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<mods:title id="ED65C0C723E962710AB6A1C45222103D">The genus Polistes Latreille, 1802 (Hymenoptera: Vespidae) of Brazil: morphological diagnoses, identification key, and geographical distribution</mods:title>
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<mods:namePart id="B30B73399EB5A516AE9E2325112CB711">Brito, Samanta</mods:namePart>
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<mods:namePart id="CF8CD3471B78BCB5699469F3B1303A2C">Carpenter, James M.</mods:namePart>
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<paragraph id="8BAA36A6FFB8FFF4CCEEF8E649C7F880" blockId="13.[151,695,1819,1881]" box="[151,695,1819,1846]" pageId="13" pageNumber="522">
<heading id="D0E281CAFFB8FFF4CCEEF8E649C7F880" bold="true" box="[151,695,1819,1846]" fontSize="11" level="1" pageId="13" pageNumber="522" reason="1">
<taxonomicName id="4C154D25FFB8FFF4CCEEF8E649C7F880" ID-CoL="4KY7S" authority="Haliday, 1836" authorityName="Haliday" authorityYear="1836" box="[151,695,1819,1846]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="522" phylum="Arthropoda" rank="species" species="actaeon" subGenus="Epicnemius">
<emphasis id="B961EAB4FFB8FFF4CCEEF8E649C7F880" bold="true" box="[151,695,1819,1846]" pageId="13" pageNumber="522">
<emphasis id="B961EAB4FFB8FFF4CCEEF8E64B84F883" bold="true" box="[151,244,1819,1846]" italics="true" pageId="13" pageNumber="522">Polistes</emphasis>
(
<emphasis id="B961EAB4FFB8FFF4CD7CF8E64AE4F880" bold="true" box="[261,404,1819,1845]" italics="true" pageId="13" pageNumber="522">Epicnemius</emphasis>
)
<emphasis id="B961EAB4FFB8FFF4CDDFF8E64975F880" bold="true" box="[422,517,1819,1845]" italics="true" pageId="13" pageNumber="522">actaeon</emphasis>
<bibRefCitation id="EF844B57FFB8FFF4CE75F8E649C7F880" author="Haliday, A. H." box="[524,695,1819,1845]" pageId="13" pageNumber="522" pagination="316 - 331" refId="ref26860" refString="Haliday, A. H. (1836) Descriptions of Hymenoptera collected by Capt. King in the survey of Straits of Magellan. Transactions of the Linnaean Society. London, 17, 316 - 331." type="journal article" year="1836">Haliday, 1836</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BAA36A6FFB8FFF4CCEEF8C24A57F8EC" blockId="13.[151,695,1819,1881]" box="[151,295,1855,1881]" pageId="13" pageNumber="522">
(
<figureCitation id="132E2A23FFB8FFF4CCE6F8C24A6FF8EC" box="[159,287,1855,1881]" captionStart="FIGURES 3744" captionStartId="15.[152,270,1896,1920]" captionTargetBox="[158,1428,189,1865]" captionTargetId="figure-17@15.[151,1436,181,1872]" captionText="FIGURES 3744. Polistes spp. 37, 38. P. (Epicnemius) actaeon Haliday. 39, 40. P. (E.) angulinus Richards. 41, 42. P. (E.) atrox Richards. 43, 44. P. (E.) bequaertianus Willink. 37, 39, 41, 43. Dorsal view. 38, 40, 42, 44. Lateral view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13736639" httpUri="https://zenodo.org/record/13736639/files/figure.png" pageId="13" pageNumber="522">Figs 37, 38</figureCitation>
)
</paragraph>
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<subSubSection id="C30F652DFFB8FFF4CCEEF880495BF821" box="[151,555,1917,1941]" pageId="13" pageNumber="522" type="reference_group">
<paragraph id="8BAA36A6FFB8FFF4CCEEF880495BF821" blockId="13.[151,555,1917,1941]" box="[151,555,1917,1941]" pageId="13" pageNumber="522">
<treatmentCitationGroup id="AB051188FFB8FFF4CCEEF880495BF821" box="[151,555,1917,1941]" pageId="13" pageNumber="522">
<treatmentCitation id="0AB410B7FFB8FFF4CCEEF8804978F820" author="Haliday, A. H." box="[151,520,1917,1941]" page="323" pageId="13" pageNumber="522" year="1836">
<taxonomicName id="4C154D25FFB8FFF4CCEEF8804978F820" ID-CoL="4KY7S" authority="Haliday, 1836: 323" authorityName="Haliday" authorityPageNumber="323" authorityYear="1836" box="[151,520,1917,1941]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="522" phylum="Arthropoda" rank="species" species="actaeon">
<emphasis id="B961EAB4FFB8FFF4CCEEF8804A4EF820" box="[151,318,1917,1941]" italics="true" pageId="13" pageNumber="522">Polistes actaeon</emphasis>
<bibRefCitation id="EF844B57FFB8FFF4CD3DF8804978F820" author="Haliday, A. H." box="[324,520,1917,1941]" pageId="13" pageNumber="522" pagination="316 - 331" refId="ref26860" refString="Haliday, A. H. (1836) Descriptions of Hymenoptera collected by Capt. King in the survey of Straits of Magellan. Transactions of the Linnaean Society. London, 17, 316 - 331." type="journal article" year="1836">Haliday, 1836: 323</bibRefCitation>
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,
<specimenCount id="9D13FD2FFFB8FFF4CE6BF8804956F821" box="[530,550,1917,1940]" count="1" pageId="13" pageNumber="522" type="female"></specimenCount>
.
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</paragraph>
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<subSubSection id="C30F652DFFB8FFF7CCEEF84148A2FE3C" lastPageId="14" lastPageNumber="523" pageId="13" pageNumber="522" type="diagnosis">
<paragraph id="8BAA36A6FFB8FFF7CCEEF84148A2FE3C" blockId="13.[151,1437,1980,2043]" lastBlockId="14.[151,1437,151,573]" lastPageId="14" lastPageNumber="523" pageId="13" pageNumber="522">
<emphasis id="B961EAB4FFB8FFF4CCEEF8414A06F862" bold="true" box="[151,374,1980,2007]" pageId="13" pageNumber="522">Diagnosis. Female.</emphasis>
Black with yellow spots and wings hyaline to fuscous. Relatively small size, maximum
<quantity id="4CED9B43FFB8FFF4C937F8414EEDF862" box="[1358,1437,1980,2007]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="522" unit="mm" value="15.0">15 mm</quantity>
in total length. Clypeus slightly wider than long or as long as wide, covered by short bristles at its base becoming long toward the apex, with medium-sized and deep punctures, present only in the apical region, apex rounded, clypeus-eye touching for an extent equal to or greater than width of antennal socket; gena width at midlength equal to or slightly wider than eye; occipital carina strong, extending to middle of the eye; humeri slightly projecting; mesepisternum with short bristles, and small and shallow punctation; propodeal concavity and striae absent; tergum I longer than wide, with short and dense bristles, punctures strong, medium sized, and, deep; tergum II as wide as long or wider than long, with short bristles, weakly punctate.
<emphasis id="B961EAB4FFBBFFF7CF29FEB648E4FED0" bold="true" box="[848,916,331,357]" pageId="14" pageNumber="523">Male.</emphasis>
Similar to females, except for the propodeum with concavity present, widely extended in the center and with fine striae.
</paragraph>
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<emphasis id="B961EAB4FFBBFFF7CCBEFE6F4ADAFE19" bold="true" box="[199,426,402,428]" pageId="14" pageNumber="523">Material examined</emphasis>
.
<specimenCount id="9D13FD2FFFBBFFF7CDC1FE6E4A9BFE18" box="[440,491,403,429]" count="21" pageId="14" pageNumber="523" type="female">21♀</specimenCount>
,
<specimenCount id="9D13FD2FFFBBFFF7CD8EFE6E496BFE18" box="[503,539,403,429]" count="3" pageId="14" pageNumber="523" type="male">3♂</specimenCount>
<materialsCitation id="3B7D3CFBFFBBFFF7CE5BFE6E4808FE18" box="[546,888,403,429]" country="Brazil" location="Rio de Janeiro" pageId="14" pageNumber="523" specimenCount="1" stateProvince="Rio de Janeiro">
in total.
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:
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,
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<specimenCount id="9D13FD2FFFBBFFF7CFFAFE6E48D6FE18" box="[899,934,403,429]" count="1" pageId="14" pageNumber="523" type="female">1♀</specimenCount>
,
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(
<collectionCode id="ED04AE63FFBBFFF7CF9FFE6E4F31FE18" box="[998,1089,403,429]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="14" pageNumber="523">AMNH</collectionCode>
)
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,
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<specimenCount id="9D13FD2FFFBBFFF7C82CFE6F4F08FE18" box="[1109,1144,402,429]" count="5" pageId="14" pageNumber="523" type="female">5♀</specimenCount>
(
<collectionCode id="ED04AE63FFBBFFF7C8FEFE6E4F93FE18" box="[1159,1251,403,429]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="523">MNHN</collectionCode>
);
<collectingRegion id="49D1F844FFBBFFF7C88EFE6E4BCFFE64" country="Brazil" name="Rio Grande do Sul" pageId="14" pageNumber="523">Rio Grande do Sul</collectingRegion>
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,
<materialsCitation id="3B7D3CFBFFBBFFF7CCB3FE4A4A3BFE64" box="[202,331,439,465]" collectionCode="NHM" country="Brazil" pageId="14" pageNumber="523" specimenCount="2" specimenCount-female="2" stateProvince="Rio Grande do Sul">
<specimenCount id="9D13FD2FFFBBFFF7CCB3FE4A4B9DFE64" box="[202,237,439,465]" count="2" pageId="14" pageNumber="523" type="female">2♀</specimenCount>
(
<collectionCode id="ED04AE63FFBBFFF7CC85FE4A4A33FE64" box="[252,323,439,465]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="14" pageNumber="523" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7D3CFBFFBBFFF7CD2FFE4A49FDFE64" box="[342,653,439,465]" collectionCode="CESC" country="Brazil" location="Santa Catarina" pageId="14" pageNumber="523" specimenCount="1" specimenCount-female="1" stateProvince="Santa Catarina">
<specimenCount id="9D13FD2FFFBBFFF7CD2FFE4A4A08FE64" box="[342,376,439,465]" count="1" pageId="14" pageNumber="523" type="female">1♀</specimenCount>
(
<collectionCode id="ED04AE63FFBBFFF7CDFEFE4A4AA3FE64" box="[391,467,439,465]" pageId="14" pageNumber="523">CESC</collectionCode>
);
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,
<materialsCitation id="3B7D3CFBFFBBFFF7CEE1FE4A48FCFE64" box="[664,908,439,465]" collectionCode="NHM" country="Brazil" pageId="14" pageNumber="523" specimenCount="1" specimenCount-female="1" stateProvince="Santa Catarina" typeStatus="holotype">
<specimenCount id="9D13FD2FFFBBFFF7CEE1FE4A49DDFE64" box="[664,685,439,465]" count="1" pageId="14" pageNumber="523" type="female"></specimenCount>
[
<typeStatus id="54AE8804FFBBFFF7CEC3FE4A4858FE64" box="[698,808,439,465]" pageId="14" pageNumber="523" type="holotype">Holotype</typeStatus>
] (
<collectionCode id="ED04AE63FFBBFFF7CF44FE4A48F4FE64" box="[829,900,439,465]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="14" pageNumber="523" type="Herbarium">NHM</collectionCode>
)
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,
<materialsCitation id="3B7D3CFBFFBBFFF7CFEEFE4A4FACFE64" box="[919,1244,439,465]" collectionCode="AMNH" country="Brazil" location="Sao Paulo" pageId="14" pageNumber="523" specimenCount="11" specimenCount-female="9" specimenCount-male="2" stateProvince="Sao Paulo" typeStatus="holotype">
<specimenCount id="9D13FD2FFFBBFFF7CFEEFE4A48CBFE64" box="[919,955,439,465]" count="9" pageId="14" pageNumber="523" type="female">9♀</specimenCount>
,
<specimenCount id="9D13FD2FFFBBFFF7CFB1FE4A489AFE64" box="[968,1002,439,465]" count="2" pageId="14" pageNumber="523" type="male">2♂</specimenCount>
(
<collectionCode id="ED04AE63FFBBFFF7CF80FE4A4F25FE64" box="[1017,1109,439,465]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="14" pageNumber="523">AMNH</collectionCode>
);
<collectingRegion id="49D1F844FFBBFFF7C811FE4A4FACFE64" box="[1128,1244,439,465]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="523">São Paulo</collectingRegion>
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,
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<specimenCount id="9D13FD2FFFBBFFF7C89FFE4A4E79FE64" box="[1254,1289,439,465]" count="1" pageId="14" pageNumber="523" type="female">1♀</specimenCount>
(
<collectionCode id="ED04AE63FFBBFFF7C961FE4A4E17FE64" box="[1304,1383,439,465]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="14" pageNumber="523" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7D3CFBFFBBFFF7C900FE4A4B9FFE40" collectionCode="NHM" country="Brazil" pageId="14" pageNumber="523" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="holotype">
<specimenCount id="9D13FD2FFFBBFFF7C900FE4A4EECFE64" box="[1401,1436,439,465]" count="1" pageId="14" pageNumber="523" type="female">1♀</specimenCount>
(
<collectionCode id="ED04AE63FFBBFFF7CCE6FE264B97FE40" box="[159,231,475,501]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="14" pageNumber="523" type="Herbarium">NHM</collectionCode>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="C30F652DFFBBFFF7CCBEFE0349C5FD88" pageId="14" pageNumber="523" type="distribution">
<paragraph id="8BAA36A6FFBBFFF7CCBEFE0349C5FD88" blockId="14.[151,1437,151,573]" pageId="14" pageNumber="523">
<emphasis id="B961EAB4FFBBFFF7CCBEFE034A11FDAD" bold="true" box="[199,353,510,536]" pageId="14" pageNumber="523">Distribution.</emphasis>
<collectingCountry id="F3027636FFBBFFF7CD1EFE024AC0FDAC" box="[359,432,511,537]" name="Brazil" pageId="14" pageNumber="523">Brazil</collectingCountry>
: Distrito Federal,
<collectingRegion id="49D1F844FFBBFFF7CE01FE02486AFDAC" box="[632,794,511,537]" country="Brazil" name="Espirito Santo" pageId="14" pageNumber="523">Espírito Santo</collectingRegion>
,
<collectingRegion id="49D1F844FFBBFFF7CF5DFE024818FDAC" box="[804,872,511,537]" country="Brazil" name="Goias" pageId="14" pageNumber="523">Goiás</collectingRegion>
,
<collectingRegion id="49D1F844FFBBFFF7CF0AFE024F7CFDAC" box="[883,1036,511,537]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="523">Minas Gerais</collectingRegion>
,
<collectingRegion id="49D1F844FFBBFFF7C86EFE024FCBFDAC" box="[1047,1211,511,537]" country="Brazil" name="Rio de Janeiro" pageId="14" pageNumber="523">Rio de Janeiro</collectingRegion>
,
<collectingRegion id="49D1F844FFBBFFF7C8BCFE024EE8FDAC" box="[1221,1432,511,537]" country="Brazil" name="Rio Grande do Sul" pageId="14" pageNumber="523">Rio Grande do Sul</collectingRegion>
,
<collectingRegion id="49D1F844FFBBFFF7CCEEFDDE4A4EFD88" box="[151,318,547,573]" country="Brazil" name="Santa Catarina" pageId="14" pageNumber="523">Santa Catarina</collectingRegion>
,
<collectingRegion id="49D1F844FFBBFFF7CD30FDDE4ACDFD88" box="[329,445,547,573]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="523">São Paulo</collectingRegion>
;
<collectingCountry id="F3027636FFBBFFF7CDB0FDDE4945FD88" box="[457,565,547,573]" name="Paraguay" pageId="14" pageNumber="523">Paraguay</collectingCountry>
;
<collectingCountry id="F3027636FFBBFFF7CE47FDDE49C1FD88" box="[574,689,547,573]" name="Argentina" pageId="14" pageNumber="523">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>