446 lines
67 KiB
XML
446 lines
67 KiB
XML
<document id="5605AEFF2D41AD1F87A0A01383C2D5C7" ID-DOI="10.11646/zootaxa.4991.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5042656" ID-ZooBank="F0FAE07E-4A90-4602-BCC5-6D80E158BB6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624989159860" checkinUser="plazi" docAuthor="Suksai, Benjamart, Gustafson, Grey T., Sites, Robert W. & Sangpradub, Narumon" docDate="2021" docId="03BC87C1FFF022500CC07970FA43513A" docLanguage="en" docName="zootaxa.4991.3.8.pdf" docOrigin="Zootaxa 4991 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patrus landaisi Regimbart 1892" docType="key" docVersion="3" lastPageNumber="588" masterDocId="FF85FFB9FFEA224B0C577A23FFDF537D" masterDocTitle="Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)" masterLastPageNumber="591" masterPageNumber="561" pageNumber="587" updateTime="1698978743576" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1F0E880A14B97AA6ECED7A863AB3F082">Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)</mods:title>
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<mods:namePart id="EABC466EB12D61F0488274875B40111D">Suksai, Benjamart</mods:namePart>
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<mods:nameIdentifier id="482F74127F5BA93AFCCE24886658EF9B" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:namePart id="1973F5F80A76EBC53B95C814CEFAAA0D">Gustafson, Grey T.</mods:namePart>
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<mods:affiliation id="F96CC8CDEAC33C7C1162F7273CDF5DFC">Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, USA. & Department of Biological Sciences, Northern Arizona University, Flagstaff, Arizona, 86011, USA.</mods:affiliation>
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<mods:namePart id="8C34E04FCF097CC15775F80C05BAC67B">Sites, Robert W.</mods:namePart>
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<mods:nameIdentifier id="0A43BEF8A9FBB643B1C9F2215E54A234" type="ORCID">0000-0002-3895-813X</mods:nameIdentifier>
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<mods:affiliation id="6DA0E639A530B9127373B0D8C9F7B9A7">Enns Entomology Museum, Division of Plant Science, University of Missouri-Columbia, Missouri 65211, USA. SitesR @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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<mods:namePart id="D428B4B4DFBADB19AFB06B46240DCBC8">Sangpradub, Narumon</mods:namePart>
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<mods:affiliation id="8D0CAE5586485119B19D4A186E8AEF21">Applied Taxonomic Research Center, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand & Corresponding Author: narumon @ kku. ac. th; https: // orcid. org / 0000 - 0003 - 0228 - 8696</mods:affiliation>
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<mods:nameIdentifier id="4502BC7E83F3A5B9FA5A0924F8820689" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:title id="666F52342ACC3E1FFA6004A7BE96C3DC">Zootaxa</mods:title>
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<mods:date id="E1BEDAE16E50F51B22E65D52B3CEF42A">2021</mods:date>
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<mods:number id="CF67DFF152787B60CFD725450CFCCBBD">2021-06-25</mods:number>
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<mods:number id="38D6E22286EDD108401598C1AFC3918E">4991</mods:number>
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<mods:identifier id="83DFF3C6306845B21ECEFCE976A910F1" type="DOI">10.11646/zootaxa.4991.3.8</mods:identifier>
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Key to species of adult
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<taxonomicName id="4C154D54FFF022510DE07970FDB15013" authorityName="Regimbart" authorityYear="1892" box="[439,622,851,878]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="landaisi">
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<emphasis id="B961EAC5FFF022510DE07970FDB15013" bold="true" box="[439,622,851,878]" italics="true" pageId="26" pageNumber="587">Patrus landaisi</emphasis>
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group within
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<collectingCountry id="F3027647FFF022510F497970FC4F5010" box="[798,912,851,877]" name="Thailand" pageId="26" pageNumber="587">Thailand</collectingCountry>
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<keyLead id="30E495E2FFF022510CC079BBFA4250D3" box="[151,1437,920,942]" pageId="26" pageNumber="587">
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1. Protarsus laterally expanded, with setose pad on posterior face.......................................... 2
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<emphasis id="B961EAC5FFF02251090579BBFA4250D3" bold="true" box="[1362,1437,920,942]" pageId="26" pageNumber="587">(males)</emphasis>
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<keyLead id="30E495E2FFF022510CC07995FA4350B6" box="[151,1436,949,971]" pageId="26" pageNumber="587">
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- Protarsus narrow, not laterally expanded, without setose pad on posterior face............................ 9
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<emphasis id="B961EAC5FFF0225109687996FA4350B6" bold="true" box="[1343,1436,949,971]" pageId="26" pageNumber="587">(females)</emphasis>
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<paragraph id="8BAA36D7FFF022510CC079F0FA435778" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
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<keyLead id="30E495E2FFF022510CC079F0FA435778" pageId="26" pageNumber="587">
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2. Elytral glabrous region along suture extending ca. 7/9 length of elytra; elytral apex without apicolateral sinuation; species found at elevations higher than
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a.s.l...................................................................... 3
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</keyLead>
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<paragraph id="8BAA36D7FFF022510CC07E2DFA43573C" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
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<keyLead id="30E495E2FFF022510CC07E2DFA43573C" pageId="26" pageNumber="587">
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- Elytral glabrous region along suture extending ca. 6/7 length of elytra; elytral apex with apicolateral sinuation (
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<figureCitation id="132E2A52FFF0225109737E2EFA4E575E" box="[1316,1425,1037,1059]" captionStart="FIGURE 6" captionStartId="10.[151,260,1117,1143]" captionTargetBox="[180,1401,191,1085]" captionTargetId="figure-25@10.[176,1410,179,1093]" captionTargetPageId="10" captionText="FIGURE 6. Patrus corpulentus: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) female pronotum and elytra, dorsal view, scale bar = 4 mm; (D) gonocoxae, ventral view; (E) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (F) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042674" httpUri="https://zenodo.org/record/5042674/files/figure.png" pageId="26" pageNumber="587">Fig. 6A, C</figureCitation>
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); species found at elevations lower than
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a.s.l..............................
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<taxonomicName id="4C154D54FFF02251087F7E08FA43573C" authority="(Regimbart, 1884)" authorityName="Borneo" baseAuthorityName="Regimbart" baseAuthorityYear="1884" box="[1064,1436,1066,1089]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="corpulentus">
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<emphasis id="B961EAC5FFF02251087F7E08FB3A573D" bold="true" box="[1064,1253,1066,1088]" italics="true" pageId="26" pageNumber="587">Patrus corpulentus</emphasis>
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(
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<bibRefCitation id="EF844B26FFF0225108A67E08FA4A573C" author="Regimbart, M." box="[1265,1429,1067,1089]" pageId="26" pageNumber="587" pagination="381 - 482" refId="ref22884" refString="Regimbart, M. (1884) Essai monographique de la famille des Gyrinidae. 3 e partie. Annales de la Societe entomologique de France, 6, 381 - 482, pls. 311 - 314." type="journal article" year="1884">Régimbart, 1884</bibRefCitation>
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)
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<paragraph id="8BAA36D7FFF022510CC07E6BFA435720" blockId="26.[151,1437,920,2027]" box="[151,1436,1096,1118]" pageId="26" pageNumber="587">
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<keyLead id="30E495E2FFF022510CC07E6BFA435720" box="[151,1436,1096,1118]" pageId="26" pageNumber="587">
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3. Protibia constricted basally, appearing hatchet-shaped (
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<figureCitation id="132E2A52FFF022510EA47E6BFC875723" box="[755,856,1096,1118]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="26" pageNumber="587">Fig. 4F–G</figureCitation>
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).............................................. 4
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</keyLead>
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</paragraph>
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<paragraph id="8BAA36D7FFF022510CC07E45FA435706" blockId="26.[151,1437,920,2027]" box="[151,1436,1125,1147]" pageId="26" pageNumber="587">
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<keyLead id="30E495E2FFF022510CC07E45FA435706" box="[151,1436,1125,1147]" pageId="26" pageNumber="587">
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- Protibia not constricted basally, appearing either wedge- or club-shaped (
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<figureCitation id="132E2A52FFF022510FC87E46FBFD5706" box="[927,1058,1125,1147]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="26" pageNumber="587">Fig. 4A–E, H</figureCitation>
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)............................. 5
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E12E72FFF022510CC07EA0FA435657" pageId="26" pageNumber="587">
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<paragraph id="8BAA36D7FFF022510CC07EA0FA4357AE" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
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<keyLead id="30E495E2FFF022510CC07EA0FA4357AE" pageId="26" pageNumber="587">
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4. Elytral apex weakly rounded in appearance, epipleural angle indistinct; elytral glabrous region laterally expanded in anterior 2/3 to nearly same width as pronotal glabrous region (
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<figureCitation id="132E2A52FFF022510EA17E83FC8F57CB" box="[758,848,1184,1206]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="26" pageNumber="587">Fig. 11A</figureCitation>
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); protibia constricted in basal 1/2 (
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<figureCitation id="132E2A52FFF0225108F07E83FB2957CB" box="[1191,1270,1184,1206]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="26" pageNumber="587">Fig. 4G</figureCitation>
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); median lobe of aedeagus constricted medially (
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<figureCitation id="132E2A52FFF022510E477E9EFDBA57AE" box="[528,613,1213,1235]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="26" pageNumber="587">Fig. 11B</figureCitation>
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)...........................................
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<taxonomicName id="4C154D54FFF0225108D47E9DFA4357AE" authority="(Ochs, 1940)" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[1155,1436,1213,1235]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="salvazai">
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<emphasis id="B961EAC5FFF0225108D47E9DFAC657AE" bold="true" box="[1155,1305,1213,1235]" italics="true" pageId="26" pageNumber="587">Patrus salvazai</emphasis>
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(
|
||
<bibRefCitation id="EF844B26FFF0225109717E9EFA4A57AE" author="Ochs" box="[1318,1429,1213,1235]" pageId="26" pageNumber="587" pagination="1 - 22" refId="ref22563" refString="Ochs, G. (1940) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Gyrinidae, gesammelt von Rene Malaise. Arkiv for Zoologi, 32 A, 1 - 22." type="journal article" year="1940">Ochs, 1940</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF022510CC07EF8FA435657" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07EF8FA435657" pageId="26" pageNumber="587">
|
||
- Elytral apex obliquely truncate, epipleural angle distinct; elytral glabrous region laterally expanded in anterior 2/3 to wider than pronotal glabrous region (
|
||
<figureCitation id="132E2A52FFF022510E5A7EDBFDB85673" box="[525,615,1272,1294]" captionStart="FIGURE 10" captionStartId="20.[151,260,1837,1863]" captionTargetBox="[160,1431,892,1805]" captionTargetId="figure-325@20.[151,1436,882,1813]" captionTargetPageId="20" captionText="FIGURE 10. Patrus phetchabunensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042684" httpUri="https://zenodo.org/record/5042684/files/figure.png" pageId="26" pageNumber="587">Fig. 10A</figureCitation>
|
||
); protibia constricted in basal 2/5 (
|
||
<figureCitation id="132E2A52FFF022510FEF7EDBFBDD5673" box="[952,1026,1272,1294]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="26" pageNumber="587">Fig. 4F</figureCitation>
|
||
); median lobe of aedeagus without medial constriction (
|
||
<figureCitation id="132E2A52FFF022510D377F36FE695656" box="[352,438,1301,1323]" captionStart="FIGURE 10" captionStartId="20.[151,260,1837,1863]" captionTargetBox="[160,1431,892,1805]" captionTargetId="figure-325@20.[151,1436,882,1813]" captionTargetPageId="20" captionText="FIGURE 10. Patrus phetchabunensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042684" httpUri="https://zenodo.org/record/5042684/files/figure.png" pageId="26" pageNumber="587">Fig. 10B</figureCitation>
|
||
).......................................................
|
||
<emphasis id="B961EAC5FFF02251080C7F35FA435657" bold="true" box="[1115,1436,1301,1323]" pageId="26" pageNumber="587">
|
||
<taxonomicName id="4C154D54FFF02251080C7F35FA955656" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[1115,1354,1301,1323]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="phetchabunensis" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFF02251080C7F35FA955656" bold="true" box="[1115,1354,1301,1323]" italics="true" pageId="26" pageNumber="587">Patrus phetchabunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25257BEFFF0225109077F36FA435657" box="[1360,1436,1301,1322]" pageId="26" pageNumber="587" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF022510CC07F10FA4356DC" pageId="26" pageNumber="587">
|
||
<paragraph id="8BAA36D7FFF022510CC07F10FA435618" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07F10FA435618" pageId="26" pageNumber="587">
|
||
5. Elytral apex rounded, epipleural angle indistinct (
|
||
<figureCitation id="132E2A52FFF022510EE97F10FCD45634" box="[702,779,1331,1353]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="26" pageNumber="587">Fig. 7A</figureCitation>
|
||
); body form in lateral view strongly dorsoventrally convex........................................................................................
|
||
<emphasis id="B961EAC5FFF0225108EC7F73FA435618" bold="true" box="[1211,1436,1360,1382]" pageId="26" pageNumber="587">
|
||
<taxonomicName id="4C154D54FFF0225108EC7F73FA95561B" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[1211,1354,1360,1382]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="garuda" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFF0225108EC7F73FA95561B" bold="true" box="[1211,1354,1360,1382]" italics="true" pageId="26" pageNumber="587">Patrus garuda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25257BEFFF0225109077F73FA435618" box="[1360,1436,1360,1381]" pageId="26" pageNumber="587" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF022510CC07F4DFA4356DC" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07F4DFA4356DC" pageId="26" pageNumber="587">
|
||
- Elytral apex obliquely truncate, epipleural angle distinct (
|
||
<figureCitation id="132E2A52FFF022510EAF7F4EFC9356FE" box="[760,844,1389,1411]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="26" pageNumber="587">Figs. 5A</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFF022510F047F4EFCAD56FF" box="[851,882,1389,1410]" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="26" pageNumber="587">8A</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFF022510F2E7F4EFC4656FF" box="[889,921,1389,1410]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="26" pageNumber="587">9A</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFF022510FF77F4EFC1256FE" box="[928,973,1389,1411]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="26" pageNumber="587">12A</figureCitation>
|
||
); body form in lateral view weakly dorsoventrally convex............................................................................................. 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF022510CC07F8BFA435685" pageId="26" pageNumber="587">
|
||
<paragraph id="8BAA36D7FFF022510CC07F8BFA4356A6" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07F8BFA4356A6" pageId="26" pageNumber="587">
|
||
6. Protibia club-shaped (
|
||
<figureCitation id="132E2A52FFF022510DE27F8BFDDD56C3" box="[437,514,1448,1470]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="26" pageNumber="587">Fig. 4D</figureCitation>
|
||
); median lobe of aedeagus without strongly constricted apex (
|
||
<figureCitation id="132E2A52FFF0225108017F8BFB4056C3" box="[1110,1183,1448,1470]" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="26" pageNumber="587">Fig. 8B</figureCitation>
|
||
)...........................................................................................
|
||
<taxonomicName id="4C154D54FFF0225108187FE5FA4356A6" authority="(Regimbart, 1892)" baseAuthorityName="Regimbart" baseAuthorityYear="1892" box="[1103,1436,1477,1499]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="landaisi">
|
||
<emphasis id="B961EAC5FFF0225108187FE5FB3A56A6" bold="true" box="[1103,1253,1477,1499]" italics="true" pageId="26" pageNumber="587">Patrus landaisi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF844B26FFF0225108A67FE6FA4A56A6" author="Regimbart, M." box="[1265,1429,1477,1499]" pageId="26" pageNumber="587" pagination="663 - 752" refId="ref23063" refString="Regimbart, M. (1892) Essai monographique de la famille des Gyrinidae. 2 e supplement. Annales de la Societe entomologique de France, 60 (1891), 663 - 752, pls. 618 - 619." type="journal article" year="1892">Régimbart, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF022510CC07FC0FA435685" blockId="26.[151,1437,920,2027]" box="[151,1436,1507,1529]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07FC0FA435685" box="[151,1436,1507,1529]" pageId="26" pageNumber="587">
|
||
- Protibia wedge-shaped (
|
||
<figureCitation id="132E2A52FFF022510D9D7FC0FD905685" box="[458,591,1507,1529]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="26" pageNumber="587">Fig. 4A, E, H</figureCitation>
|
||
); median lobe of aedeagus with strongly constricted apex...................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF022510CC07C23FA435510" pageId="26" pageNumber="587">
|
||
<paragraph id="8BAA36D7FFF022510CC07C23FA43554F" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07C23FA43554F" pageId="26" pageNumber="587">
|
||
7. Protarsus ca. 1/2 the length of the protibia, protarsomeres width gradually decreasing apically (
|
||
<figureCitation id="132E2A52FFF0225108F67C23FB2F556B" box="[1185,1264,1536,1558]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="26" pageNumber="587">Fig. 4E</figureCitation>
|
||
); median lobe of aedeagus with needle-like apex (
|
||
<figureCitation id="132E2A52FFF022510E4B7C3EFDBA554E" box="[540,613,1565,1587]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="26" pageNumber="587">Fig. 9B</figureCitation>
|
||
)..............................................
|
||
<emphasis id="B961EAC5FFF0225108FD7C3DFA43554F" bold="true" box="[1194,1436,1565,1587]" pageId="26" pageNumber="587">
|
||
<taxonomicName id="4C154D54FFF0225108FD7C3DFA96554F" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[1194,1353,1565,1587]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFF0225108FD7C3DFA96554F" bold="true" box="[1194,1353,1565,1587]" italics="true" pageId="26" pageNumber="587">Patrus nanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25257BEFFF0225109077C3EFA43554F" box="[1360,1436,1565,1586]" pageId="26" pageNumber="587" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF022510CC07C18FA435510" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07C18FA435510" pageId="26" pageNumber="587">
|
||
- Protarsus> 1/2 length of protibia, protarsomeres width not gradually decreasing apically (
|
||
<figureCitation id="132E2A52FFF0225108607C18FB7F552D" box="[1079,1184,1595,1617]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="26" pageNumber="587">Fig. 4A, H</figureCitation>
|
||
); median lobe of aedeagus with or without needle-like apex......................................................................... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF022510CC07C56FA4355B8" pageId="26" pageNumber="587">
|
||
<paragraph id="8BAA36D7FFF022510CC07C56FA4355F6" blockId="26.[151,1437,920,2027]" box="[151,1436,1653,1675]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07C56FA4355F6" box="[151,1436,1653,1675]" pageId="26" pageNumber="587">
|
||
8. Mesoventrite with median pit; median lobe of aedeagus with needle-like apex (
|
||
<figureCitation id="132E2A52FFF022510F8D7C56FBFC55F6" box="[986,1059,1653,1675]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="26" pageNumber="587">Fig. 5B</figureCitation>
|
||
)..
|
||
<taxonomicName id="4C154D54FFF0225108057C55FA4355F6" authority="(Regimbart, 1891)" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[1106,1436,1653,1675]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="apicalis">
|
||
<emphasis id="B961EAC5FFF0225108057C55FB3A55F6" bold="true" box="[1106,1253,1653,1675]" italics="true" pageId="26" pageNumber="587">Patrus apicalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF844B26FFF0225108A67C56FA4A55F6" author="Regimbart, M." box="[1265,1429,1653,1675]" pageId="26" pageNumber="587" pagination="537 - 554" refId="ref23003" refString="Regimbart, M. (1891) Viaggio di Leonardo Fea in Birmania e regioni vicine. XXXIII. Enumeration des Dytiscidae et Gyrinidae recueillis par M. Leonardo Fea, dans ses voyages en Birmanie et regions voisines. Annali del Museo Civico di Storia Naturale Giacomo Doria Genova, 10 (2), 537 - 554." type="journal article" year="1891">Régimbart, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF022510CC07CB0FA4355B8" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07CB0FA4355B8" pageId="26" pageNumber="587">
|
||
- Mesoventrite without median pit; median lobe of aedeagus without needle-like apex, apex triangular (
|
||
<figureCitation id="132E2A52FFF0225108E47CB0FAD655D4" box="[1203,1289,1683,1705]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="26" pageNumber="587">Fig. 12B</figureCitation>
|
||
)............................................................................
|
||
<taxonomicName id="4C154D54FFF0225108557C93FAE455BB" authority="(Ochs, 1930)" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[1026,1339,1712,1734]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="subapicalis" status="stat. nov.">
|
||
<emphasis id="B961EAC5FFF0225108557C93FB6755BB" bold="true" box="[1026,1208,1712,1734]" italics="true" pageId="26" pageNumber="587">Patrus subapicalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF844B26FFF0225108937C93FAEB55BB" author="Ochs" box="[1220,1332,1712,1734]" pageId="26" pageNumber="587" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs, 1930</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B961EAC5FFF0225109167C93FA4355B8" bold="true" box="[1345,1436,1712,1733]" pageId="26" pageNumber="587">
|
||
<taxonomicNameLabel id="A25257BEFFF0225109167C93FA4355B8" box="[1345,1436,1712,1733]" pageId="26" pageNumber="587">stat. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF022510CC07CEEFA43540B" pageId="26" pageNumber="587">
|
||
<paragraph id="8BAA36D7FFF022510CC07CEEFA435460" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07CEEFA435460" pageId="26" pageNumber="587">
|
||
9. Elytral glabrous region along suture extending ca. 7/9 length of elytra, lateral pubescence convex posteriorly towards suture in association with medial acumination of glabrous region; elytral apex without apicolateral sinuation (
|
||
<figureCitation id="132E2A52FFF0225108FA7CC8FAE9547C" box="[1197,1334,1771,1793]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="26" pageNumber="587">Fig. 3A, C–H</figureCitation>
|
||
). Found at elevations higher than 450 meters....................................................................... 10
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF022510CC07D05FA43540B" blockId="26.[151,1437,920,2027]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07D05FA43540B" pageId="26" pageNumber="587">
|
||
- Elytral glabrous region along suture extending ca. 6/7 length of elytra, lateral pubescence without convex posterior region extending towards suture, instead concave throughout (
|
||
<figureCitation id="132E2A52FFF022510EAE7D60FC9A5424" box="[761,837,1859,1881]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="26" pageNumber="587">Fig. 3B</figureCitation>
|
||
); elytral apex with apicolateral sinuation. Found at elevations lower than
|
||
<quantity id="4CED9B32FFF022510D197D43FE545408" box="[334,395,1888,1910]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="26" pageNumber="587" unit="m" value="450.0">450 m</quantity>
|
||
a.s.l....................................................
|
||
<taxonomicName id="4C154D54FFF02251087F7D43FA43540B" authority="(Regimbart, 1884)" authorityName="Borneo" baseAuthorityName="Regimbart" baseAuthorityYear="1884" box="[1064,1436,1888,1910]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="corpulentus">
|
||
<emphasis id="B961EAC5FFF02251087F7D43FB3A540B" bold="true" box="[1064,1253,1888,1910]" italics="true" pageId="26" pageNumber="587">Patrus corpulentus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF844B26FFF0225108A67D43FA4A540B" author="Regimbart, M." box="[1265,1429,1888,1910]" pageId="26" pageNumber="587" pagination="381 - 482" refId="ref22884" refString="Regimbart, M. (1884) Essai monographique de la famille des Gyrinidae. 3 e partie. Annales de la Societe entomologique de France, 6, 381 - 482, pls. 311 - 314." type="journal article" year="1884">Régimbart, 1884</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF022510CC07D5EFA4354CD" pageId="26" pageNumber="587">
|
||
<paragraph id="8BAA36D7FFF022510CC07D5EFA4354EF" blockId="26.[151,1437,920,2027]" box="[151,1436,1917,1939]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07D5EFA4354EF" box="[151,1436,1917,1939]" pageId="26" pageNumber="587">
|
||
10. Elytral glabrous region cordiform (
|
||
<figureCitation id="132E2A52FFF022510E657D5EFD4854EE" box="[562,663,1917,1939]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="26" pageNumber="587">Fig. 3F–G</figureCitation>
|
||
)............................................................. 11
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF022510CC07DB8FA4354CD" blockId="26.[151,1437,920,2027]" box="[151,1436,1947,1969]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07DB8FA4354CD" box="[151,1436,1947,1969]" pageId="26" pageNumber="587">
|
||
- Elytral glabrous region not cordiform (
|
||
<figureCitation id="132E2A52FFF022510E007DB8FD2954CD" box="[599,758,1947,1969]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="26" pageNumber="587">Fig. 3A, C–E, H</figureCitation>
|
||
)..................................................... 12
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF022510CC07D9BFA435497" pageId="26" pageNumber="587">
|
||
<paragraph id="8BAA36D7FFF022510CC07D9BFA4354B3" blockId="26.[151,1437,920,2027]" box="[151,1436,1976,1998]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07D9BFA4354B3" box="[151,1436,1976,1998]" pageId="26" pageNumber="587">
|
||
11. Elytral apex weakly rounded in appearance, epipleural angle indistinct (
|
||
<figureCitation id="132E2A52FFF022510FC47D9BFC3954B3" box="[915,998,1976,1998]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="26" pageNumber="587">Fig. 11E</figureCitation>
|
||
)............
|
||
<taxonomicName id="4C154D54FFF0225108D47D9BFA4354B3" authority="(Ochs, 1940)" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[1155,1436,1976,1998]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="salvazai">
|
||
<emphasis id="B961EAC5FFF0225108D47D9BFAC654B3" bold="true" box="[1155,1305,1976,1998]" italics="true" pageId="26" pageNumber="587">Patrus salvazai</emphasis>
|
||
(
|
||
<bibRefCitation id="EF844B26FFF0225109717D9BFA4A54B3" author="Ochs" box="[1318,1429,1976,1998]" pageId="26" pageNumber="587" pagination="1 - 22" refId="ref22563" refString="Ochs, G. (1940) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Gyrinidae, gesammelt von Rene Malaise. Arkiv for Zoologi, 32 A, 1 - 22." type="journal article" year="1940">Ochs, 1940</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF022510CC07DF5FA435497" blockId="26.[151,1437,920,2027]" box="[151,1436,2005,2027]" pageId="26" pageNumber="587">
|
||
<keyLead id="30E495E2FFF022510CC07DF5FA435497" box="[151,1436,2005,2027]" pageId="26" pageNumber="587">
|
||
- Elytral apex obliquely truncate, epipleural angle distinct, weakly rounded (
|
||
<figureCitation id="132E2A52FFF022510FE77DF6FBDA5496" box="[944,1029,2005,2027]" captionStart="FIGURE 10" captionStartId="20.[151,260,1837,1863]" captionTargetBox="[160,1431,892,1805]" captionTargetId="figure-325@20.[151,1436,882,1813]" captionTargetPageId="20" captionText="FIGURE 10. Patrus phetchabunensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042684" httpUri="https://zenodo.org/record/5042684/files/figure.png" pageId="26" pageNumber="587">Fig. 10E</figureCitation>
|
||
).....
|
||
<emphasis id="B961EAC5FFF02251080C7DF5FA435497" bold="true" box="[1115,1436,2005,2027]" pageId="26" pageNumber="587">
|
||
<taxonomicName id="4C154D54FFF02251080C7DF5FA955496" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[1115,1354,2005,2027]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="587" phylum="Arthropoda" rank="species" species="phetchabunensis" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFF02251080C7DF5FA955496" bold="true" box="[1115,1354,2005,2027]" italics="true" pageId="26" pageNumber="587">Patrus phetchabunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25257BEFFF0225109077DF6FA435497" box="[1360,1436,2005,2026]" pageId="26" pageNumber="587" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF122500CC07ABBFA435278" pageId="27" pageNumber="588">
|
||
<paragraph id="8BAA36D7FFF122500CC07ABBFA4353B7" blockId="27.[151,1437,151,584]" pageId="27" pageNumber="588">
|
||
<keyLead id="30E495E2FFF122500CC07ABBFA4353B7" pageId="27" pageNumber="588">
|
||
12. Elytral apex rounded, epiplural angle indistinct, elytral glabrous region “nycteriform” (batwing-shaped) (
|
||
<figureCitation id="132E2A52FFF1225008907AB4FACD53D3" box="[1223,1298,151,174]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="27" pageNumber="588">Fig. 3C</figureCitation>
|
||
); reproductive tract with additional sclerotized structure between gonocoxae (
|
||
<figureCitation id="132E2A52FFF122500F6B7A96FC5B53B7" box="[828,900,181,203]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="27" pageNumber="588">Fig. 7F</figureCitation>
|
||
)........................
|
||
<emphasis id="B961EAC5FFF1225008EC7A96FA4353B7" bold="true" box="[1211,1436,180,202]" pageId="27" pageNumber="588">
|
||
<taxonomicName id="4C154D54FFF1225008EC7A96FA9553B7" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[1211,1354,180,202]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="588" phylum="Arthropoda" rank="species" species="garuda" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFF1225008EC7A96FA9553B7" bold="true" box="[1211,1354,180,202]" italics="true" pageId="27" pageNumber="588">Patrus garuda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25257BEFFF1225009077A96FA4353B7" box="[1360,1436,181,202]" pageId="27" pageNumber="588" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF122500CC07AF0FA435278" blockId="27.[151,1437,151,584]" pageId="27" pageNumber="588">
|
||
<keyLead id="30E495E2FFF122500CC07AF0FA435278" pageId="27" pageNumber="588">- Elytral apex obliquely truncate, epiplural angle distinct, reproductive tract without additional sclerotized structure between gonocoxae......................................................................................... 13</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF122500CC07B2EFA435223" pageId="27" pageNumber="588">
|
||
<paragraph id="8BAA36D7FFF122500CC07B2EFA43525F" blockId="27.[151,1437,151,584]" box="[151,1436,269,291]" pageId="27" pageNumber="588">
|
||
<keyLead id="30E495E2FFF122500CC07B2EFA43525F" box="[151,1436,269,291]" pageId="27" pageNumber="588">
|
||
13. Elytral glabrous region with posterolateral corners produced into points (
|
||
<figureCitation id="132E2A52FFF122500FC97B2EFBFF525F" box="[926,1056,269,291]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="27" pageNumber="588">Fig. 3A, E, H</figureCitation>
|
||
)............................ 14
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF122500CC07B08FA435223" blockId="27.[151,1437,151,584]" pageId="27" pageNumber="588">
|
||
<keyLead id="30E495E2FFF122500CC07B08FA435223" pageId="27" pageNumber="588">
|
||
- Elytral glabrous region with posterolateral corners squared-off, not produced posteriorly (
|
||
<figureCitation id="132E2A52FFF1225008697B09FB56523D" box="[1086,1161,298,320]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="27" pageNumber="588">Fig. 3D</figureCitation>
|
||
).............................................................................................
|
||
<taxonomicName id="4C154D54FFF1225008187B6BFA435223" authority="(Regimbart, 1892)" baseAuthorityName="Regimbart" baseAuthorityYear="1892" box="[1103,1436,327,350]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="588" phylum="Arthropoda" rank="species" species="landaisi">
|
||
<emphasis id="B961EAC5FFF1225008187B6BFB3A5220" bold="true" box="[1103,1253,327,349]" italics="true" pageId="27" pageNumber="588">Patrus landaisi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF844B26FFF1225008A67B64FA4A5223" author="Regimbart, M." box="[1265,1429,327,350]" pageId="27" pageNumber="588" pagination="663 - 752" refId="ref23063" refString="Regimbart, M. (1892) Essai monographique de la famille des Gyrinidae. 2 e supplement. Annales de la Societe entomologique de France, 60 (1891), 663 - 752, pls. 618 - 619." type="journal article" year="1892">Régimbart, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF122500CC07B46FA4352AE" pageId="27" pageNumber="588">
|
||
<paragraph id="8BAA36D7FFF122500CC07B46FA4352EA" blockId="27.[151,1437,151,584]" pageId="27" pageNumber="588">
|
||
<keyLead id="30E495E2FFF122500CC07B46FA4352EA" pageId="27" pageNumber="588">
|
||
14. Glabrous region of elytra narrower, occupying 9/11 elytral width; posterior lateral pubescence of elytra unevenly convex towards suture, extending obliquely with a subtruncate appearance (
|
||
<figureCitation id="132E2A52FFF122500F387BA1FC1E52EA" box="[879,961,386,408]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="27" pageNumber="588">Figs. 3E</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFF122500F9B7BA1FC3852E5" box="[972,999,386,408]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="27" pageNumber="588">9E</figureCitation>
|
||
)..............
|
||
<emphasis id="B961EAC5FFF1225008FD7BA1FA4352EA" bold="true" box="[1194,1436,386,407]" pageId="27" pageNumber="588">
|
||
<taxonomicName id="4C154D54FFF1225008FD7BA1FA9652EA" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[1194,1353,386,407]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="588" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFF1225008FD7BA1FA9652EA" bold="true" box="[1194,1353,386,407]" italics="true" pageId="27" pageNumber="588">Patrus nanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25257BEFFF1225009077BA1FA4352EA" box="[1360,1436,386,407]" pageId="27" pageNumber="588" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF122500CC07B83FA4352AE" blockId="27.[151,1437,151,584]" pageId="27" pageNumber="588">
|
||
<keyLead id="30E495E2FFF122500CC07B83FA4352AE" pageId="27" pageNumber="588">
|
||
- Glabrous region of elytra broader, occupying 10/11 elytral width; posterior margin of elytral glabrous region evenly convex towards suture (
|
||
<figureCitation id="132E2A52FFF122500D2B7B9EFE3B52AF" box="[380,484,445,467]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="27" pageNumber="588">Fig. 3A, H</figureCitation>
|
||
)............................................................................ 15
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E72FFF122500CC07BF9FA43513A" pageId="27" pageNumber="588">
|
||
<paragraph id="8BAA36D7FFF122500CC07BF9FA435173" blockId="27.[151,1437,151,584]" pageId="27" pageNumber="588">
|
||
<keyLead id="30E495E2FFF122500CC07BF9FA435173" pageId="27" pageNumber="588">
|
||
15. In lateral view, width of elytral pubescent margin at the humeral region nearly equal to that extending posterior; in dorsal view, lateral margin of elytral glabrous region more arcuate in appearance (
|
||
<figureCitation id="132E2A52FFF122500F2C7BD4FC1C5170" box="[891,963,503,525]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="27" pageNumber="588">Fig. 5E</figureCitation>
|
||
)..........
|
||
<taxonomicName id="4C154D54FFF1225008057BDBFA435173" authority="(Regimbart, 1891)" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[1106,1436,503,526]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="588" phylum="Arthropoda" rank="species" species="apicalis">
|
||
<emphasis id="B961EAC5FFF1225008057BDBFB3A5170" bold="true" box="[1106,1253,503,525]" italics="true" pageId="27" pageNumber="588">Patrus apicalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF844B26FFF1225008A67BD4FA4A5173" author="Regimbart, M." box="[1265,1429,503,526]" pageId="27" pageNumber="588" pagination="537 - 554" refId="ref23003" refString="Regimbart, M. (1891) Viaggio di Leonardo Fea in Birmania e regioni vicine. XXXIII. Enumeration des Dytiscidae et Gyrinidae recueillis par M. Leonardo Fea, dans ses voyages en Birmanie et regions voisines. Annali del Museo Civico di Storia Naturale Giacomo Doria Genova, 10 (2), 537 - 554." type="journal article" year="1891">Régimbart, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFF122500CC07836FA43513A" blockId="27.[151,1437,151,584]" pageId="27" pageNumber="588">
|
||
<keyLead id="30E495E2FFF122500CC07836FA43513A" pageId="27" pageNumber="588">
|
||
- In lateral view, width of elytral pubescent margin at the humeral region twice that extending posterior; in dorsal view, lateral margin of elytral glabrous region less arcuate in appearance (
|
||
<figureCitation id="132E2A52FFF122500F7B7811FC5F5135" box="[812,896,562,584]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="27" pageNumber="588">Fig. 12E</figureCitation>
|
||
)........
|
||
<taxonomicName id="4C154D54FFF1225008557811FAE45135" authority="(Ochs, 1930)" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[1026,1339,562,584]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="588" phylum="Arthropoda" rank="species" species="subapicalis" status="stat. nov.">
|
||
<emphasis id="B961EAC5FFF1225008557811FB675135" bold="true" box="[1026,1208,562,584]" italics="true" pageId="27" pageNumber="588">Patrus subapicalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF844B26FFF1225008937811FAEB5135" author="Ochs" box="[1220,1332,562,584]" pageId="27" pageNumber="588" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs, 1930</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B961EAC5FFF1225009167811FA43513A" bold="true" box="[1345,1436,562,583]" pageId="27" pageNumber="588">
|
||
<taxonomicNameLabel id="A25257BEFFF1225009167811FA43513A" box="[1345,1436,562,583]" pageId="27" pageNumber="588">stat. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |