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<document id="544891D572C03D73AA2C3AED4338BFFB" ID-DOI="10.5281/zenodo.180818" ID-GBIF-Dataset="850fc9b3-67a6-4ced-b77b-3cc2bebe8857" ID-ISSN="1175-5326" ID-Zenodo-Dep="180818" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459816636239" checkinUser="plazi" docAuthor="Ivie, Michael A. &amp; Philips, Keith" docDate="2008" docId="03BC87DCAF3BFFB6FF68F8D0FA77490F" docLanguage="en" docName="zt01701p014.pdf" docOrigin="Zootaxa 1701" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Canthonella Chapin" docType="key" docVersion="8" lastPageNumber="8" masterDocId="FF85FFA4AF3DFFB1FFFFFFE8FFEC4D74" masterDocTitle="Three new species of Canthonella Chapin from Hispaniola, with new records and Nomenclatural changes for West Indian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="7" updateTime="1698221501712" updateUser="plazi">
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<mods:title id="E108BFF810D6FADC3307C5490B76A546">Three new species of Canthonella Chapin from Hispaniola, with new records and Nomenclatural changes for West Indian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="D1B203D6949693AF36ABFC6278D18E24">Ivie, Michael A.</mods:namePart>
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<mods:namePart id="7439F1BB5F0F331F45E73575EE98EC59">Philips, Keith</mods:namePart>
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<mods:date id="0C7843829341E977CB572ABC5AF34E42">2008</mods:date>
<mods:detail id="4011B8F2F1E36B2C805FD573D8DBAE49" type="volume">
<mods:number id="15C867B48580A50B2981948A5A8AB9B3">1701</mods:number>
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<mods:identifier id="87472D801B35D82BE6542BDE7B9D6AFE" type="DOI">10.5281/zenodo.180818</mods:identifier>
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<mods:identifier id="2533A22459517B272691345F9D6AFCAB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A4139B0315DAA7EE046ABD761D3EFAC4" type="Zenodo-Dep">180818</mods:identifier>
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<treatment id="03BC87DCAF3BFFB6FF68F8D0FA77490F" ID-DOI="http://doi.org/10.5281/zenodo.6233225" ID-GBIF-Taxon="119354874" ID-Zenodo-Dep="6233225" LSID="urn:lsid:plazi:treatment:03BC87DCAF3BFFB6FF68F8D0FA77490F" httpUri="http://treatment.plazi.org/id/03BC87DCAF3BFFB6FF68F8D0FA77490F" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection id="C30F6541AF3BFFB7FF68F8D0FD3D4A06" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BAA36CAAF3BFFB7FF68F8D0FD664A26" blockId="6.[151,721,1848,1906]" box="[151,650,1848,1874]" pageId="6" pageNumber="7">
<heading id="D0E281A6AF3BFFB7FF68F8D0FD664A26" bold="true" box="[151,650,1848,1874]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B961EAD8AF3BFFB7FF68F8D0FD664A26" bold="true" box="[151,650,1848,1874]" pageId="6" pageNumber="7">
Key to West Indian
<taxonomicName id="4C154D49AF3BFFB7FE6FF8D0FD664A26" ID-CoL="3GWB" authority="Chapin" authorityName="Chapin" box="[400,650,1848,1874]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAD8AF3BFFB7FE6FF8D0FDCA4A26" bold="true" box="[400,550,1848,1874]" italics="true" pageId="6" pageNumber="7">Canthonella</emphasis>
Chapin
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36CAAF3BFFB7FF68F8B3FD3D4A06" blockId="6.[151,721,1848,1906]" box="[151,721,1883,1906]" pageId="6" pageNumber="7">
(Modified from
<bibRefCitation id="EF844B3BAF3BFFB7FEBBF8B3FE004A06" author="Matthews" box="[324,492,1883,1906]" pageId="6" pageNumber="13" refString="Matthews, E. G. (1966) A taxonomic and zoogeographic survey of the Scarabaeinae of the Antilles (Coleoptera: Scarabaeidae). Memoirs of the American Entomological Society, 21, 1 - 134." type="journal article" year="1966">Matthews 1966</bibRefCitation>
,
<bibRefCitation id="EF844B3BAF3BFFB7FE09F8B3FD264A06" author="Ivie" box="[502,714,1883,1906]" pageId="6" pageNumber="13" refString="Ivie, M. A. &amp; Philips, K. (1990) New species of Canthonella Chapin from Hispaniola (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the New York Entomological Society, 98, 429 - 433." type="journal article" year="1990">Ivie &amp; Philips 1990</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F6541AF3BFFB6FF68F84DFA77490F" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="key">
<keyStep id="30E12E6FAF3BFFB7FF68F84DFA774A93" pageId="6" pageNumber="7">
<paragraph id="8BAA36CAAF3BFFB7FF68F84DFA704ACB" blockId="6.[151,1436,1957,2023]" box="[151,1436,1957,1983]" pageId="6" pageNumber="7">
<keyLead id="30E495FFAF3BFFB7FF68F84DFA704ACB" box="[151,1436,1957,1983]" pageId="6" pageNumber="7">1. Protibia with 3 teeth on outer margin...........................................................................................................2</keyLead>
</paragraph>
<paragraph id="8BAA36CAAF3BFFB7FF68F825FA774A93" blockId="6.[151,1436,1957,2023]" box="[151,1435,1997,2023]" pageId="6" pageNumber="7">
<keyLead id="30E495FFAF3BFFB7FF68F825FA774A93" box="[151,1435,1997,2023]" pageId="6" pageNumber="7">- Protibia with 2 teeth on outer margin...........................................................................................................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E6FAF3AFFB6FF68FF70FA774C5E" pageId="7" pageNumber="8">
<paragraph id="8BAA36CAAF3AFFB6FF68FF70FA704DAE" blockId="7.[151,1436,152,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FF70FA704DAE" pageId="7" pageNumber="8">
2. Elytral interstriae uneven in height, especially on declivity; protibia very narrow;
<collectingCountry id="F302765AAF3AFFB6FB80FF70FAE64DC6" box="[1151,1290,152,178]" name="Puerto Rico" pageId="7" pageNumber="8">Puerto Rico</collectingCountry>
and the
<collectingCountry id="F302765AAF3AFFB6FA92FF70FEAB4DAE" name="Virgin Islands" pageId="7" pageNumber="8">Virgin Islands</collectingCountry>
<emphasis id="B961EAD8AF3AFFB6FEB6FF28FAD34DAD" box="[329,1343,192,217]" italics="true" pageId="7" pageNumber="8">
............................................................................................................
<taxonomicName id="4C154D49AF3AFFB6FB9BFF28FAD34DAD" ID-CoL="QPFS" authority="Chapin" authorityName="Chapin" box="[1124,1343,192,217]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="parva">Canthonella parva</taxonomicName>
</emphasis>
Chapin
</keyLead>
</paragraph>
<paragraph id="8BAA36CAAF3AFFB6FF68FF00FA774C5E" blockId="7.[151,1436,152,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FF00FA774C5E" pageId="7" pageNumber="8">
- Elytral interstriae flat, even in height; protibia widened apically; Hispaniola ...............................................
<emphasis id="B961EAD8AF3AFFB6FF33FEF8FB5F4C5D" box="[204,1203,272,297]" italics="true" pageId="7" pageNumber="8">
.....................................................................................................
<taxonomicName id="4C154D49AF3AFFB6FC4AFEF8FB5F4C5D" ID-CoL="QPFJ" authority="Matthews" authorityName="Matthews" box="[949,1203,272,297]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="constans">Canthonella constans</taxonomicName>
</emphasis>
Matthews (
<figureCitation id="132E2A4FAF3AFFB6FAC2FEF8FA7F4C5E" box="[1341,1427,272,298]" captionStart="FIGURES 19 – 22" captionStartId="6.[151,269,1258,1282]" captionTargetBox="[291,1311,196,1233]" captionTargetId="figure@6.[275,1312,168,1234]" captionTargetPageId="6" captionText="FIGURES 19 – 22. Dorsal habiti. 19. Canthonella jarmilea; 20. Canthonella quesquaya; 21. Canthonella sikesi; 22. Canthonella constans." httpUri="https://zenodo.org/record/180820/files/figure.png" pageId="7" pageNumber="8">Fig. 22</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E6FAF3AFFB6FF68FED0FA774CD6" pageId="7" pageNumber="8">
<paragraph id="8BAA36CAAF3AFFB6FF68FED0FA774C26" blockId="7.[151,1436,152,1147]" box="[151,1435,312,338]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FED0FA774C26" box="[151,1435,312,338]" pageId="7" pageNumber="8">
3. Elytral macula yellow-orange, reaching base of elytron (
<figureCitation id="132E2A4FAF3AFFB6FCA4FED0FC4F4C26" box="[859,931,312,338]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
); Hispaniola..............................................4
</keyLead>
</paragraph>
<paragraph id="8BAA36CAAF3AFFB6FF68FE88FA774CD6" blockId="7.[151,1436,152,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FE88FA774CD6" pageId="7" pageNumber="8">
- Elytral macula whitish-yellow, not reaching base of elytron, with at least a narrow black margin at base (
<figureCitation id="132E2A4FAF3AFFB6FF30FE60FED34CD6" box="[207,319,392,418]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="7" pageNumber="8">Figs. 7, 8</figureCitation>
);
<collectingCountry id="F302765AAF3AFFB6FEA9FE60FE784CD6" box="[342,404,392,418]" name="Cuba" pageId="7" pageNumber="8">Cuba</collectingCountry>
and Hispaniola.................................................................................................................5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E6FAF3AFFB6FF68FE58FA774F30" pageId="7" pageNumber="8">
<paragraph id="8BAA36CAAF3AFFB6FF68FE58FA704C86" blockId="7.[151,1436,152,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FE58FA704C86" pageId="7" pageNumber="8">
4. Mesad edge of elytral spot reaching or nearly reaching suture; eighth elytral stria strongly punctate ..........
<emphasis id="B961EAD8AF3AFFB6FF33FE30FB5E4C85" box="[204,1202,472,497]" italics="true" pageId="7" pageNumber="8">
.....................................................................................................
<taxonomicName id="4C154D49AF3AFFB6FC47FE30FB5E4C85" ID-CoL="QPFN" authority="Matthews" authorityName="Matthews" box="[952,1202,472,497]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="isabellae">Canthonella isabellae</taxonomicName>
</emphasis>
Matthews (
<figureCitation id="132E2A4FAF3AFFB6FAC2FE30FA7E4C86" box="[1341,1426,472,498]" captionStart="FIGURES 23 – 25" captionStartId="8.[151,269,1586,1610]" captionTargetBox="[274,1244,494,1561]" captionTargetId="figure@8.[265,1321,469,1562]" captionTargetPageId="8" captionText="FIGURES 23 – 25. Dorsal habiti. 23. Canthonella isabellae; 24. Canthonella baorucensis; 25. Canthonella howdeni." httpUri="https://zenodo.org/record/180821/files/figure.png" pageId="7" pageNumber="8">Fig. 23</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BAA36CAAF3AFFB6FF68FDEAFA774F30" blockId="7.[151,1436,152,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FDEAFA774F30" pageId="7" pageNumber="8">
- Mesad edge of elytral spot reaching to 2nd stria; eighth elytral stria impunctate............................................
<emphasis id="B961EAD8AF3AFFB6FF33FDC3FB764F30" box="[204,1178,555,580]" italics="true" pageId="7" pageNumber="8">
.......................................................................................................
<taxonomicName id="4C154D49AF3AFFB6FC38FDC3FB764F30" ID-CoL="QPFV" box="[967,1178,555,580]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sikesi" status="sp. nov.">Canthonella sikesi</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A25257A3AF3AFFB6FB5FFDC2FAC14F30" box="[1184,1325,554,580]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="132E2A4FAF3AFFB6FAC2FDC2FA7E4F30" box="[1341,1426,554,580]" captionStart="FIGURES 19 – 22" captionStartId="6.[151,269,1258,1282]" captionTargetBox="[291,1311,196,1233]" captionTargetId="figure@6.[275,1312,168,1234]" captionTargetPageId="6" captionText="FIGURES 19 – 22. Dorsal habiti. 19. Canthonella jarmilea; 20. Canthonella quesquaya; 21. Canthonella sikesi; 22. Canthonella constans." httpUri="https://zenodo.org/record/180820/files/figure.png" pageId="7" pageNumber="8">Fig. 21</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E6FAF3AFFB6FF68FDBDFA774F9F" pageId="7" pageNumber="8">
<paragraph id="8BAA36CAAF3AFFB6FF68FDBDFA704FE3" blockId="7.[151,1436,152,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FDBDFA704FE3" pageId="7" pageNumber="8">
5. Elytral spot ending at or laterad to 3rd stria (
<figureCitation id="132E2A4FAF3AFFB6FD33FDBDFCDA4F1B" box="[716,822,597,623]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="7" pageNumber="8">Figs 7, 8</figureCitation>
); labium rounded, truncate or emarginate anteriorly (
<figureCitation id="132E2A4FAF3AFFB6FF30FD95FED24FE3" box="[207,318,637,663]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="7" pageNumber="8">Figs. 1, 2</figureCitation>
and
<bibRefCitation id="EF844B3BAF3AFFB6FE89FD95FDB34FE3" author="Ivie" box="[374,607,637,663]" pageId="7" pageNumber="13" refString="Ivie, M. A. &amp; Philips, K. (1990) New species of Canthonella Chapin from Hispaniola (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the New York Entomological Society, 98, 429 - 433." type="journal article" year="1990">Ivie &amp; Philips 1990</bibRefCitation>
,
<figureCitation id="132E2A4FAF3AFFB6FD94FD95FD474FE3" box="[619,683,637,663]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="7" pageNumber="8">Fig.1</figureCitation>
);
<collectingCountry id="F302765AAF3AFFB6FD3EFD95FD134FE3" box="[705,767,637,663]" name="Cuba" pageId="7" pageNumber="8">Cuba</collectingCountry>
and Hispaniola................................................................6
</keyLead>
</paragraph>
<paragraph id="8BAA36CAAF3AFFB6FF68FD41FA774F9F" blockId="7.[151,1436,152,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FD41FA774F9F" pageId="7" pageNumber="8">
- Elytral spot extending at least half-way across 2nd interstia (
<bibRefCitation id="EF844B3BAF3AFFB6FC54FD41FAC34FB7" author="Ivie" box="[939,1327,681,707]" pageId="7" pageNumber="13" refString="Ivie, M. A. &amp; Philips, K. (1990) New species of Canthonella Chapin from Hispaniola (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the New York Entomological Society, 98, 429 - 433." type="journal article" year="1990">
Ivie &amp; Philips 1990,
<figureCitation id="132E2A4FAF3AFFB6FB4AFD41FAC34FB7" box="[1205,1327,681,707]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="7" pageNumber="8">Figs. 3, 4</figureCitation>
</bibRefCitation>
); labium obtusely produced anteriorly; Hispaniola.....................................................................................................8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E6FAF3AFFB6FF68FD11FA704E4F" pageId="7" pageNumber="8">
<paragraph id="8BAA36CAAF3AFFB6FF68FD11FA774E67" blockId="7.[151,1436,152,1147]" box="[151,1435,761,787]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FD11FA774E67" box="[151,1435,761,787]" pageId="7" pageNumber="8">
6. Labium narrow, subtruncate (
<bibRefCitation id="EF844B3BAF3AFFB6FDE0FD11FCD04E67" author="Matthews" box="[543,828,761,787]" pageId="7" pageNumber="13" refString="Matthews, E. G. (1966) A taxonomic and zoogeographic survey of the Scarabaeinae of the Antilles (Coleoptera: Scarabaeidae). Memoirs of the American Entomological Society, 21, 1 - 134." type="journal article" year="1966">Matthews 1966, Fig. 84</bibRefCitation>
);
<collectingCountry id="F302765AAF3AFFB6FCAEFD11FC634E67" box="[849,911,761,787]" name="Cuba" pageId="7" pageNumber="8">Cuba</collectingCountry>
<emphasis id="B961EAD8AF3AFFB6FC6CFD11FBDC4E66" box="[915,1072,761,786]" italics="true" pageId="7" pageNumber="8">.....................</emphasis>
<taxonomicName id="4C154D49AF3AFFB6FBCFFD11FA774E67" ID-CoL="QPFT" authority="(Harold)" authorityName="Harold" baseAuthorityName="Harold" box="[1072,1435,761,787]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="B961EAD8AF3AFFB6FBCFFD11FADC4E66" box="[1072,1328,761,786]" italics="true" pageId="7" pageNumber="8">Canthonella pygmaea</emphasis>
(Harold)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAA36CAAF3AFFB6FF68FCC9FA704E4F" blockId="7.[151,1436,152,1147]" box="[151,1436,801,827]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FCC9FA704E4F" box="[151,1436,801,827]" pageId="7" pageNumber="8">
- Labium broader, truncate or emarginate (
<figureCitation id="132E2A4FAF3AFFB6FD58FCC9FCFA4E4F" box="[679,790,801,827]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="7" pageNumber="8">Figs. 1, 2</figureCitation>
); Hispaniola.................................................................7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E6FAF3AFFB6FF68FCA1FA774977" pageId="7" pageNumber="8">
<paragraph id="8BAA36CAAF3AFFB6FF68FCA1FA774EC7" blockId="7.[151,1436,152,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FCA1FA774EC7" pageId="7" pageNumber="8">
7. Elytral ground color castaneous, darker pitchy around elytral spot and along suture, pronotal disc usually castaneous with pitchy base; labium truncate apically (
<figureCitation id="132E2A4FAF3AFFB6FCB7FC99FC674EFF" box="[840,907,881,907]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
)...................................................................... .................................................................................................
<taxonomicName id="4C154D49AF3AFFB6FC61FC71FB754EC6" ID-CoL="QPFP" box="[926,1177,921,946]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="jarmilae" status="sp. nov.">
<emphasis id="B961EAD8AF3AFFB6FC61FC71FB754EC6" box="[926,1177,921,946]" italics="true" pageId="7" pageNumber="8">Canthonella jarmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257A3AF3AFFB6FB5FFC71FAC14EC7" box="[1184,1325,921,947]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="132E2A4FAF3AFFB6FAC2FC71FA7E4EC7" box="[1341,1426,921,947]" captionStart="FIGURES 19 – 22" captionStartId="6.[151,269,1258,1282]" captionTargetBox="[291,1311,196,1233]" captionTargetId="figure@6.[275,1312,168,1234]" captionTargetPageId="6" captionText="FIGURES 19 – 22. Dorsal habiti. 19. Canthonella jarmilea; 20. Canthonella quesquaya; 21. Canthonella sikesi; 22. Canthonella constans." httpUri="https://zenodo.org/record/180820/files/figure.png" pageId="7" pageNumber="8">Fig. 19</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BAA36CAAF3AFFB6FF68FC29FA774977" blockId="7.[151,1436,152,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FC29FA774977" pageId="7" pageNumber="8">
- Elytral and pronotal ground color black; labium emarginate apically (
<figureCitation id="132E2A4FAF3AFFB6FC0CFC29FBDA4EAF" box="[1011,1078,961,987]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
)...............................................
<emphasis id="B961EAD8AF3AFFB6FF33FC01FB754976" box="[204,1177,1001,1026]" italics="true" pageId="7" pageNumber="8">
...............................................................................................
<taxonomicName id="4C154D49AF3AFFB6FC73FC01FB754976" ID-CoL="68XGV" box="[908,1177,1001,1026]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="quesqueya" status="sp. nov.">Canthonella quesqueya</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A25257A3AF3AFFB6FB5FFC01FAC14977" box="[1184,1325,1001,1027]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="132E2A4FAF3AFFB6FAC2FC01FA7F4977" box="[1341,1427,1001,1027]" captionStart="FIGURES 19 – 22" captionStartId="6.[151,269,1258,1282]" captionTargetBox="[291,1311,196,1233]" captionTargetId="figure@6.[275,1312,168,1234]" captionTargetPageId="6" captionText="FIGURES 19 – 22. Dorsal habiti. 19. Canthonella jarmilea; 20. Canthonella quesquaya; 21. Canthonella sikesi; 22. Canthonella constans." httpUri="https://zenodo.org/record/180820/files/figure.png" pageId="7" pageNumber="8">Fig. 20</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E6FAF3AFFB6FF68FBF9FA77490F" pageId="7" pageNumber="8">
<paragraph id="8BAA36CAAF3AFFB6FF68FBF9FA774927" blockId="7.[151,1436,152,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FBF9FA774927" pageId="7" pageNumber="8">
8. Elytron with triangular spot at apex of interstriae 1, 2, and 3; a smaller (occasionally obsolete) macula at apex of stria 5; 3.3–4.8 mm
<emphasis id="B961EAD8AF3AFFB6FDF0FBD1FB6E4926" box="[527,1154,1081,1106]" italics="true" pageId="7" pageNumber="8">
.............................................
<taxonomicName id="4C154D49AF3AFFB6FCA2FBD1FB6E4926" ID-CoL="QPFF" authority="Ivie &amp; Philips" authorityName="Ivie &amp; Philips" box="[861,1154,1081,1106]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="baorucensis">Canthonella baorucensis</taxonomicName>
</emphasis>
Ivie &amp; Philips (
<figureCitation id="132E2A4FAF3AFFB6FAC2FBD1FA7F4927" box="[1341,1427,1081,1107]" captionStart="FIGURES 23 – 25" captionStartId="8.[151,269,1586,1610]" captionTargetBox="[274,1244,494,1561]" captionTargetId="figure@8.[265,1321,469,1562]" captionTargetPageId="8" captionText="FIGURES 23 – 25. Dorsal habiti. 23. Canthonella isabellae; 24. Canthonella baorucensis; 25. Canthonella howdeni." httpUri="https://zenodo.org/record/180821/files/figure.png" pageId="7" pageNumber="8">Fig. 24</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BAA36CAAF3AFFB6FF68FB89FA77490F" blockId="7.[151,1436,152,1147]" box="[151,1435,1121,1147]" pageId="7" pageNumber="8">
<keyLead id="30E495FFAF3AFFB6FF68FB89FA77490F" box="[151,1435,1121,1147]" pageId="7" pageNumber="8">
- Elytron immaculate in apical ½; 2.3–3.4 mm
<emphasis id="B961EAD8AF3AFFB6FD2AFB89FB6D490E" box="[725,1153,1121,1146]" italics="true" pageId="7" pageNumber="8">
........................
<taxonomicName id="4C154D49AF3AFFB6FC77FB89FB6D490E" ID-CoL="QPFL" authority="Ivie &amp; Philips" authorityName="Ivie &amp; Philips" box="[904,1153,1121,1146]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="howdeni">Canthonella howdeni</taxonomicName>
</emphasis>
Ivie &amp; Philips (
<figureCitation id="132E2A4FAF3AFFB6FAC2FB89FA7E490F" box="[1341,1426,1121,1147]" captionStart="FIGURES 23 – 25" captionStartId="8.[151,269,1586,1610]" captionTargetBox="[274,1244,494,1561]" captionTargetId="figure@8.[265,1321,469,1562]" captionTargetPageId="8" captionText="FIGURES 23 – 25. Dorsal habiti. 23. Canthonella isabellae; 24. Canthonella baorucensis; 25. Canthonella howdeni." httpUri="https://zenodo.org/record/180821/files/figure.png" pageId="7" pageNumber="8">Fig. 25</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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