AB994:8ECA60104333FF85FBF9A731FB2EBC57" box="[1025,1188,798,822]" country="Portugal" municipality="Gruta da Ceramica" name="Serra de Sico" pageId="2" pageNumber="50">Serra de Sicó</location>
,
<collectingMunicipality id="6BCEACB14333FF85FB4EA731FA12BC57" box="[1206,1432,798,822]" pageId="2" pageNumber="50">
Gruta
<collectorName id="26E0531D4333FF85FAFAA731FA12BC57" box="[1282,1432,798,822]" pageId="2" pageNumber="50">da Cerâmica</collectorName>
</collectingMunicipality>
,
<date id="FFAB100B4333FF85FF6FA76DFE9DBC3B" box="[151,279,834,858]" pageId="2" pageNumber="50" value="2010-03-21">
<collectingDate id="EFEFE9E34333FF85FF6FA76DFE9DBC3B" box="[151,279,834,858]" pageId="2" pageNumber="50" value="2010-03-21">21.III.2010</collectingDate>
</date>
,
<collectorName id="26E0531D4333FF85FEDCA76DFE3ABC3B" box="[292,432,834,858]" pageId="2" pageNumber="50">S. Reboleira</collectorName>
leg., deposited at SR
</materialsCitation>
;
<materialsCitation id="3B7D3C964333FF85FD4AA76DFEF0BC1F" collectingDate="2010-12-27" collectorName="da Ceramica &amp; S. Reboleira" country="Portugal" location="Serra de Sico" municipality="Gruta da Ceramica" pageId="2" pageNumber="50" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D13FD424333FF85FD4AA76DFD54BC3B" box="[690,734,834,858]" pageId="2" pageNumber="50" type="male">3 ♂</specimenCount>
,
<location id="8ECA60104333FF85FD14A76DFC0CBC3B" LSID="urn:lsid:plazi:treatment:03BC87DD4333FF83FF6FA6A0FD4AB994:8ECA60104333FF85FD14A76DFC0CBC3B" box="[748,902,834,858]" country="Portugal" municipality="Gruta da Ceramica" name="Serra de Sico" pageId="2" pageNumber="50">Serra de Sicó</location>
,
<collectingMunicipality id="6BCEACB14333FF85FC6BA76DFBE6BC3B" box="[915,1132,834,858]" pageId="2" pageNumber="50">
Gruta
<collectorName id="26E0531D4333FF85FC23A76DFBE6BC3B" box="[987,1132,834,858]" pageId="2" pageNumber="50">da Cerâmica</collectorName>
</collectingMunicipality>
,
<date id="FFAB100B4333FF85FB80A76DFA89BC3B" box="[1144,1283,834,858]" pageId="2" pageNumber="50" value="2010-12-27">
<collectingDate id="EFEFE9E34333FF85FB80A76DFA89BC3B" box="[1144,1283,834,858]" pageId="2" pageNumber="50" value="2010-12-27">27.XII.2010</collectingDate>
</date>
,
<collectorName id="26E0531D4333FF85FAE8A76DFA16BC3B" box="[1296,1436,834,858]" pageId="2" pageNumber="50">S. Reboleira</collectorName>
leg, deposited at SR
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F65404333FF83FF3FA7A5FCD8B9CC" lastPageId="4" lastPageNumber="52" pageId="2" pageNumber="50" type="description">
<paragraph id="8BAA36CB4333FF85FF3FA7A5FDCDBCA7" blockId="2.[151,1436,762,966]" pageId="2" pageNumber="50">
<emphasis id="B961EAD94333FF85FF3FA7A5FED2BCC2" bold="true" box="[199,344,906,931]" inLineHeading="true" pageId="2" pageNumber="50" reason="1">Description.</emphasis>
Microphtalmous, wingless, reddish brown and elongate body (fig. 2). Total body length:
<quantity id="4CED9B2E4333FF85FAB7A7A5FF48BCA7" metricMagnitude="-3" metricUnit="m" metricValue="9.24" metricValueMax="9.48" metricValueMin="9.0" pageId="2" pageNumber="50" unit="mm" value="9.24" valueMax="9.48" valueMin="9.0">9–9.48 mm</quantity>
(a contracted specimen
<quantity id="4CED9B2E4333FF85FE2BA781FDB0BCA7" box="[467,570,942,966]" metricMagnitude="-3" metricUnit="m" metricValue="7.56" pageId="2" pageNumber="50" unit="mm" value="7.56">7.56 mm</quantity>
).
</paragraph>
<caption id="DF6A66434333FF85FF6FA230FB82B954" ID-DOI="http://doi.org/10.5281/zenodo.5290607" ID-Zenodo-Dep="5290607" box="[151,1032,1567,1590]" httpUri="https://zenodo.org/record/5290607/files/figure.png" pageId="2" pageNumber="50" startId="2.[151,250,1567,1590]" targetBox="[189,1412,1037,1535]" targetPageId="2">
<paragraph id="8BAA36CB4333FF85FF6FA230FB82B954" blockId="2.[151,1032,1567,1590]" box="[151,1032,1567,1590]" pageId="2" pageNumber="50">
<emphasis id="B961EAD94333FF85FF6FA230FE99B957" bold="true" box="[151,275,1567,1590]" pageId="2" pageNumber="50">FIGURE 2.</emphasis>
Habitus of
<taxonomicName id="4C154D484333FF85FE71A20FFDC2B954" authority="Reboleira &amp; Gonçalves &amp; Oromí, 2011" authorityName="Reboleira &amp; Gonçalves &amp; Oromí" authorityYear="2011" box="[393,584,1567,1589]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="species" species="lusitanica" status="sp. nov.">
<emphasis id="B961EAD94333FF85FE71A20FFDC2B954" box="[393,584,1567,1589]" italics="true" pageId="2" pageNumber="50">Domene lusitanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257A24333FF85FDB6A20FFD0FB954" box="[590,645,1568,1589]" pageId="2" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
Reboleira &amp; Oromí. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BAA36CB4333FF85FF3DA24DFB1DB886" blockId="2.[151,1437,1634,2024]" pageId="2" pageNumber="50">
<emphasis id="B961EAD94333FF85FF3DA24DFE8FB91B" bold="true" box="[197,261,1634,1658]" pageId="2" pageNumber="50">Head</emphasis>
(fig. 3): subquadrate/orbicular (1.15 times longer than wide), wider than pronotum; length
<quantity id="4CED9B2E4333FF85FAF5A24DFA1DB918" box="[1293,1431,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="2" pageNumber="50" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 mm</quantity>
, width
<quantity id="4CED9B2E4333FF85FF27A2A9FE0DB9FF" box="[223,391,1670,1694]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" metricValueMax="1.14" metricValueMin="1.02" pageId="2" pageNumber="50" unit="mm" value="1.08" valueMax="1.14" valueMin="1.02">1.02–1.14 mm</quantity>
; eyes reduced in size and pigmentation, without ommatidia; long and thin seta inserted in a supraocular, small semicircular depression (fig. 3a); two dark spots in the vertex indicating the insertions of the dorsal arms of the tentorium; gular sutures well defined, converging towards neck; neck distinct; head with dorsal punctuation well defined, similar to that of pronotum, but distinctly finer than that of elytra; antennae filiform,
<quantity id="4CED9B2E4333FF85FA81A2DDFF48B851" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="50" unit="mm" value="2.7">2.7 mm</quantity>
long, directed backwards and reaching base of pronotum; all antennomeres longer than wide, the 1
<superScript id="7C609B834333FF85FAE3A339FAAFB845" attach="left" box="[1307,1317,1814,1828]" fontSize="6" pageId="2" pageNumber="50">st</superScript>
longest of all; 3
<superScript id="7C609B834333FF85FF2CA312FF6BB82A" attach="left" box="[212,225,1853,1867]" fontSize="6" pageId="2" pageNumber="50">rd</superScript>
antennomere 1.5 times, 11
<superScript id="7C609B834333FF85FDDAA312FDA4B82A" attach="left" box="[546,558,1853,1867]" fontSize="6" pageId="2" pageNumber="50">th</superScript>
antennomere 1.3 times, longer than the remaining subequal antennomeres; labrum deeply emarginate with large setae at the margin; mandibles symmetrical with two distinct teeth in its inner edge (
<figureCitation id="132E2A4E4333FF85FF24A3A8FEA5B8FE" box="[220,303,1927,1951]" captionStart="FIGURE 3" captionStartId="3.[151,250,1320,1343]" captionTargetBox="[159,1426,294,1292]" captionTargetId="figure-122@3.[151,1436,277,1299]" captionTargetPageId="3" captionText="FIGURE 3. Head of Domene lusitanica n. sp. Reboleira &amp; Oromí. a) detail of the reduced eye and chaetotaxy in lateral view; b) detail of mouthparts and gular suture in ventral view. Scale: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5290609" httpUri="https://zenodo.org/record/5290609/files/figure.png" pageId="2" pageNumber="50">Fig. 3b</figureCitation>
); maxillary palpus with four segments, second and third subequal, apical segment reduced and conical; internal lobe of maxilla with tufts of setae; labium with bilobed glossae, framed by densely hairy paraglossae; labial palpus with three segments, the second being the largest and the last one the thinnest.
</paragraph>
<paragraph id="8BAA36CB4332FF84FF3DA4B7FBEFBF96" blockId="3.[151,1436,151,247]" pageId="3" pageNumber="51">
<emphasis id="B961EAD94332FF84FF3DA4B7FEB4BFD1" bold="true" box="[197,318,152,176]" pageId="3" pageNumber="51">Pronotum</emphasis>
: length
<quantity id="4CED9B2E4332FF84FE60A4B8FDB6BFCE" box="[408,572,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="3" pageNumber="51" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.20–1.30 mm</quantity>
, width
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; as long as the head and 1.3x longer than wide; anterior angles weak, posterior angles marked but rounded; with a slight groove at the posterior third; basal margin well defined; evenly distributed punctuation except for a smooth midline in its anterior part.
</paragraph>
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<paragraph id="8BAA36CB4332FF84FF6FA107FCFCBA3C" blockId="3.[151,1436,1320,1373]" pageId="3" pageNumber="51">
<emphasis id="B961EAD94332FF84FF6FA107FE99BA5E" bold="true" box="[151,275,1320,1343]" pageId="3" pageNumber="51">FIGURE 3.</emphasis>
Head of
<taxonomicName id="4C154D484332FF84FE8AA106FDBABA5F" authority="Reboleira &amp; Gonçalves &amp; Oromí, 2011" authorityName="Reboleira &amp; Gonçalves &amp; Oromí" authorityYear="2011" box="[370,560,1320,1342]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="51" phylum="Arthropoda" rank="species" species="lusitanica" status="sp. nov.">
<emphasis id="B961EAD94332FF84FE8AA106FDBABA5F" box="[370,560,1320,1342]" italics="true" pageId="3" pageNumber="51">Domene lusitanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257A24332FF84FDCFA106FDE4BA5F" box="[567,622,1321,1342]" pageId="3" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
Reboleira &amp; Oromí. a) detail of the reduced eye and chaetotaxy in lateral view; b) detail of mouthparts and gular suture in ventral view. Scale: 0.25 mm.
</paragraph>
</caption>
<caption id="DF6A66434332FF84FF6FA3E6FAE8B8BE" ID-DOI="http://doi.org/10.5281/zenodo.5290613" ID-Zenodo-Dep="5290613" box="[151,1378,1993,2016]" httpUri="https://zenodo.org/record/5290613/files/figure.png" pageId="3" pageNumber="51" startId="3.[151,250,1993,2016]" targetBox="[475,1101,1432,1958]" targetPageId="3">
<paragraph id="8BAA36CB4332FF84FF6FA3E6FAE8B8BE" blockId="3.[151,1378,1993,2016]" box="[151,1378,1993,2016]" pageId="3" pageNumber="51">
<emphasis id="B961EAD94332FF84FF6FA3E6FE98B881" bold="true" box="[151,274,1993,2016]" pageId="3" pageNumber="51">FIGURE 4.</emphasis>
Male sternite VIII in ventral view, with detailed microsculpture of the glabrous anterior part. Scale: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BAA36CB4335FF83FF3DA4B7FE00BF96" blockId="4.[151,1436,151,503]" pageId="4" pageNumber="52">
<emphasis id="B961EAD94335FF83FF3DA4B7FE90BFD1" bold="true" box="[197,282,152,176]" pageId="4" pageNumber="52">Elytra:</emphasis>
1.00–
<quantity id="4CED9B2E4335FF83FE9EA4B8FE44BFCE" box="[358,462,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="52" unit="mm" value="1.1">1.10 mm</quantity>
long, maximum width
<quantity id="4CED9B2E4335FF83FD12A4B8FC1FBFCE" box="[746,917,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.3" metricValueMin="4.2" pageId="4" pageNumber="52" unit="mm" value="0.425" valueMax="0.43" valueMin="0.42">0.42–0.43 mm</quantity>
; 2.5x longer than wide, approximately 1.5x shorter than head or pronotum; flat in dorsal view; lateral sides of elytra parallel; with coarse, rugose and confluent punctation. Wingless.
</paragraph>
<paragraph id="8BAA36CB4335FF83FF3DA529FBCEBECE" blockId="4.[151,1436,151,503]" pageId="4" pageNumber="52">
<emphasis id="B961EAD94335FF83FF3DA529FEC9BE7E" bold="true" box="[197,323,262,287]" pageId="4" pageNumber="52">Abdomen:</emphasis>
Maximum width at sternite VII (the 5
<superScript id="7C609B834335FF83FD0CA52BFC8ABE73" attach="left" box="[756,768,260,274]" fontSize="6" pageId="4" pageNumber="52">th</superScript>
visible); tergites with confluent punctuation and microsculpture composed of transverse meshes in the intersegmental membrane. Male: sternite VIII with a glabrous anterior strip with microsculpture as in figure 4, and the posterior margin with a relatively broad U-shaped incision, surrounded by a small glabrous area; genital segment in ventral view as in figure 5a. Female: with a large glabrous area in the centre of the genital segment in dorsal view; ventral view as in figure 5b.
</paragraph>
<paragraph id="8BAA36CB4335FF83FF3DA594FE67BE96" blockId="4.[151,1436,151,503]" pageId="4" pageNumber="52">
<emphasis id="B961EAD94335FF83FF3DA594FE8CBEB2" bold="true" box="[197,262,443,467]" pageId="4" pageNumber="52">Legs:</emphasis>
elongate, forelegs with antennal cleaning organ composed by two complementary scooped-out structures located on the femur and tibia.
</paragraph>
<caption id="DF6A66434335FF83FF6FA012FCA2BB35" box="[151,808,1085,1108]" pageId="4" pageNumber="52" startId="4.[151,250,1085,1108]">
<paragraph id="8BAA36CB4335FF83FF6FA012FCA2BB35" blockId="4.[151,808,1085,1108]" box="[151,808,1085,1108]" pageId="4" pageNumber="52">
<emphasis id="B961EAD94335FF83FF6FA012FE98BB32" bold="true" box="[151,274,1085,1108]" pageId="4" pageNumber="52">FIGURE 5.</emphasis>
Genital segment in ventral view. a) male; b) female.
</paragraph>
</caption>
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<paragraph id="8BAA36CB4335FF83FF6FA1C9FAB4BA9D" blockId="4.[151,1342,1510,1533]" box="[151,1342,1510,1533]" pageId="4" pageNumber="52">
<emphasis id="B961EAD94335FF83FF6FA1C9FE98BA9C" bold="true" box="[151,274,1510,1533]" pageId="4" pageNumber="52">FIGURE 6.</emphasis>
Aedeagus of
<taxonomicName id="4C154D484335FF83FE66A1C8FDD7BA9C" authority="Reboleira &amp; Gonçalves &amp; Oromí, 2011" authorityName="Reboleira &amp; Gonçalves &amp; Oromí" authorityYear="2011" box="[414,605,1511,1533]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="52" phylum="Arthropoda" rank="species" species="lusitanica" status="sp. nov.">
<emphasis id="B961EAD94335FF83FE66A1C8FDD7BA9C" box="[414,605,1511,1533]" italics="true" pageId="4" pageNumber="52">Domene lusitanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257A24335FF83FD9BA1C8FD10BA9D" box="[611,666,1511,1532]" pageId="4" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
a) lateral view; b) dorsal view; c) ventral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BAA36CB4335FF83FF3DA206FCD8B9CC" blockId="4.[151,1436,1577,1781]" pageId="4" pageNumber="52">
<emphasis id="B961EAD94335FF83FF3DA206FEB2B923" bold="true" box="[197,312,1577,1602]" pageId="4" pageNumber="52">Aedeagus</emphasis>
(fig. 6): with fused lateral lobes (fig. 6c); ellipse-shaped in dorsal view, presenting a v-shaped scletotized blade covering the upper part and the apical edge of the aedeagus (
<figureCitation id="132E2A4E4335FF83FBEBA262FBE2B904" box="[1043,1128,1613,1637]" captionStart="FIGURE 6" captionStartId="4.[151,250,1510,1533]" captionTargetBox="[566,1019,1159,1485]" captionTargetId="figure-330@4.[558,1029,1156,1489]" captionTargetPageId="4" captionText="FIGURE 6. Aedeagus of Domene lusitanica n. sp. a) lateral view; b) dorsal view; c) ventral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290615" httpUri="https://zenodo.org/record/5290615/files/figure.png" pageId="4" pageNumber="52">Fig. 6b</figureCitation>
); a well sclerotized ventral blade, with a pointed apex, not exceeding the edge of the aedeagus; median lobe with two internal small finger-like structures and internal sac with two plates of scales (fig. 6a,b).
</paragraph>
</subSubSection>
<subSubSection id="C30F65404335FF83FF3FA296FD4AB994" pageId="4" pageNumber="52" type="etymology">
<paragraph id="8BAA36CB4335FF83FF3FA296FD4AB994" blockId="4.[151,1436,1577,1781]" pageId="4" pageNumber="52">
<emphasis id="B961EAD94335FF83FF3FA296FEC4B9B0" bold="true" box="[199,334,1721,1745]" inLineHeading="true" pageId="4" pageNumber="52" reason="1">Etymology.</emphasis>
The specific epithet is related to the distribution of the species, which is located in the Lusitanic biospeleological district in the centre of
<collectingCountry id="F302765B4335FF83FDA2A2F2FD36B994" box="[602,700,1757,1781]" name="Portugal" pageId="4" pageNumber="52">Portugal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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