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<mods:title id="D251A637405E5ABE0B69397625E5ED83">New synonymies of Coprophilus (Zonyptilus) pentatoma Fauvel, 1897 (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<taxonomicName id="4C154D47CA1ABA681BECC8F10AB8FE45" ID-CoL="5ZYJ6" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" box="[151,756,436,463]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
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<emphasis id="B961EAD6CA1ABA681BECC8F10A02FE44" bold="true" box="[151,590,436,462]" italics="true" pageId="1" pageNumber="131">Coprophilus (Zonyptilus) pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1ABA68192EC8F00AB8FE45" author="Fauvel" box="[597,756,437,463]" pageId="1" pageNumber="131" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
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<figureCitation id="132E2A41CA1ABA681BE4C8920946FE7A" box="[159,266,471,496]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1254,1277]" captionTargetBox="[157,1411,459,1214]" captionTargetId="figure@2.[151,1436,447,1230]" captionTargetPageId="2" captionText="FIGURES 1 3. Coprophilus (Zonyptilus) pentatoma Fauvel, 1897 (lectotype, male): 1 — body, dorsal view; 2 — aedeagus, lateral view; 3 — aedeagus, ventral view. Scale bar: fig. 1 — 1 mm; figs. 2, 3 — 0.25 mm." httpUri="https://zenodo.org/record/843165/files/figure.png" pageId="1" pageNumber="131">Figs. 13</figureCitation>
,
<figureCitation id="132E2A41CA1ABA681A6DC89D096AFE65" box="[278,294,472,495]" captionStart="FIGURES 6 7" captionStartId="4.[151,264,1982,2005]" captionTargetBox="[164,1422,949,1948]" captionTargetId="figure@4.[151,1437,934,1961]" captionTargetPageId="4" captionText="FIGURES 6 7. Coprophilus spp.: 6 — C. (Z.) rufitarsis Kashcheev, 1999 (holotype?, female), body, dorsal view; 7 — C. (Z.) pentatoma Fauvel, 1897 (“ Kebin ”, female), body, dorsal view. Scale bar: figs. 6, 7 — 1 mm." httpUri="https://zenodo.org/record/843169/files/figure.png" pageId="1" pageNumber="131">7</figureCitation>
,
<figureCitation id="132E2A41CA1ABA681A49C89D0973FE7A" box="[306,319,472,496]" captionStart="FIGURES 8 9" captionStartId="5.[151,264,1227,1250]" captionTargetBox="[170,1410,206,1188]" captionTargetId="figure@5.[151,1436,193,1206]" captionTargetPageId="5" captionText="FIGURES 8 9. Coprophilus spp., the labels: 8 — C. pentatoma (lectotype, male); 9 — C. pentatoma (paralectotype, female). FIGURES 10 11. Coprophilus spp., the labels: 10 — C. pentatoma (“ Kebin ”, female); 11 — C. longicornis (holotype, male). FIGURES 12 13. Coprophilus spp., the labels: 12 — C. bimaculatus (lectotype, male); 13 — C. rufitarsis (holotype?, female)." httpUri="https://zenodo.org/record/843171/files/figure.png" pageId="1" pageNumber="131">8</figureCitation>
10)
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: 227
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:
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: 329
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<emphasis id="B961EAD6CA1ABA681BECCB1809CAFDF9" box="[151,390,605,627]" italics="true" pageId="1" pageNumber="131">Coprophilus pentatoma</emphasis>
:
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: 89
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<emphasis id="B961EAD6CA1ABA681BECCB3909CAFD18" box="[151,390,636,658]" italics="true" pageId="1" pageNumber="131">Coprophilus pentatoma</emphasis>
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<emphasis id="B961EAD6CA1ABA681BECCBDF09CAFD3A" box="[151,390,666,688]" italics="true" pageId="1" pageNumber="131">Coprophilus pentatoma</emphasis>
:
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: 72
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<emphasis id="B961EAD6CA1ABA681BECCBFC09CAFD45" box="[151,390,697,719]" italics="true" pageId="1" pageNumber="131">Coprophilus pentatoma</emphasis>
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<emphasis id="B961EAD6CA1ABA681BECCB9D0A45FD64" box="[151,521,728,750]" italics="true" pageId="1" pageNumber="131">Coprophilus (Zonoptilus) pentatoma</emphasis>
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: 94
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(
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<emphasis id="B961EAD6CA1ABA681BECCBB30A45FC87" box="[151,521,758,781]" italics="true" pageId="1" pageNumber="131">Coprophilus (Zonoptilus) pentatoma</emphasis>
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<emphasis id="B961EAD6CA1ABA681BECCA500A45FCA1" box="[151,521,789,811]" italics="true" pageId="1" pageNumber="131">Coprophilus (Zonoptilus) pentatoma</emphasis>
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<emphasis id="B961EAD6CA1ABA681BECCA7109CAFCC0" box="[151,390,820,842]" italics="true" pageId="1" pageNumber="131">Coprophilus pentatoma</emphasis>
:
<treatmentCitation id="0AB410D5CA1ABA681AE8CA700A2FFCC0" author="Herman" box="[403,611,820,842]" page="1315" pageId="1" pageNumber="131" year="2001">
<bibRefCitation id="EF844B35CA1ABA681AE8CA700A6BFCC0" author="Herman" box="[403,551,820,842]" pageId="1" pageNumber="131" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1785 to the end of the second millennium. Bulletin of the American Museum of Natural History, 265, 1309 - 1319." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 1315
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA36C4CA1ABA681BECCA170B2DFC0D" blockId="1.[151,923,544,903]" pageId="1" pageNumber="131">
<treatmentCitationGroup id="AB0511EACA1ABA681BECCA170B2DFC0D" pageId="1" pageNumber="131">
<taxonomicName id="4C154D47CA1ABA681BECCA170A92FCE2" ID-CoL="5ZYJ6" authority=": Smetana, 2004: 511" authorityName="Smetana" authorityPageNumber="511" authorityYear="2004" box="[151,734,850,872]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1ABA681BECCA170A4BFCE2" box="[151,519,850,872]" italics="true" pageId="1" pageNumber="131">Coprophilus (Zonyptilus) pentatoma</emphasis>
:
<treatmentCitation id="0AB410D5CA1ABA68196ECA170A92FCE2" author="Smetana" box="[533,734,850,872]" page="511" pageId="1" pageNumber="131" year="2004">
<bibRefCitation id="EF844B35CA1ABA68196ECA170AFCFCE2" author="Smetana" box="[533,688,850,872]" pageId="1" pageNumber="131" refString="Smetana, A. (2004) Oxytelinae. In Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Hydrophiloidea- Histeroidea-Staphylinoidea, 5 (2), 511 - 535. [Apollo Books, Stenstrup]" type="journal article" year="2004">Smetana, 2004</bibRefCitation>
: 511
</treatmentCitation>
</taxonomicName>
<taxonomicName id="4C154D47CA1ABA681BECCA340B2DFC0D" ID-CoL="5ZYJ6" authority=": Schulke and Smetana, 2015: 767" authorityName="Schulke and Smetana" authorityPageNumber="767" authorityYear="2015" box="[151,865,881,903]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1ABA681BECCA340A4BFC0D" box="[151,519,881,903]" italics="true" pageId="1" pageNumber="131">Coprophilus (Zonyptilus) pentatoma</emphasis>
:
<treatmentCitation id="0AB410D5CA1ABA68196ECA340B2DFC0D" author="Schulke" box="[533,865,881,903]" page="767" pageId="1" pageNumber="131" year="2015">
<bibRefCitation id="EF844B35CA1ABA68196ECA340B7EFC0D" author="Schulke" box="[533,818,881,903]" pageId="1" pageNumber="131" refString="Schulke, M. &amp; Smetana, A. (2015) Oxytelinae. In Lobl, I. &amp; Lobl, D. (Ed.), Catalogue of Palaearctic Coleoptera, Hydrophiloidea-Staphylinoidea, 5 (1), pp. 757 - 795. [Brill, Leiden &amp; Boston]" type="journal article" year="2015">Schülke and Smetana, 2015</bibRefCitation>
: 767
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30F654FCA1ABA681BECCAF30C72F94C" pageId="1" pageNumber="131" type="materials_examined">
<paragraph id="8BAA36C4CA1ABA681BECCAF309B0FB0C" blockId="1.[151,1437,947,2022]" pageId="1" pageNumber="131">
<emphasis id="B961EAD6CA1ABA681BECCAF309F2FC45" bold="true" box="[151,446,950,975]" pageId="1" pageNumber="131">Type material examined.</emphasis>
Lectotype ♂ “M ts Alai Turkestan” “Col. et det. A. Fauvel |
<taxonomicName id="4C154D47CA1ABA681F2FCAF30DDCFC45" authority="Fvl." authorityName="Fauvel" authorityYear="1897" box="[1108,1424,950,975]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1ABA681F2FCAF30D10FC45" box="[1108,1372,950,975]" italics="true" pageId="1" pageNumber="131">Coprophilus pentatoma</emphasis>
Fvl.
</taxonomicName>
| R.I.Sc.N.B. 17.479” “Lectotypus ♂ 1990
<taxonomicName id="4C154D47CA1ABA681915CA9E0C7EFC79" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" box="[622,1074,986,1011]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonoptilus">
<emphasis id="B961EAD6CA1ABA681915CA9E0ACBFC78" box="[622,647,987,1010]" italics="true" pageId="1" pageNumber="131">C.</emphasis>
(
<emphasis id="B961EAD6CA1ABA6819E3CA9F0B5CFC79" box="[664,784,986,1011]" italics="true" pageId="1" pageNumber="131">Zonoptilus</emphasis>
)
<emphasis id="B961EAD6CA1ABA68185ACA990BDBFC79" box="[801,919,988,1011]" italics="true" pageId="1" pageNumber="131">pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1ABA6818E4CA9F0C7EFC79" author="Fauvel" box="[927,1074,986,1011]" pageId="1" pageNumber="131" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. dr. Tóth L.” “
<taxonomicName id="4C154D47CA1ABA681E6ACA9F0A02FB90" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1ABA681E6ACA9F09E9FB93" italics="true" pageId="1" pageNumber="131">Coprophilus (Zonyptilus) pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1ABA681ACFCD440A02FB90" author="Fauvel" box="[436,590,1025,1050]" pageId="1" pageNumber="131" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. M. Gildenkov, 2014” (IRSNB). Paralectotypes: 1 ♀ “M t Alai Turkestan” &lt;Col. et det. A. Fauvel |
<taxonomicName id="4C154D47CA1ABA68194ACD600B22FBB4" authority="Fvl." authorityName="Fauvel" authorityYear="1897" box="[561,878,1061,1086]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1ABA68194ACD600B76FBB7" box="[561,826,1061,1086]" italics="true" pageId="1" pageNumber="131">Coprophilus pentatoma</emphasis>
Fvl.
</taxonomicName>
| R.I.Sc.N.B. 17.479” “Paralectotypus ♂ 1990
<taxonomicName id="4C154D47CA1ABA681EF9CD630A71FBE8" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonoptilus">
<emphasis id="B961EAD6CA1ABA681EF9CD630DD0FBB7" box="[1410,1436,1062,1085]" italics="true" pageId="1" pageNumber="131">C.</emphasis>
(
<emphasis id="B961EAD6CA1ABA681BDBCD0C0954FBE8" box="[160,280,1097,1122]" italics="true" pageId="1" pageNumber="131">Zonoptilus</emphasis>
)
<emphasis id="B961EAD6CA1ABA681A51CD0F09ECFBEB" box="[298,416,1098,1121]" italics="true" pageId="1" pageNumber="131">pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1ABA681AD2CD0C0A71FBE8" author="Fauvel" box="[425,573,1097,1122]" pageId="1" pageNumber="131" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. dr. Tóth L.” “
<taxonomicName id="4C154D47CA1ABA681858CD0C0D17FBE8" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" box="[803,1371,1097,1122]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1ABA681858CD0C0BE2FBE8" box="[803,942,1097,1122]" italics="true" pageId="1" pageNumber="131">Coprophilus</emphasis>
(
<emphasis id="B961EAD6CA1ABA6818BBCD0C0C7AFBE8" box="[960,1078,1097,1122]" italics="true" pageId="1" pageNumber="131">Zonyptilus</emphasis>
)
<emphasis id="B961EAD6CA1ABA681F32CD0F0CF3FBEB" box="[1097,1215,1098,1121]" italics="true" pageId="1" pageNumber="131">pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1ABA681FBCCD0C0D17FBE8" author="Fauvel" box="[1223,1371,1097,1122]" pageId="1" pageNumber="131" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. M. Gildenkov, 2014” (IRSNB).
</paragraph>
<paragraph id="8BAA36C4CA1ABA681BBCCDD40C72F94C" blockId="1.[151,1437,947,2022]" pageId="1" pageNumber="131">
<emphasis id="B961EAD6CA1ABA681BBCCDD40A0CFB20" bold="true" box="[199,576,1169,1194]" pageId="1" pageNumber="131">Additional material examined.</emphasis>
1 ♀ “Kebin” “Kebin Turkestan” “Col. et det. A. Fauvel |
<taxonomicName id="4C154D47CA1ABA681E6ACDD4090FFB44" authority="Fvl." authorityName="Fauvel" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1ABA681E6ACDD40941FB44" italics="true" pageId="1" pageNumber="131">Coprophilus pentatoma</emphasis>
Fvl.
</taxonomicName>
| R.I.Sc.N.B. 17.479” “Paralectotypus ♂ 1990
<taxonomicName id="4C154D47CA1ABA68181DCDF30D7FFB44" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" box="[870,1331,1205,1230]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonoptilus">
<emphasis id="B961EAD6CA1ABA68181DCDF30BCCFB47" box="[870,896,1206,1229]" italics="true" pageId="1" pageNumber="131">C.</emphasis>
(
<emphasis id="B961EAD6CA1ABA6818E8CDF00C47FB44" box="[915,1035,1205,1230]" italics="true" pageId="1" pageNumber="131">Zonoptilus</emphasis>
)
<emphasis id="B961EAD6CA1ABA681F65CDF20CD8FB44" box="[1054,1172,1207,1230]" italics="true" pageId="1" pageNumber="131">pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1ABA681FE5CDF00D7FFB44" author="Fauvel" box="[1182,1331,1205,1230]" pageId="1" pageNumber="131" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. dr. Tóth L.” “
<taxonomicName id="4C154D47CA1ABA681A67CD9C0B3DFB78" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" box="[284,881,1241,1266]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonyptilus">
<emphasis id="B961EAD6CA1ABA681A67CD9C0A8AFB7B" box="[284,710,1241,1266]" italics="true" pageId="1" pageNumber="131">Coprophilus (Zonyptilus) pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1ABA6819ADCD9C0B3DFB78" author="Fauvel" box="[726,881,1241,1266]" pageId="1" pageNumber="131" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. M. Gildenkov, 2014” (IRSNB); 1 ♂, Kyrgyzstan, Talass valley “[… illegible] Talass Thal | E.Fischer” “
<taxonomicName id="4C154D47CA1ABA6818EECDB80CFAFA9C" authority="Solsky" authorityName="Solsky" box="[917,1206,1277,1302]" class="Insecta" family="Staphylinidae" genus="Zonoptilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B961EAD6CA1ABA6818EECDB80C12FA9F" box="[917,1118,1277,1302]" italics="true" pageId="1" pageNumber="131">Zonoptilus piceus</emphasis>
Solsky
</taxonomicName>
” (ZMUM); 3 spec., Uzbekistan “Fergana. Skobelev. |
<date id="FFAB1004CA1ABA681936CC640A8AFAB0" box="[589,710,1313,1338]" pageId="1" pageNumber="131" value="1919-08-25">25.VIII.19</date>
. P.Archangelskij” (ZMUM; 1 spec.—cMG); 5 spec. “E KAZAKHSTAN, S Tarbagatay Mts.,
<quantity id="4CED9B21CA1ABA68192FCC030ADFFAD4" box="[596,659,1349,1374]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="131" unit="km" value="4.0">4 km</quantity>
N Alexeevka,
<quantity id="4CED9B21CA1ABA681837CC030BE5FAD7" box="[844,937,1350,1374]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="131" unit="m" value="1000.0">1000 m</quantity>
,
<date id="FFAB1004CA1ABA6818C0CC000C2EFAD4" box="[955,1122,1349,1374]" pageId="1" pageNumber="131" value="2001-06-24" valueMax="2001-06-25" valueMin="2001-06-24">24-25.VI.2001</date>
, leg. W. SCHAWALLER” (SMNS; 1 spec.—cMG); 5 spec. “S. Kazakhstan, Almaty reg., slp. Zailiyskyi Alatau, B. Almatinskoe Lake env. H- 2500,
<date id="FFAB1004CA1ABA681B9BCCC809CBFA2C" box="[224,391,1421,1446]" pageId="1" pageNumber="131" value="2002-06-15" valueMax="2002-06-30" valueMin="2002-06-15">15-30.VI.2002</date>
” (HNHM; 2 spec.—cMG); Kyrgyzstan, Talass valley 1 ♂ “Turkestan occ. Unllis Talass | Fischer” “
<taxonomicName id="4C154D47CA1ABA681A72CCF40A79FA40" authority="Solsky" authorityName="Solsky" authorityYear="1867" box="[265,565,1457,1482]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B961EAD6CA1ABA681A72CCF409AEFA43" box="[265,482,1457,1482]" italics="true" pageId="1" pageNumber="131">Coprophilus piceus</emphasis>
Solsky
</taxonomicName>
| Coll. Reitter” “
<taxonomicName id="4C154D47CA1ABA681989CCF40D68FA40" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" box="[754,1316,1457,1482]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonoptilus">
<emphasis id="B961EAD6CA1ABA681989CCF40CC7FA43" box="[754,1163,1457,1482]" italics="true" pageId="1" pageNumber="131">Coprophilus (Zonoptilus) pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1ABA681FE9CCF40D68FA40" author="Fauvel" box="[1170,1316,1457,1482]" pageId="1" pageNumber="131" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. Tóth L. 1990” (HNHM); Kyrgyzstan, Talass valley 1 ♂ “Talass-Thal WestTurkestan | Fischer” “
<taxonomicName id="4C154D47CA1ABA681FC4CC9008AFF998" authority="Solsky" authorityName="Solsky" authorityYear="1867" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B961EAD6CA1ABA681FC4CC900DD7FA67" box="[1215,1435,1493,1518]" italics="true" pageId="1" pageNumber="131">Coprophilus piceus</emphasis>
Solsky
</taxonomicName>
| Coll. Reitter” “
<taxonomicName id="4C154D47CA1ABA681ADCCCBC0BACF998" authority="Fauvel, 1897" authorityName="Fauvel" authorityYear="1897" box="[423,992,1529,1554]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma" subGenus="Zonoptilus">
<emphasis id="B961EAD6CA1ABA681ADCCCBC0B0FF99B" box="[423,835,1529,1554]" italics="true" pageId="1" pageNumber="131">Coprophilus (Zonoptilus) pentatoma</emphasis>
<bibRefCitation id="EF844B35CA1ABA681837CCBC0BACF998" author="Fauvel" box="[844,992,1529,1554]" pageId="1" pageNumber="131" refString="Fauvel, A. (1897) Tableau des especes palearctiques du genre Coprophilus Latr. (Zonoptilus Mots. - Homalotrichus Sol.). Revue d'Entomologie, 16, 226 - 229." type="journal article" year="1897">Fauvel, 1897</bibRefCitation>
</taxonomicName>
| det. Tóth L. 1990” (HNHM); 3 spec. “
<date id="FFAB1004CA1ABA681BDACF5B096EF9BC" box="[161,290,1566,1590]" pageId="1" pageNumber="131" value="2009-07-29">29.07.2009</date>
| Kazakhstan, Charyn riv., Aktogai | V. Kastcheev” (ZMAS); 1 spec. “
<date id="FFAB1004CA1ABA681F32CF580C85F9BC" box="[1097,1225,1565,1590]" pageId="1" pageNumber="131" value="2005-07-21">21.07.2005</date>
| S-E Kazakhstan, Charyn riv., ur. Sartogai | V. Kastcheev” (ZMAS); 1 spec. “[in Russian]
<date id="FFAB1004CA1ABA6818ACCF040C1AF9D0" box="[983,1110,1601,1626]" pageId="1" pageNumber="131" value="1988-05-15">15.05.1988</date>
| Kazakhstan, hr. Ketmen, r. Dolaity | V. Kastcheev” (ZMAS); 2 spec. “
<date id="FFAB1004CA1ABA6819F5CF230B43F9F4" box="[654,783,1638,1662]" pageId="1" pageNumber="131" value="1987-08-23">23.08.1987</date>
| SE Kazakhstan, Dzhungar mnts., Keskenterek riv. | V. Kastcheev” (ZMAS); 1 spec. “
<date id="FFAB1004CA1ABA681A88CFCC0A3FF928" box="[499,627,1673,1698]" pageId="1" pageNumber="131" value="1986-05-05">05.05.1986</date>
| Kazakhstan, Almaty, Kamenka | V. Kastcheev” (ZMAS); 1 ♂, 1 ♀ “P Fischer | Talass Thal
<date id="FFAB1004CA1ABA681AFFCFE809A2F94C" box="[388,494,1709,1734]" pageId="1" pageNumber="131" value="2007-06-15">15-VI-07</date>
” “Kastch. |
<taxonomicName id="4C154D47CA1ABA68190FCFE80B8FF94C" authority="Fauv." authorityName="Fauvel" authorityYear="1897" box="[628,963,1709,1734]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1ABA68190FCFE80B30F94C" box="[628,892,1709,1734]" italics="true" pageId="1" pageNumber="131">Coprophilus pentatoma</emphasis>
Fauv.
</taxonomicName>
” (ZMAS).
</paragraph>
</subSubSection>
<subSubSection id="C30F654FCA1ABA6B1BBCCF940A7FFE02" lastPageId="2" lastPageNumber="132" pageId="1" pageNumber="131" type="discussion">
<paragraph id="8BAA36C4CA1ABA6B1BBCCF94089AFF5E" blockId="1.[151,1437,947,2022]" lastBlockId="2.[151,1437,151,392]" lastPageId="2" lastPageNumber="132" pageId="1" pageNumber="131">
<emphasis id="B961EAD6CA1ABA681BBCCF940909F960" bold="true" box="[199,325,1745,1770]" pageId="1" pageNumber="131">Discussion</emphasis>
. Type specimens and other material, including males, were examined. The aedeagus has a characteristic structure (
<figureCitation id="132E2A41CA1ABA681ADFCFB00A43F884" box="[420,527,1781,1806]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1254,1277]" captionTargetBox="[157,1411,459,1214]" captionTargetId="figure@2.[151,1436,447,1230]" captionTargetPageId="2" captionText="FIGURES 1 3. Coprophilus (Zonyptilus) pentatoma Fauvel, 1897 (lectotype, male): 1 — body, dorsal view; 2 — aedeagus, lateral view; 3 — aedeagus, ventral view. Scale bar: fig. 1 — 1 mm; figs. 2, 3 — 0.25 mm." httpUri="https://zenodo.org/record/843165/files/figure.png" pageId="1" pageNumber="131">Figs. 23</figureCitation>
) that shows no apparent variation. The females are similar to males in coloration and body shape, but differ by the smaller pronotum. The species is distributed in Tien Shan Mt. range and probably Karakorum Mt. range (cited from
<bibRefCitation id="EF844B35CA1ABA681967CE780AF4F8DC" author="Gridelli" box="[540,696,1853,1878]" pageId="1" pageNumber="131" refString="Gridelli, E. (1934) Materiali zoologici raccolti dalla spedizione italiana al Karakoram (1929 - Anno VII). Coleoptera - Staphylinidae. Atti del Museo Civico di Storia Naturale di Trieste, 12, 69 - 85." type="journal article" year="1934">Gridelli 1934</bibRefCitation>
: 72). In my opinion, the record of the species from Azerbaijan is erroneous. This conclusion is based on the following facts. Fauvel described the species from material in three locations “Turkestan (Monts Alaï), Baku, and Dobrudja”. Tóth revised the type material of
<taxonomicName id="4C154D47CA1ABA681FBACEC30D17F814" authorityName="Fauvel" authorityYear="1897" box="[1217,1371,1926,1950]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1ABA681FBACEC30D17F814" box="[1217,1371,1926,1950]" italics="true" pageId="1" pageNumber="131">C. pentatoma</emphasis>
</taxonomicName>
from IRSNB and designated the lectotype (
<bibRefCitation id="EF844B35CA1ABA681933CEEC0AA2F848" author="Toth" box="[584,750,1961,1986]" pageId="1" pageNumber="131" refString="Toth, L. (1991) New data to the knowledge of the Palearctic Coprophilus Latreille, 1829 species (Coleoptera, Staphylinidae). Annales Historico-naturales Musei Nationalis Hungarici, 83, 91 - 101." type="journal article" year="1991">Tóth 1991: 94</bibRefCitation>
) from Mons Alai. In the same publication,
<bibRefCitation id="EF844B35CA1ABA681F9DCEEC0DD0F848" author="Toth" box="[1254,1436,1961,1986]" pageId="1" pageNumber="131" refString="Toth, L. (1991) New data to the knowledge of the Palearctic Coprophilus Latreille, 1829 species (Coleoptera, Staphylinidae). Annales Historico-naturales Musei Nationalis Hungarici, 83, 91 - 101." type="journal article" year="1991">Tóth (1991: 95)</bibRefCitation>
described one male (syntype of
<taxonomicName id="4C154D47CA1ABA681979CE8B0AD6F86C" authorityName="Fauvel" authorityYear="1897" box="[514,666,1998,2022]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA1ABA681979CE8B0AD6F86C" box="[514,666,1998,2022]" italics="true" pageId="1" pageNumber="131">C. pentatoma</emphasis>
</taxonomicName>
) from Cernavoda, Dobrudja as
<taxonomicName id="4C154D47CA1ABA681F7FCE8B0D64F86C" authority="Toth, 1991" authorityName="Toth" authorityYear="1991" box="[1028,1320,1997,2022]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="131" phylum="Arthropoda" rank="species" species="drugmandi">
<emphasis id="B961EAD6CA1ABA681F7FCE8B0CEDF86C" box="[1028,1185,1997,2022]" italics="true" pageId="1" pageNumber="131">C. drugmandi</emphasis>
<bibRefCitation id="EF844B35CA1ABA681FD2CE880D64F86C" author="Toth" box="[1193,1320,1997,2022]" pageId="1" pageNumber="131" refString="Toth, L. (1991) New data to the knowledge of the Palearctic Coprophilus Latreille, 1829 species (Coleoptera, Staphylinidae). Annales Historico-naturales Musei Nationalis Hungarici, 83, 91 - 101." type="journal article" year="1991">Tóth, 1991</bibRefCitation>
</taxonomicName>
. Also two syntypes of
<taxonomicName id="4C154D47CA19BA6B1A5EC9DD098CFF3A" authorityName="Fauvel" authorityYear="1897" box="[293,448,152,176]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="132" phylum="Arthropoda" rank="species" species="pentatoma">
<emphasis id="B961EAD6CA19BA6B1A5EC9DD098CFF3A" box="[293,448,152,176]" italics="true" pageId="2" pageNumber="132">C. pentatoma</emphasis>
</taxonomicName>
from Baku (
<bibRefCitation id="EF844B35CA19BA6B1922C9D20A98FF3A" author="Toth" box="[601,724,151,176]" pageId="2" pageNumber="132" refString="Toth, L. (1992) Coprophilus (Zonoptilus) fauveli sp. n. from Azerbaidzhan (Coleoptera, Staphylinidae: Oxytelinae). Acta Zoologica Hungarica, 38, 373 - 376." type="journal article" year="1992">Tóth 1992</bibRefCitation>
) were later described as a separate species—
<taxonomicName id="4C154D47CA19BA6B1F9EC9DD089FFF5E" authority="Toth, 1992" authorityName="Toth" authorityYear="1992" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="132" phylum="Arthropoda" rank="species" species="fauveli">
<emphasis id="B961EAD6CA19BA6B1F9EC9DD0D19FF3A" box="[1253,1365,151,176]" italics="true" pageId="2" pageNumber="132">C. fauveli</emphasis>
<bibRefCitation id="EF844B35CA19BA6B1E1BC9D2089FFF5E" author="Toth" pageId="2" pageNumber="132" refString="Toth, L. (1992) Coprophilus (Zonoptilus) fauveli sp. n. from Azerbaidzhan (Coleoptera, Staphylinidae: Oxytelinae). Acta Zoologica Hungarica, 38, 373 - 376." type="journal article" year="1992">Tóth, 1992</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAA36C4CA19BA6B1BBCC99A0A7FFE02" blockId="2.[151,1437,151,392]" pageId="2" pageNumber="132">
<emphasis id="B961EAD6CA19BA6B1BBCC99A0978FF72" bold="true" box="[199,308,223,248]" pageId="2" pageNumber="132">Remarks</emphasis>
.
<bibRefCitation id="EF844B35CA19BA6B1A3FC99A09B6FF72" author="Toth" box="[324,506,223,248]" pageId="2" pageNumber="132" refString="Toth, L. (1991) New data to the knowledge of the Palearctic Coprophilus Latreille, 1829 species (Coleoptera, Staphylinidae). Annales Historico-naturales Musei Nationalis Hungarici, 83, 91 - 101." type="journal article" year="1991">Tóth (1991: 94)</bibRefCitation>
who designated the lectotype and paralectotypes indicated that the lectotype is a male, while the paralectotypes are females; the labels of the paralectotypes (figs. 910), however, state that they are males. The examination of these specimens confirmed that they are females. Moreover, one of the paralectotypes designated by Tóth (1 ♀ “Kebin”) cannot be considered as such, because the original description gives no indication of the locality of “Kebin”.
</paragraph>
</subSubSection>
</treatment>
</document>