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<document id="C0E21717B94F3512C08D0B0B61175FAA" ID-CLB-Dataset="29024" ID-DOI="10.11646/zootaxa.4459.2.2" ID-GBIF-Dataset="634e23da-84c4-46a4-8bb8-94fcf1637160" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458548" ID-ZooBank="73510C3B-6482-4760-8173-B6E701839940" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539288809625" checkinUser="plazi" docAuthor="Sheth, Sayali D., Ghate, Hemant V. &amp; Hájek, Jiří" docDate="2018" docId="03BC3747FF96FF9EFF5E11D4FDCEF9AA" docLanguage="en" docName="zootaxa.4459.2.2.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copelatus neelumae Vazirani 1973" docType="treatment" docVersion="7" lastPageNumber="248" masterDocId="FF854F3FFF9BFF93FFC91143FFF8FFA8" masterDocTitle="Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="260" masterPageNumber="235" pageNumber="248" updateTime="1698578320975" updateUser="ExternalLinkService">
<mods:mods id="E2DBC0FC07F68E6070F255B92984D73C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="470DC63D75E1FA8D82E0F195528DF2FC">
<mods:title id="A8167C24009793A0AE69009D56B6835E">Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name id="9EFCFF90EEF1D4EDCFACA0C0AA641524" type="personal">
<mods:role id="66681416102F92780B5010E06F7080D5">
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</mods:role>
<mods:namePart id="D39D7876207639F279FE6850F2CFDF47">Sheth, Sayali D.</mods:namePart>
</mods:name>
<mods:name id="20157CBC741FF387EEF6AE2B9A0A1AE4" type="personal">
<mods:role id="5717E6E907C0080703CCB93873B84E96">
<mods:roleTerm id="E0ACCA39926B9D603E04143B1A261A6B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D4C1AE8E47E830E84DE62E5688AFC916">Ghate, Hemant V.</mods:namePart>
</mods:name>
<mods:name id="1907A45DF2A93830CF9B4EBB88295B46" type="personal">
<mods:role id="5B7803D69FD0274644DC457BA3846AFC">
<mods:roleTerm id="5315909ED7EAB98D616986FC510C2525">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7BE4C5CB3C740EA603E3CDCD6DD4501C">Hájek, Jiří</mods:namePart>
</mods:name>
<mods:typeOfResource id="76391FFDA74C29191BD054ED8233D5D9">text</mods:typeOfResource>
<mods:relatedItem id="4CCB55B29696A0B14CC53AAA875E90E2" type="host">
<mods:titleInfo id="28349041C4CCE6B85619A2BB0F8D2B54">
<mods:title id="61CDE21449F24E8F258DBC38C32CB793">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7B04690D8829294702D3E68A1F0A4A6E">
<mods:date id="39A5C62C7317A73E70E0CE8B408E46D3">2018</mods:date>
<mods:detail id="0CB5D5E476E3DEDC396C8354565C3B64" type="pubDate">
<mods:number id="E8CDDEB860C24BF696133081FCEF2A6D">2018-08-15</mods:number>
</mods:detail>
<mods:detail id="FE23BE093043578491F961EF6819D52C" type="volume">
<mods:number id="059FF52EA5DEFB8C8DE7732FBB742B5E">4459</mods:number>
</mods:detail>
<mods:detail id="65B7DB47B880638E483613345D23507D" type="issue">
<mods:number id="1AEBA255D9087DD4E90B92FF5F7AC028">2</mods:number>
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<mods:start id="25FA49418EB67815A53F4B7016E0D264">235</mods:start>
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<mods:classification id="E50F0BB55AE57372E63B1674F129529D">journal article</mods:classification>
<mods:identifier id="10912CB9626DC4D8E5AD08E06AD6AADA" type="CLB-Dataset">29024</mods:identifier>
<mods:identifier id="FB4FD8E9E7659E62E7400879C06FB36C" type="DOI">10.11646/zootaxa.4459.2.2</mods:identifier>
<mods:identifier id="508105F808D322C2F1220CE044A60890" type="GBIF-Dataset">634e23da-84c4-46a4-8bb8-94fcf1637160</mods:identifier>
<mods:identifier id="2F2BF7B36BB11224BA61FA11612FA4D6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="23E658CC682B10DF72AC194ACBF2A666" type="Zenodo-Dep">1458548</mods:identifier>
<mods:identifier id="4833AD101364A53CAA82B041C5E4A8F5" type="ZooBank">73510C3B-6482-4760-8173-B6E701839940</mods:identifier>
</mods:mods>
<treatment id="03BC3747FF96FF9EFF5E11D4FDCEF9AA" ID-DOI="http://doi.org/10.5281/zenodo.5966535" ID-GBIF-Taxon="148692673" ID-Zenodo-Dep="5966535" LSID="urn:lsid:plazi:treatment:03BC3747FF96FF9EFF5E11D4FDCEF9AA" httpUri="http://treatment.plazi.org/id/03BC3747FF96FF9EFF5E11D4FDCEF9AA" lastPageNumber="248" pageId="13" pageNumber="248">
<subSubSection id="C30FD5DAFF96FF9EFF5E11D4FE94FF7B" pageId="13" pageNumber="248" type="nomenclature">
<paragraph id="8BAA8651FF96FF9EFF5E11D4FDBFFF1A" blockId="13.[151,583,151,211]" box="[151,583,151,178]" pageId="13" pageNumber="248">
<heading id="D0E2313DFF96FF9EFF5E11D4FDBFFF1A" bold="true" box="[151,583,151,178]" fontSize="11" level="1" pageId="13" pageNumber="248" reason="1">
<taxonomicName id="4C15FDD2FF96FF9EFF5E11D4FDBFFF1A" ID-CoL="Y3CS" authority="Vazirani, 1973" authorityName="Vazirani" authorityYear="1973" box="[151,583,151,178]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="neelumae">
<emphasis id="B9615A43FF96FF9EFF5E11D4FDBFFF1A" bold="true" box="[151,583,151,178]" pageId="13" pageNumber="248">
<emphasis id="B9615A43FF96FF9EFF5E11D4FE74FF19" bold="true" box="[151,396,151,177]" italics="true" pageId="13" pageNumber="248">Copelatus neelumae</emphasis>
<bibRefCitation id="EF84FBA0FF96FF9EFE5A11DBFDBFFF1A" box="[403,583,152,178]" pageId="13" pageNumber="248" refId="ref16556" refString="Vazirani, T. G. (1973) Contribution to the study of aquatic beetles. 14. Copelatus neelumae sp. nov. (Dytiscidae) from India. Journal of the Bombay Natural History Society, 70 (1972), 224 - 226." type="journal article">Vazirani, 1973</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAA8651FF96FF9EFF5E11F9FE94FF7B" blockId="13.[151,583,151,211]" box="[151,364,186,211]" pageId="13" pageNumber="248">
(
<figureCitation id="132E9AD4FF96FF9EFF5611F9FEE9FF7B" box="[159,273,186,211]" captionStart="FIGURES 9–12" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[204,1385,231,1922]" captionTargetId="figure@5.[150,1436,193,1959]" captionTargetPageId="5" captionText="FIGURES 9–12. Habitus of Copelatus. 9–11. C. neelumae: 9. male, pale form (Karnataka); 10. male, dark form (Maharashtra, Lonavala); 11. female, immature specimen of dark form (Maharashtra, Mulshi); 12. C. schereri, male (Sadawaghapur)." httpUri="https://zenodo.org/record/1458554/files/figure.png" pageId="13" pageNumber="248">Figs 9–11</figureCitation>
,
<figureCitation id="132E9AD4FF96FF9EFED511F8FE9CFF7B" box="[284,356,187,211]" captionStart="FIGURES 23–30" captionStartId="14.[151,263,1513,1536]" captionTargetBox="[159,1413,213,1460]" captionTargetId="figure@14.[151,1436,193,1492]" captionTargetPageId="14" captionText="FIGURES 23–30. Male genitalia of Copelatus. 23–24. C. bezdeki sp. nov.; 25–26. C. indicus; 27–28. C. neelumae; 29–30. C. schereri. 23, 25, 27, 29. median lobe in lateral view (27a. redrawn from original description of C. neelumae); 24, 26, 28, 30. paramere. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1458560/files/figure.png" pageId="13" pageNumber="248">27–28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF96FF9EFF5E1041FDC1FEB0" box="[151,569,258,280]" pageId="13" pageNumber="248" type="reference_group">
<paragraph id="8BAA8651FF96FF9EFF5E1041FDC1FEB0" blockId="13.[151,569,258,280]" box="[151,569,258,280]" pageId="13" pageNumber="248">
<treatmentCitationGroup id="AB05A17FFF96FF9EFF5E1041FDC1FEB0" box="[151,569,258,280]" pageId="13" pageNumber="248">
<taxonomicName id="4C15FDD2FF96FF9EFF5E1041FDCDFEB0" ID-CoL="Y3CS" authority="Vazirani, 1973: 224" authorityName="Vazirani" authorityPageNumber="224" authorityYear="1973" box="[151,565,258,280]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="neelumae">
<emphasis id="B9615A43FF96FF9EFF5E1041FE9DFEB0" box="[151,357,258,280]" italics="true" pageId="13" pageNumber="248">Copelatus neelumae</emphasis>
<treatmentCitation id="0AB4A040FF96FF9EFEA21041FDCDFEB0" author="" box="[363,565,258,280]" page="224" pageId="13" pageNumber="248">
<bibRefCitation id="EF84FBA0FF96FF9EFEA21041FDFAFEB0" box="[363,514,258,280]" pageId="13" pageNumber="248" refId="ref16556" refString="Vazirani, T. G. (1973) Contribution to the study of aquatic beetles. 14. Copelatus neelumae sp. nov. (Dytiscidae) from India. Journal of the Bombay Natural History Society, 70 (1972), 224 - 226." type="journal article">Vazirani, 1973</bibRefCitation>
: 224
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF96FF9EFF5E1006FAB8FD9E" pageId="13" pageNumber="248" type="materials_examined">
<paragraph id="8BAA8651FF96FF9EFF5E1006FC65FEF6" blockId="13.[151,1437,325,1538]" box="[151,925,325,350]" pageId="13" pageNumber="248">
<materialsCitation id="3B7D8C0CFF96FF9EFF5E1006FC65FEF6" ID-GBIF-Occurrence="1917355307" box="[151,925,325,350]" country="India" location="Tiruchirapally District" municipality="Ottokovil" pageId="13" pageNumber="248" specimenCount="1" stateProvince="Tamilnadu" typeStatus="holotype">
<emphasis id="B9615A43FF96FF9EFF5E1006FECDFEF6" bold="true" box="[151,309,325,350]" pageId="13" pageNumber="248">
<typeStatus id="54AE38F3FF96FF9EFF5E1006FF2AFEF6" box="[151,210,325,350]" pageId="13" pageNumber="248">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F302C6C1FF96FF9EFEF51006FE82FEF6" box="[316,378,325,350]" name="India" pageId="13" pageNumber="248">India</collectingCountry>
: &quot;
<collectingRegion id="49D148B3FF96FF9EFE441006FDF4FEF6" box="[397,524,325,350]" country="India" name="Tamil Nadu" pageId="13" pageNumber="248">Tamilnadu</collectingRegion>
:
<collectingMunicipality id="6BCE1C2BFF96FF9EFDDF1006FD7CFEF6" box="[534,644,325,350]" pageId="13" pageNumber="248">Ottokovil</collectingMunicipality>
,
<location id="8ECAD08AFF96FF9EFD461006FC68FEF6" LSID="urn:lsid:plazi:treatment:03BC3747FF96FF9EFF5E11D4FDCEF9AA:8ECAD08AFF96FF9EFD461006FC68FEF6" box="[655,912,325,350]" country="India" municipality="Ottokovil" name="Tiruchirapally District" pageId="13" pageNumber="248" stateProvince="Tamilnadu">Tiruchirapally District</location>
&quot;.
</materialsCitation>
</paragraph>
<paragraph id="8BAA8651FF96FF9EFF0E102AFF0EFE0E" blockId="13.[151,1437,325,1538]" pageId="13" pageNumber="248">
<emphasis id="B9615A43FF96FF9EFF0E102AFE8BFE2A" bold="true" box="[199,371,361,386]" pageId="13" pageNumber="248">
<typeStatus id="54AE38F3FF96FF9EFF0E102AFEF9FE2A" box="[199,257,361,386]" pageId="13" pageNumber="248">Type</typeStatus>
material.
</emphasis>
<taxonomicName id="4C15FDD2FF96FF9EFEB2102AFDA4FE2A" authorityName="Vazirani" authorityYear="1973" box="[379,604,361,386]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="neelumae">
<emphasis id="B9615A43FF96FF9EFEB2102AFDA4FE2A" box="[379,604,361,386]" italics="true" pageId="13" pageNumber="248">Copelatus neelumae</emphasis>
</taxonomicName>
:
<typeStatus id="54AE38F3FF96FF9EFDA5102AFD2AFE2A" box="[620,722,361,386]" pageId="13" pageNumber="248" type="holotype">Holotype</typeStatus>
<specimenCount id="9D134DD8FF96FF9EFD131029FD08FE29" box="[730,752,362,385]" pageId="13" pageNumber="248" type="male">♂</specimenCount>
, deposited in Zoological Survey of
<collectingCountry id="F302C6C1FF96FF9EFB4B102AFB45FE2A" box="[1154,1213,361,386]" name="India" pageId="13" pageNumber="248">India</collectingCountry>
, Kolkata,
<collectingCountry id="F302C6C1FF96FF9EFAF9102AFA90FE2A" box="[1328,1384,361,386]" name="India" pageId="13" pageNumber="248">India</collectingCountry>
(not studied).
</paragraph>
<paragraph id="8BAA8651FF96FF9EFF0E10F2FAB8FD9E" blockId="13.[151,1437,325,1538]" pageId="13" pageNumber="248">
<materialsCitation id="3B7D8C0CFF96FF9EFF0E10F2FE56FE46" ID-GBIF-Occurrence="1917355257" collectingDate="2005-09-24" collectingDateMax="2005-09-28" collectingDateMin="2005-09-24" collectorName="J. Bezdek" country="India" elevation="500" location="Bhushi" pageId="13" pageNumber="248" specimenCount="1" specimenCount-male="1" stateProvince="Maharashtra">
<emphasis id="B9615A43FF96FF9EFF0E10F2FDCCFE62" bold="true" box="[199,564,433,458]" pageId="13" pageNumber="248">Additional material examined.</emphasis>
<collectingCountry id="F302C6C1FF96FF9EFD8910F2FD83FE62" box="[576,635,433,458]" name="India" pageId="13" pageNumber="248">India</collectingCountry>
:
<collectingRegion id="49D148B3FF96FF9EFD4510F2FCEFFE62" box="[652,791,433,458]" country="India" name="Maharashtra" pageId="13" pageNumber="248">Maharashtra</collectingRegion>
:
<specimenCount id="9D134DD8FF96FF9EFCEE10F2FCB2FE61" box="[807,842,433,457]" pageId="13" pageNumber="248" type="male">1♂</specimenCount>
,
<locationDeviation id="9A189F73FF96FF9EFC9410F2FBD1FE62" box="[861,1065,433,458]" pageId="13" pageNumber="248">
<quantity id="4CED2BB4FF96FF9EFC9410F2FC62FE62" box="[861,922,433,458]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="248" unit="km" value="4.0">4 km</quantity>
S Lonavala
</locationDeviation>
,
<location id="8ECAD08AFF96FF9EFBF310F2FB7FFE62" LSID="urn:lsid:plazi:treatment:03BC3747FF96FF9EFF5E11D4FDCEF9AA:8ECAD08AFF96FF9EFBF310F2FB7FFE62" box="[1082,1159,433,458]" country="India" name="Bhushi" pageId="13" pageNumber="248" stateProvince="Maharashtra">Bhushi</location>
dam env.,
<elevation id="00386162FF96FF9EFAC610F2FAA4FE61" box="[1295,1372,433,458]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="248" unit="m" value="500.0">
<quantity id="4CED2BB4FF96FF9EFAC610F2FAA4FE61" box="[1295,1372,433,458]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="248" unit="m" value="500.0">500 m</quantity>
</elevation>
,
<collectingDate id="EFEF5979FF96FF9EFAA410F2FEF5FE46" pageId="13" pageNumber="248" value="2005-09-24" valueMax="2005-09-28" valueMin="2005-09-24">
<date id="FFABA091FF96FF9EFAA410F2FEF5FE46" pageId="13" pageNumber="248" value="2005-09-24" valueMax="2005-09-28" valueMin="2005-09-24">24.– 28.ix.2005</date>
</collectingDate>
,
<collectorName id="26E0E387FF96FF9EFED11095FE7AFE46" box="[280,386,469,494]" pageId="13" pageNumber="248">J. Bezděk</collectorName>
lgt.
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF96FF9EFE701095FB8DFE46" ID-GBIF-Occurrence="1917355508" box="[441,1141,469,494]" collectingDate="2005-10-07" collectingDateMax="2005-10-11" collectingDateMin="2005-10-07" collectorName="J. Bezdek" country="India" location="Mulshi" municipality="Pune" pageId="13" pageNumber="248" specimenCount="1" specimenCount-male="1" stateProvince="Maharashtra">
<specimenCount id="9D134DD8FF96FF9EFE701095FE24FE46" box="[441,476,470,494]" pageId="13" pageNumber="248" type="male">1♂</specimenCount>
,
<quantity id="4CED2BB4FF96FF9EFE201095FDD6FE46" box="[489,558,469,494]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="248" unit="km" value="40.0">40 km</quantity>
W of
<collectingMunicipality id="6BCE1C2BFF96FF9EFDBA1095FD54FE45" box="[627,684,470,493]" pageId="13" pageNumber="248">Pune</collectingMunicipality>
,
<location id="8ECAD08AFF96FF9EFD7E1096FCFCFE46" LSID="urn:lsid:plazi:treatment:03BC3747FF96FF9EFF5E11D4FDCEF9AA:8ECAD08AFF96FF9EFD7E1096FCFCFE46" box="[695,772,469,494]" country="India" municipality="Pune" name="Mulshi" pageId="13" pageNumber="248" stateProvince="Maharashtra">Mulshi</location>
env.,
<collectingDate id="EFEF5979FF96FF9EFC8D1096FC2BFE46" box="[836,979,469,494]" pageId="13" pageNumber="248" value="2005-10-07" valueMax="2005-10-11" valueMin="2005-10-07">
<date id="FFABA091FF96FF9EFC8D1096FC2BFE46" box="[836,979,469,494]" pageId="13" pageNumber="248" value="2005-10-07" valueMax="2005-10-11" valueMin="2005-10-07">7.–11.x.2005</date>
</collectingDate>
,
<collectorName id="26E0E387FF96FF9EFC171095FBB0FE46" box="[990,1096,469,494]" pageId="13" pageNumber="248">J. Bezděk</collectorName>
lgt.
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF96FF9EFB491095FCEAFDB9" ID-GBIF-Occurrence="1917355494" collectingDate="2005-10-07" collectingDateMax="2005-10-08" collectingDateMin="2005-10-07" collectionCode="NMPC" collectorName="L. Borowiec" country="India" county="Pune Distr." location="Mulshi Lake" municipality="Mulshi" pageId="13" pageNumber="248" specimenCount="1" specimenCount-female="1" stateProvince="Maharashtra">
<specimenCount id="9D134DD8FF96FF9EFB491095FB5BFE46" box="[1152,1187,470,494]" pageId="13" pageNumber="248" type="female">1♀</specimenCount>
,
<collectingCounty id="62CBFEDDFF96FF9EFB791095FAD3FE46" box="[1200,1323,469,494]" pageId="13" pageNumber="248">Pune Distr.</collectingCounty>
,
<collectingMunicipality id="6BCE1C2BFF96FF9EFAFC1096FA7AFE46" box="[1333,1410,469,494]" pageId="13" pageNumber="248">Mulshi</collectingMunicipality>
at
<location id="8ECAD08AFF96FF9EFF5E10BAFEDFFDBA" LSID="urn:lsid:plazi:treatment:03BC3747FF96FF9EFF5E11D4FDCEF9AA:8ECAD08AFF96FF9EFF5E10BAFEDFFDBA" box="[151,295,505,530]" country="India" county="Pune Distr." municipality="Mulshi" name="Mulshi Lake" pageId="13" pageNumber="248" stateProvince="Maharashtra">Mulshi Lake</location>
,
<collectingDate id="EFEF5979FF96FF9EFEFC10BAFE40FDBA" box="[309,440,505,530]" pageId="13" pageNumber="248" value="2005-10-07" valueMax="2005-10-08" valueMin="2005-10-07">
<date id="FFABA091FF96FF9EFEFC10BAFE40FDBA" box="[309,440,505,530]" pageId="13" pageNumber="248" value="2005-10-07" valueMax="2005-10-08" valueMin="2005-10-07">7.–8.x.2005</date>
</collectingDate>
,
<collectorName id="26E0E387FF96FF9EFE0E10B9FDAAFDBA" box="[455,594,505,530]" pageId="13" pageNumber="248">L. Borowiec</collectorName>
lgt. (all
<collectionCode id="ED041E94FF96FF9EFD7110BAFCF2FDB9" box="[696,778,505,529]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="13" pageNumber="248">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7D8C0CFF96FF9EFCD610BAFE89FD9E" ID-GBIF-Occurrence="1917355434" collectingDate="2000-05-26" collectingDateMax="2000-05-28" collectingDateMin="2000-05-26" collectionCode="NMPC" collectorName="D. Hauck" country="India" location="Goa" pageId="13" pageNumber="248" specimenCount="1" specimenCount-male="1" stateProvince="Goa">
<collectingRegion id="49D148B3FF96FF9EFCD610BAFCA8FDB9" box="[799,848,505,529]" country="India" name="Goa" pageId="13" pageNumber="248">Goa</collectingRegion>
:
<specimenCount id="9D134DD8FF96FF9EFC9610BAFC7AFDB9" box="[863,898,505,529]" pageId="13" pageNumber="248" type="male">1♂</specimenCount>
, NW of Panaji, Baga Beach,
<collectingDate id="EFEF5979FF96FF9EFB1E10BAFA8BFDBA" box="[1239,1395,505,530]" pageId="13" pageNumber="248" value="2000-05-26" valueMax="2000-05-28" valueMin="2000-05-26">
<date id="FFABA091FF96FF9EFB1E10BAFA8BFDBA" box="[1239,1395,505,530]" pageId="13" pageNumber="248" value="2000-05-26" valueMax="2000-05-28" valueMin="2000-05-26">26.–28.v.2000</date>
</collectingDate>
,
<collectorName id="26E0E387FF96FF9EFA4B10B9FF26FD9E" pageId="13" pageNumber="248">D. Hauck</collectorName>
lgt. (
<collectionCode id="ED041E94FF96FF9EFEDC135DFE91FD9E" box="[277,361,542,566]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="13" pageNumber="248">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7D8C0CFF96FF9EFEB2135EFAC4FD9E" ID-GBIF-Occurrence="1917355345" box="[379,1340,541,566]" collectingDate="2006-05-26" collectingDateMax="2006-05-29" collectingDateMin="2006-05-26" collectionCode="NMPC" collectorName="Z. Kejval" country="India" location="Udipi Distr." pageId="13" pageNumber="248" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<collectingRegion id="49D148B3FF96FF9EFEB2135EFE16FD9E" box="[379,494,541,566]" country="India" name="Karnataka" pageId="13" pageNumber="248">Karnataka</collectingRegion>
:
<specimenCount id="9D134DD8FF96FF9EFE31135DFDE3FD9E" box="[504,539,542,566]" pageId="13" pageNumber="248" type="male">1♂</specimenCount>
,
<location id="8ECAD08AFF96FF9EFDEE135EFD52FD9E" LSID="urn:lsid:plazi:treatment:03BC3747FF96FF9EFF5E11D4FDCEF9AA:8ECAD08AFF96FF9EFDEE135EFD52FD9E" box="[551,682,541,566]" country="India" name="Udipi Distr." pageId="13" pageNumber="248" stateProvince="Karnataka">Udipi Distr.</location>
, E of Bhatkal, Kollur,
<collectingDate id="EFEF5979FF96FF9EFC55135EFBCFFD9E" box="[924,1079,541,566]" pageId="13" pageNumber="248" value="2006-05-26" valueMax="2006-05-29" valueMin="2006-05-26">
<date id="FFABA091FF96FF9EFC55135EFBCFFD9E" box="[924,1079,541,566]" pageId="13" pageNumber="248" value="2006-05-26" valueMax="2006-05-29" valueMin="2006-05-26">26.–29.v.2006</date>
</collectingDate>
,
<collectorName id="26E0E387FF96FF9EFB8B135DFB51FD9E" box="[1090,1193,541,566]" pageId="13" pageNumber="248">Z. Kejval</collectorName>
lgt. (
<collectionCode id="ED041E94FF96FF9EFB29135DFACCFD9E" box="[1248,1332,542,566]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="13" pageNumber="248">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF96FF9EFF0E1302FB50FC6A" pageId="13" pageNumber="248" type="diagnosis">
<paragraph id="8BAA8651FF96FF9EFF0E1302FB50FC6A" blockId="13.[151,1437,325,1538]" pageId="13" pageNumber="248">
<emphasis id="B9615A43FF96FF9EFF0E1302FEB9FDF2" bold="true" box="[199,321,577,602]" pageId="13" pageNumber="248">Diagnosis.</emphasis>
Medium sized (TL:
<quantity id="4CED2BB4FF96FF9EFDEE1302FD49FDF1" box="[551,689,577,602]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="13" pageNumber="248" unit="mm" value="5.35" valueMax="5.6" valueMin="5.1">5.1–5.6 mm</quantity>
), oblong oval species. Species highly variable in elytral coloration: elytra of
<typeStatus id="54AE38F3FF96FF9EFEC91325FEC8FDD6" box="[256,304,614,638]" pageId="13" pageNumber="248">type</typeStatus>
specimens largely yellow with only blackish, oval sutural spot extending to elytral stria 3, but not reaching base or apex of elytra (
<bibRefCitation id="EF84FBA0FF96FF9EFDCA13CAFD5FFD0A" box="[515,679,649,674]" pageId="13" pageNumber="248" refId="ref16556" refString="Vazirani, T. G. (1973) Contribution to the study of aquatic beetles. 14. Copelatus neelumae sp. nov. (Dytiscidae) from India. Journal of the Bombay Natural History Society, 70 (1972), 224 - 226." type="journal article">Vazirani 1973</bibRefCitation>
); specimens from
<collectingRegion id="49D148B3FF96FF9EFCBE13CAFC12FD0A" box="[887,1002,649,674]" country="India" name="Karnataka" pageId="13" pageNumber="248">Karnataka</collectingRegion>
similarly coloured, but with dark spot extended to elytral base and apex as narrow dark band along suture (
<figureCitation id="132E9AD4FF96FF9EFC5E13EEFC23FD6E" box="[919,987,685,710]" captionStart="FIGURES 9–12" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[204,1385,231,1922]" captionTargetId="figure@5.[150,1436,193,1959]" captionTargetPageId="5" captionText="FIGURES 9–12. Habitus of Copelatus. 9–11. C. neelumae: 9. male, pale form (Karnataka); 10. male, dark form (Maharashtra, Lonavala); 11. female, immature specimen of dark form (Maharashtra, Mulshi); 12. C. schereri, male (Sadawaghapur)." httpUri="https://zenodo.org/record/1458554/files/figure.png" pageId="13" pageNumber="248">Fig. 9</figureCitation>
); specimens from
<collectingRegion id="49D148B3FF96FF9EFB6213EDFB21FD6E" box="[1195,1241,686,710]" country="India" name="Goa" pageId="13" pageNumber="248">Goa</collectingRegion>
and
<collectingRegion id="49D148B3FF96FF9EFAD913EEFA64FD6E" box="[1296,1436,685,710]" country="India" name="Maharashtra" pageId="13" pageNumber="248">Maharashtra</collectingRegion>
darker, with yellow coloration confined to elytral base, interval between striae 3–4 and laterally from stria 5, and elytral apex (
<figureCitation id="132E9AD4FF96FF9EFEE713B6FE4AFCA6" box="[302,434,757,782]" captionStart="FIGURES 9–12" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[204,1385,231,1922]" captionTargetId="figure@5.[150,1436,193,1959]" captionTargetPageId="5" captionText="FIGURES 9–12. Habitus of Copelatus. 9–11. C. neelumae: 9. male, pale form (Karnataka); 10. male, dark form (Maharashtra, Lonavala); 11. female, immature specimen of dark form (Maharashtra, Mulshi); 12. C. schereri, male (Sadawaghapur)." httpUri="https://zenodo.org/record/1458554/files/figure.png" pageId="13" pageNumber="248">Figs 10–11</figureCitation>
). Elytral striae well impressed: stria 1 starting subbasally, striae 2, 3 and 5 shorter than striae 1 and 4—not reaching apex of elytra; stria 6 shortest, ending approximately in apical fifth of elytral length; submarginal stria long, starting at about elytral midlength and terminates in approximately apical fifth of elytral length. Female with few short strioles in middle third of elytra between striae 4–6 (
<figureCitation id="132E9AD4FF96FF9EFB991222FB5CFCD2" box="[1104,1188,865,890]" captionStart="FIGURES 9–12" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[204,1385,231,1922]" captionTargetId="figure@5.[150,1436,193,1959]" captionTargetPageId="5" captionText="FIGURES 9–12. Habitus of Copelatus. 9–11. C. neelumae: 9. male, pale form (Karnataka); 10. male, dark form (Maharashtra, Lonavala); 11. female, immature specimen of dark form (Maharashtra, Mulshi); 12. C. schereri, male (Sadawaghapur)." httpUri="https://zenodo.org/record/1458554/files/figure.png" pageId="13" pageNumber="248">Fig. 11</figureCitation>
). Median lobe sickleshaped in lateral view, almost uniformly broad throughout, with broad, obtusely pointed apex (
<figureCitation id="132E9AD4FF96FF9EFB7612C6FAECFC36" box="[1215,1300,901,926]" captionStart="FIGURES 23–30" captionStartId="14.[151,263,1513,1536]" captionTargetBox="[159,1413,213,1460]" captionTargetId="figure@14.[151,1436,193,1492]" captionTargetPageId="14" captionText="FIGURES 23–30. Male genitalia of Copelatus. 23–24. C. bezdeki sp. nov.; 25–26. C. indicus; 27–28. C. neelumae; 29–30. C. schereri. 23, 25, 27, 29. median lobe in lateral view (27a. redrawn from original description of C. neelumae); 24, 26, 28, 30. paramere. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1458560/files/figure.png" pageId="13" pageNumber="248">Fig. 27</figureCitation>
). Parameres moderately broad, 'C'-shaped; apex short and broad; apical lobes long, club-shaped (
<figureCitation id="132E9AD4FF96FF9EFB8112EAFB64FC6A" box="[1096,1180,937,962]" captionStart="FIGURES 23–30" captionStartId="14.[151,263,1513,1536]" captionTargetBox="[159,1413,213,1460]" captionTargetId="figure@14.[151,1436,193,1492]" captionTargetPageId="14" captionText="FIGURES 23–30. Male genitalia of Copelatus. 23–24. C. bezdeki sp. nov.; 25–26. C. indicus; 27–28. C. neelumae; 29–30. C. schereri. 23, 25, 27, 29. median lobe in lateral view (27a. redrawn from original description of C. neelumae); 24, 26, 28, 30. paramere. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1458560/files/figure.png" pageId="13" pageNumber="248">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF96FF9EFF0E128EFD8CFA12" pageId="13" pageNumber="248" type="discussion">
<paragraph id="8BAA8651FF96FF9EFF0E128EFD55FB49" blockId="13.[151,1437,325,1538]" pageId="13" pageNumber="248">
<emphasis id="B9615A43FF96FF9EFF0E128EFDECFC4E" bold="true" box="[199,532,973,998]" pageId="13" pageNumber="248">Comments on classification.</emphasis>
The species was described from the
<typeStatus id="54AE38F3FF96FF9EFC66128EFBE9FC4E" box="[943,1041,973,998]" pageId="13" pageNumber="248" type="holotype">holotype</typeStatus>
only, and remained enigmatic for a long time. Although
<bibRefCitation id="EF84FBA0FF96FF9EFE4F12B2FD42FBA2" box="[390,698,1009,1034]" pageId="13" pageNumber="248" refId="ref15798" refString="Holmen, M. &amp; Vazirani, T. G. (1990) Notes on the genera Neptosternus Sharp and Copelatus Erichson from Sri Lanka and India with the description of new species (Coleoptera: Dytiscidae). Koleopterologische Rundschau, 60, 19 - 32." type="journal article">Holmen &amp; Vazirani (1990)</bibRefCitation>
keyed the species and published the first record of
<taxonomicName id="4C15FDD2FF96FF9EFAC712B1FA64FBA2" authorityName="Vazirani" authorityYear="1973" box="[1294,1436,1009,1034]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="neelumae">
<emphasis id="B9615A43FF96FF9EFAC712B1FA64FBA2" box="[1294,1436,1009,1034]" italics="true" pageId="13" pageNumber="248">C. neelumae</emphasis>
</taxonomicName>
from
<collectingCountry id="F302C6C1FF96FF9EFF1F1556FEB0FB86" box="[214,328,1045,1070]" name="Sri Lanka" pageId="13" pageNumber="248">
<collectingRegion id="49D148B3FF96FF9EFF1F1556FF0FFB86" box="[214,247,1045,1070]" country="United Kingdom" name="Surrey" pageId="13" pageNumber="248">Sri</collectingRegion>
Lanka
</collectingCountry>
, and
<bibRefCitation id="EF84FBA0FF96FF9EFE4C1556FDD0FB86" box="[389,552,1045,1070]" pageId="13" pageNumber="248" refId="ref16293" refString="Rocchi, S. (2001) Dytiscidae raccolti in India durante due missioni entomologiche (1995, 1997) organizzate dal Museo Zoologico &quot; La Specola &quot; dell'Universita di Firenze (Coleoptera, Dytiscidae). Atti del Museo di Storia Naturale della Maremma, 19, 63 - 75." type="journal article">Rocchi (2001)</bibRefCitation>
mentioned the species from
<collectingCountry id="F302C6C1FF96FF9EFCBB1556FC40FB86" box="[882,952,1045,1070]" name="Nepal" pageId="13" pageNumber="248">Nepal</collectingCountry>
, none of these authors provided additional diagnostic characters and thus all researchers were referred to rather sketchy illustrations of elytral coloration and median lobe (
<figureCitation id="132E9AD4FF96FF9EFEFA151EFE6EFBDE" box="[307,406,1117,1142]" captionStart="FIGURES 23–30" captionStartId="14.[151,263,1513,1536]" captionTargetBox="[159,1413,213,1460]" captionTargetId="figure@14.[151,1436,193,1492]" captionTargetPageId="14" captionText="FIGURES 23–30. Male genitalia of Copelatus. 23–24. C. bezdeki sp. nov.; 25–26. C. indicus; 27–28. C. neelumae; 29–30. C. schereri. 23, 25, 27, 29. median lobe in lateral view (27a. redrawn from original description of C. neelumae); 24, 26, 28, 30. paramere. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1458560/files/figure.png" pageId="13" pageNumber="248">Fig. 27a</figureCitation>
). We have based our interpretation of the species on a specimen from
<collectingRegion id="49D148B3FF96FF9EFB74151EFAC8FBDE" box="[1213,1328,1117,1142]" country="India" name="Karnataka" pageId="13" pageNumber="248">Karnataka</collectingRegion>
in which dorsal coloration is similar to the figured
<typeStatus id="54AE38F3FF96FF9EFDA415C2FD37FB32" box="[621,719,1153,1178]" pageId="13" pageNumber="248" type="holotype">holotype</typeStatus>
and also the shape of the median lobe more or less agrees with the original illustration. Subsequently, we also assigned darker specimens from
<collectingRegion id="49D148B3FF96FF9EFBDB15E5FBB9FB16" box="[1042,1089,1190,1214]" country="India" name="Goa" pageId="13" pageNumber="248">Goa</collectingRegion>
and
<collectingRegion id="49D148B3FF96FF9EFBBE15E6FAFEFB16" box="[1143,1286,1189,1214]" country="India" name="Maharashtra" pageId="13" pageNumber="248">Maharashtra</collectingRegion>
, based on the same shape of the median lobe, to
<taxonomicName id="4C15FDD2FF96FF9EFDD31589FD5EFB4A" authorityName="Vazirani" authorityYear="1973" box="[538,678,1225,1250]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="neelumae">
<emphasis id="B9615A43FF96FF9EFDD31589FD5EFB4A" box="[538,678,1225,1250]" italics="true" pageId="13" pageNumber="248">C. neelumae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAA8651FF96FF9EFF0E15ADFB6CFA3E" blockId="13.[151,1437,325,1538]" pageId="13" pageNumber="248">
On the other hand, we had the possibility to study specimens from
<collectingCountry id="F302C6C1FF96FF9EFC7015AEFBDFFAAE" box="[953,1063,1261,1286]" name="Sri Lanka" pageId="13" pageNumber="248">
<collectingRegion id="49D148B3FF96FF9EFC7015AEFC22FAAE" box="[953,986,1261,1286]" country="United Kingdom" name="Surrey" pageId="13" pageNumber="248">Sri</collectingRegion>
Lanka
</collectingCountry>
(deposited in ZMUC), identified as
<taxonomicName id="4C15FDD2FF96FF9EFF711451FEBEFA82" authorityName="Vazirani" authorityYear="1973" box="[184,326,1297,1322]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="neelumae">
<emphasis id="B9615A43FF96FF9EFF711451FEBEFA82" box="[184,326,1297,1322]" italics="true" pageId="13" pageNumber="248">C. neelumae</emphasis>
</taxonomicName>
by T.G. Vazirani, and published by
<bibRefCitation id="EF84FBA0FF96FF9EFD211452FBB5FA82" box="[744,1101,1297,1322]" pageId="13" pageNumber="248" refId="ref15798" refString="Holmen, M. &amp; Vazirani, T. G. (1990) Notes on the genera Neptosternus Sharp and Copelatus Erichson from Sri Lanka and India with the description of new species (Coleoptera: Dytiscidae). Koleopterologische Rundschau, 60, 19 - 32." type="journal article">Holmen &amp; Vazirani (1990: 27)</bibRefCitation>
. However, the male genitalia (although the tip of the median lobe is broken) of specimens from
<collectingCountry id="F302C6C1FF96FF9EFC511476FBF0FAE6" box="[920,1032,1333,1358]" name="Sri Lanka" pageId="13" pageNumber="248">
<collectingRegion id="49D148B3FF96FF9EFC511476FC41FAE6" box="[920,953,1333,1358]" country="United Kingdom" name="Surrey" pageId="13" pageNumber="248">Sri</collectingRegion>
Lanka
</collectingCountry>
do not agree with the figure in the original description. We believe that this material belongs to another species, known to us across the Indian subcontinent, and similar to the widely distributed Afrotropical
<taxonomicName id="4C15FDD2FF96FF9EFCA8143DFB68FA3E" authority="(Klug, 1834)" authorityName="Klug" authorityYear="1834" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[865,1168,1405,1430]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="B9615A43FF96FF9EFCA8143DFC0DFA3E" box="[865,1013,1405,1430]" italics="true" pageId="13" pageNumber="248">C. pulchellus</emphasis>
(Klug, 1834)
</taxonomicName>
.
</paragraph>
<paragraph id="8BAA8651FF96FF9EFF0E14E2FD8CFA12" blockId="13.[151,1437,325,1538]" box="[199,628,1441,1466]" pageId="13" pageNumber="248">
<emphasis id="B9615A43FF96FF9EFF0E14E2FE0FFA12" bold="true" box="[199,503,1441,1466]" pageId="13" pageNumber="248">Collecting circumstances.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF96FF9EFF0E1486FDCEF9AA" pageId="13" pageNumber="248" type="distribution">
<paragraph id="8BAA8651FF96FF9EFF0E1486FDCEF9AA" blockId="13.[151,1437,325,1538]" pageId="13" pageNumber="248">
<emphasis id="B9615A43FF96FF9EFF0E1486FE98FA76" bold="true" box="[199,352,1477,1502]" pageId="13" pageNumber="248">Distribution.</emphasis>
<collectingRegion id="49D148B3FF96FF9EFEA21486FE28FA76" box="[363,464,1477,1502]" country="Sri Lanka" name="Southern" pageId="13" pageNumber="248">Southern</collectingRegion>
<collectingCountry id="F302C6C1FF96FF9EFE151486FDEEFA76" box="[476,534,1477,1502]" name="India" pageId="13" pageNumber="248">India</collectingCountry>
(
<collectingRegion id="49D148B3FF96FF9EFDE01485FDA4FA76" box="[553,604,1478,1502]" country="India" name="Goa" pageId="13" pageNumber="248">Goa</collectingRegion>
,
<collectingRegion id="49D148B3FF96FF9EFDA21486FD19FA76" box="[619,737,1477,1502]" country="India" name="Karnataka" pageId="13" pageNumber="248">Karnataka</collectingRegion>
,
<collectingRegion id="49D148B3FF96FF9EFD391486FC87FA76" box="[752,895,1477,1502]" country="India" name="Maharashtra" pageId="13" pageNumber="248">Maharashtra</collectingRegion>
,
<collectingRegion id="49D148B3FF96FF9EFC461486FBE2FA76" box="[911,1050,1477,1502]" country="India" name="Tamil Nadu" pageId="13" pageNumber="248">Tamil Nadu</collectingRegion>
) (
<figureCitation id="132E9AD4FF96FF9EFBFC1486FB75FA76" box="[1077,1165,1477,1502]" captionStart="FIGURE 45" captionStartId="21.[151,249,1915,1937]" captionTargetBox="[190,1401,196,1869]" captionTargetId="figure@21.[153,1434,158,1905]" captionTargetPageId="21" captionText="FIGURE 45. Distributional map of Copelatus species in Maharashtra." httpUri="https://zenodo.org/record/1458568/files/figure.png" pageId="13" pageNumber="248">Fig. 45</figureCitation>
). The records from
<collectingCountry id="F302C6C1FF96FF9EFAB21486FF25F9AA" name="Sri Lanka" pageId="13" pageNumber="248">
<collectingRegion id="49D148B3FF96FF9EFAB21486FA64FA76" box="[1403,1436,1477,1502]" country="United Kingdom" name="Surrey" pageId="13" pageNumber="248">Sri</collectingRegion>
Lanka
</collectingCountry>
and
<collectingCountry id="F302C6C1FF96FF9EFEDA14AAFEAEF9AA" box="[275,342,1513,1538]" name="Nepal" pageId="13" pageNumber="248">Nepal</collectingCountry>
need to be verified.
</paragraph>
</subSubSection>
</treatment>
</document>
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