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<document id="8F7904A4CA060512A2BE96BCB9C65D78" ID-DOI="10.5281/zenodo.158472" ID-GBIF-Dataset="2e1982e8-c59f-4ca0-92b4-c46e9f542752" ID-ISSN="1175­5326" ID-Zenodo-Dep="158472" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461024732819" checkinUser="plazi" docAuthor="Burks, R. A. &amp; Redak, R. A." docDate="2004" docId="03B887BBBA05FFBBFE922FF5FBDDFDD7" docLanguage="en" docName="zt00606.pdf" docOrigin="Zootaxa 606" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Meximalus skinnerensis Burks, n. sp." docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="FF81FFC3BA03FFB6FF9A2D39FFACFFE8" masterDocTitle="New species of Pteromalidae and Torymidae (Hymenoptera: Chalcidoidea) from California, with taxonomic notes" masterLastPageNumber="20" masterPageNumber="1" pageNumber="7" updateTime="1698571605235" updateUser="plazi">
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<mods:title id="99106154771C30A3057357D3848B30B2">New species of Pteromalidae and Torymidae (Hymenoptera: Chalcidoidea) from California, with taxonomic notes</mods:title>
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<mods:namePart id="F0C422E39A0E62D0DC6D601A3DDFA31F">Burks, R. A.</mods:namePart>
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<mods:namePart id="A44237FA1086FDC574611C11F67EE0F9">Redak, R. A.</mods:namePart>
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<mods:date id="ECF73A612C831861A4516C058AA25751">2004</mods:date>
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<subSubSection id="C30B6526BA05FFB0FE922FF5FD7FFD0E" box="[264,723,716,742]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BAE36ADBA05FFB0FE922FF5FD7FFD0E" blockId="6.[264,723,716,742]" box="[264,723,716,742]" pageId="6" pageNumber="7">
<heading id="D0E681C1BA05FFB0FE922FF5FD7FFD0E" bold="true" box="[264,723,716,742]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B965EABFBA05FFB0FE922FF5FD7FFD0E" bold="true" box="[264,723,716,742]" pageId="6" pageNumber="7">
<taxonomicName id="4C114D2EBA05FFB0FE922FF5FD2FFD0E" ID-CoL="73F3V" authority="Burks" authorityName="Burks" box="[264,643,716,742]" class="Insecta" family="Pteromalidae" genus="Meximalus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="skinnerensis" status="sp. nov.">
<emphasis id="B965EABFBA05FFB0FE922FF5FD82FD0E" bold="true" box="[264,558,716,742]" italics="true" pageId="6" pageNumber="7">Meximalus skinnerensis</emphasis>
Burks
</taxonomicName>
,
<taxonomicNameLabel id="A25657C4BA05FFB0FD0B2FF5FD7FFD0E" box="[657,723,716,742]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C30B6526BA05FFB0FE922E37FE5AFC9B" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BAE36ADBA05FFB0FE922E37FE5AFC9B" blockId="6.[264,1325,782,1891]" pageId="6" pageNumber="7">
<emphasis id="B965EABFBA05FFB0FE922E37FE5AFCC0" bold="true" box="[264,502,782,808]" pageId="6" pageNumber="7">Diagnosis (female).</emphasis>
Temples/dorsal eye length 0.38 (Fig. 5) [
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<taxonomicName id="4C114D2EBA05FFB0FBA72E36FB14FCC0" box="[1085,1208,783,808]" class="Insecta" family="Pteromalidae" genus="Meximalus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="heratyi">
<emphasis id="B965EABFBA05FFB0FBA72E36FB14FCC0" box="[1085,1208,783,808]" italics="true" pageId="6" pageNumber="7">M. heratyi</emphasis>
</taxonomicName>
(Fig. 4)]. Metafemur 4.4x longer than broad [3.25x in
<taxonomicName id="4C114D2EBA05FFB0FC8C2E0DFC3DFCA5" box="[790,913,820,845]" class="Insecta" family="Pteromalidae" genus="Meximalus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="heratyi">
<emphasis id="B965EABFBA05FFB0FC8C2E0DFC3DFCA5" box="[790,913,820,845]" italics="true" pageId="6" pageNumber="7">M. heratyi</emphasis>
</taxonomicName>
]. Profemur 4.8x longer than broad [3.7x in
<taxonomicName id="4C114D2EBA05FFB0FEF12E60FE49FC9A" box="[363,485,857,882]" class="Insecta" family="Pteromalidae" genus="Meximalus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="heratyi">
<emphasis id="B965EABFBA05FFB0FEF12E60FE49FC9A" box="[363,485,857,882]" italics="true" pageId="6" pageNumber="7">M. heratyi</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="C30B6526BA05FFBDFEA22E47FBDDFA04" lastPageId="11" lastPageNumber="12" pageId="6" pageNumber="7" type="description">
<paragraph id="8BAE36ADBA05FFB0FEA22E47FBFDFC56" blockId="6.[264,1325,782,1891]" pageId="6" pageNumber="7">
<emphasis id="B965EABFBA05FFB0FEA22E47FD07FC70" bold="true" box="[312,683,894,920]" pageId="6" pageNumber="7">
Description.
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female.
</emphasis>
Body length
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(
<quantity id="4CE99B48BA05FFB0FC292E46FC43FC71" box="[947,1007,895,921]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="6" pageNumber="7" unit="in" value="14.0">14 in</quantity>
relative units used below). No other female specimens known, therefore variation is not addressed.
</paragraph>
<paragraph id="8BAE36ADBA05FFB0FEA22EF0FA86FB2B" blockId="6.[264,1325,782,1891]" pageId="6" pageNumber="7">Color: Head and mesosoma black with weak metallic luster, with white, slightly thickened, setae; eyes and ocelli pinkish white; mandibles centrally light brown, peripherally dark reddish brown; maxillary and labial palps light brown; entire antenna light brown, the flagellum gradually becoming darker apically; tegula and humeral plate light brown, tegula semitranslucent; venation centrally white and peripherally light brown, stigma entirely light brown; all coxae same color as mesopleuron, most of legs light brown, except pro­ and mesofemora and tarsal claws darker brown; gaster brown with varying metallic luster.</paragraph>
<paragraph id="8BAE36ADBA05FFB0FEA229F6FE11F9B6" blockId="6.[264,1325,782,1891]" pageId="6" pageNumber="7">
Sculpture: Most of head and mesosoma deeply and broadly alveolate, except: clypeus very shallowly alveolate with even shallower indications of radiating striae that are limited to clypeal area; postocciput coriaceous with elongate meshes, sharp transition, very weakly carinate, between occiput and postocciput; frenum not differently sculptured from remainder of scutellum; upper mesepimeron entirely finely alveolate; one large and broad depression along dorsal edge of lower mesepimeron (this divided into pits in most other
<taxonomicName id="4C114D2EBA05FFB0FE922896FE04FA21" box="[264,424,1455,1481]" class="Insecta" family="Pteromalidae" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subFamily" subFamily="Pteromalinae">Pteromalinae</taxonomicName>
); acropleuron mostly coriaceous but changing to finely alveolate near tegula; visible surface of lateral panels of preaxilla and axilla shallowly alveolate and relatively glossy; prepectus finely and shallowly alveolate; dorsellum smooth; propodeum entirely transverse­alveolate, with callus much more shallowly sculpted and all sunken areas more finely sculpted.
</paragraph>
<paragraph id="8BAE36ADBA05FFB1FEA22B50FC98FDFE" blockId="6.[264,1325,782,1891]" lastBlockId="7.[264,1325,284,1878]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">Head (Figs. 5, 8) width 4.8, height 3.5, length 2.2; Dorsal IOD 3.4, ventral IOD 3.6; bottom of toruli about 0.2 above lower eye margin; ITD 0.3; VTD 2.1; MTD 1.15; ocular setae shorter than diameter of an adjacent ommatidium; clypeus shallowly emarginate, lateral incisions of clypeus no deeper or more extensive than median incision; EH 1.9; MS 1.2; MW 1.8; POL 0.9; OOL 1.1; EL 1.3; TL 0.5; malar sulcus present and deep; genal concavity vague and weak, about 0.4x malar space but not defined by a carina or change in sculpture, with a distinct narrow notch for reception of mandible base; genal carina distinct. Scape 1.9:0.4; pedicel 0.6:0.3; the two anelli tiny and strongly transverse; 1st funicular segment as long as broad, slightly shorter than pedicel, following funicular segments increasing in width and decreasing in length until 6th funicular segment twice as broad as long; funicular and claval segments with 1 row of longitudinal sensilla that slightly exceed the segment apex; flagellar setae inconspicuous and appressed to the segments; club slightly shorter than preceding three segments, sensory area tiny and at claval apex. Both mandibles with 3 distinctly separated denticles.</paragraph>
<paragraph id="8BAE36ADBA04FFB1FEA22F18FED0F973" blockId="7.[264,1325,284,1878]" pageId="7" pageNumber="8">
Mesosoma (Figs. 8, 10). Pronotal collar medial length 0.7, width 3.4; mesoscutum length:width 2.0:3.9; scutellum 2.4:2.2 (width at axillular apices); dorsellum &lt;0.1x scutellar length; propodeum length 1.2. Pronotal collar smoothly rounded anteriorly, collum steep and short, visible dorsally for a length of 0.2; lateral panel of pronotum with a deep and narrow, almost vertical and straight, scrobe beginning abruptly near ventral edge of lateral pronotal panel and ending to dorsal surface of pronotum such that the collar has a distinct lateral notch when seen from dorsal view; axillar separation 0.4; axillulae small, about 0.25x scutellar height; scutellum from dorsal view heptagonal, bluntly pointed posteriorly and projecting straight posteriorly over propodeum to end at about the level of nuchal base; frenal separation only apparent at lateral extremes, but frenum comprising ventral surface of scutellar projection; dorsal margin of prepectus shorter than tegula. Propodeum at about a 70° angle with top of scutellum, with a trident­shaped complex of complete median carina and anterior plicae that smoothly unite with the costula near propodeal midlength, ISD 2.15; anterior and posterior plical pits not evident; spiracles emarginate; postspiracular furrow deep, moderately narrow, and uninterrupted, ending in a distinct pit at supracoxal flange; supracoxal flange slightly greater in length than short axis of spiracle, ending near nuchal base; nucha convex, without basal cross­carina and with a very vague basal furrow, 0.5 (from end of median carina)x0.8, extending distinctly beyond metacoxal bases but not reaching their midlength. PRF 2.9:0.6; PRT 2.6:0.4; MSF 2.5:0.5; MST 3.0:0.5; MTF 3.1:0.7; MTT 3.6:0.4; mesotibial spur 0.5; basal mesotarsomere 0.9; metatibial spur 0.4; basal metatarsomere 0.9. FW 9:4.4; CC 3.3; MV 2.1; PMV 1.5; SV 1.2, stigma large and teardrop­shaped, broadening apically, shortest distance from uncal apex to MV 0.6, from stigmal apex to MV 1.1; uncus tiny, less than 0.2x greatest stigmal width; venational setae as in
<taxonomicName id="4C114D2EBA04FFB1FDF92845FD73FA7D" box="[611,735,1404,1429]" class="Insecta" family="Pteromalidae" genus="Meximalus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="heratyi">
<emphasis id="B965EABFBA04FFB1FDF92845FD73FA7D" box="[611,735,1404,1429]" italics="true" pageId="7" pageNumber="8">M. heratyi</emphasis>
</taxonomicName>
, with very long submarginal vein setae and three even longer parastigmal setae; costal cell with one complete row of ventral setae and additional ventral setae in apical quarter, with a row of 6 dorsal setae in apical half; speculum extending from middle of parastigma to be closed posteriorly by the complete subcubital setal track, with an occasional isolated ventral seta; five vague rows of short admarginal setae present but not exposed by speculum; basal cell sparsely setose, basal fold setose; cubital setal track ending at speculum. HW 7.0:2.4, not different from the typical pteromaline form.
</paragraph>
<paragraph id="8BAE36ADBA04FFB1FEA22B9EFABDF8BE" blockId="7.[264,1325,284,1878]" pageId="7" pageNumber="8">Petiole transverse and small. Gaster 6.0:3.5, dorsally without rigid form and tending to collapse; first 5 gastral tergites with short medial posterior notch, sternites 13 each with a deep posterior incision; hypopygium ending 7.3 from gastral base (exceeding gastral apex, which is slightly upturned), with a small lateral­apical notch on each side and a smaller ventral­apical notch; ovipositor short: total length about 4.5, not exceeding gastral apex.</paragraph>
<paragraph id="8BAE36ADBA0BFFBEFEA22C22FB81FDFE" blockId="8.[264,1324,283,1019]" pageId="8" pageNumber="9">
<emphasis id="B965EABFBA0BFFBEFEA22C22FE26FEDD" bold="true" box="[312,394,283,309]" pageId="8" pageNumber="9">Males.</emphasis>
Body (Figs. 7, 9) length
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(
<quantity id="4CE99B48BA0BFFBEFCF62C25FC5FFEDE" box="[876,1011,284,310]" metricMagnitude="-1" metricUnit="m" metricValue="2.2479" metricValueMax="2.5907999999999998" metricValueMin="1.905" pageId="8" pageNumber="9" unit="in" value="8.85" valueMax="10.2" valueMin="7.5">7.510.2 in</quantity>
relative units). With some characters given in the above discussion of the genus. Differing from female in the characters noted in the generic discussion above. Also, scape 1.21.4:0.2; pedicel 0.4:0.3; EL 0.81.0; TL 0.30.4; genal concavity weaker and only about 0.15x MS; pronotal scrobe more sinuate, without an abrupt ventral ending; scutellum 1.6:1.8; much more bluntly pointed posteriorly; legs dark brown, with basal tarsomeres and extremes of tibiae lighter; PRF 1.62.2:0.30.5; MTF 1.92.6:0.30.5; petiole 0.40.5:0.30.4.
</paragraph>
<paragraph id="8BAE36ADBA0BFFBEFEA22F18FA87FDD3" blockId="8.[264,1324,283,1019]" box="[312,1323,545,571]" pageId="8" pageNumber="9">Variation. Only one female specimen known. Males showing no noteworthy variation.</paragraph>
<paragraph id="8BAE36ADBA0BFFBEFEA22F7EFE56FD89" blockId="8.[264,1324,283,1019]" box="[312,506,583,609]" pageId="8" pageNumber="9">Host: Unknown.</paragraph>
<paragraph id="8BAE36ADBA0BFFBEFEA22F55FECCFD43" blockId="8.[264,1324,283,1019]" pageId="8" pageNumber="9">Distribution: Riverside County, northeast of Lake Skinner, all from coastal sage scrub habitat.</paragraph>
<paragraph id="8BAE36ADBA0BFFBEFEA22F8EFBB3FD39" blockId="8.[264,1324,283,1019]" box="[312,1055,695,721]" pageId="8" pageNumber="9">
Etymology: After Lake Skinner, which is near the
<typeStatus id="54AA880FBA0BFFBEFC122F8EFC16FD39" box="[904,954,695,721]" pageId="8" pageNumber="9">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BAE36ADBA0BFFBEFEA22FE5FE5CFC13" blockId="8.[264,1324,283,1019]" pageId="8" pageNumber="9">
<typeStatus id="54AA880FBA0BFFBEFEA22FE5FE09FD1E" box="[312,421,732,758]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
Ψ
<emphasis id="B965EABFBA0BFFBEFE572FE5FD0BFD1E" bold="true" box="[461,679,732,758]" pageId="8" pageNumber="9">
<collectingCountry id="F306763DBA0BFFBEFE572FE5FDA0FD1E" box="[461,524,732,758]" name="United States of America" pageId="8" pageNumber="9">USA</collectingCountry>
: California:
</emphasis>
Riverside Co. Lake Skinner/Shipley Res.
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NE Lake Skinner. 33°38 N 117°0 W. elev.
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.
</specimenCount>
<date id="FFAF106DBA0BFFBEFCB42E38FC70FCF3" box="[814,988,769,795]" pageId="8" pageNumber="9" value="2000-09-17" valueMax="2000-09-20" valueMin="2000-09-17">1720.ix.2000</date>
. Redak Lab, UCR. voucher: hym562 b6p3#23 (USNM). Allotype ɗ Riverside Co. Lake Skinner (NE End) 33°
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117°
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2
<specimenCount id="9D17FD24BA0BFFBEFECE2E75FE23FC8E" box="[340,399,844,870]" pageId="8" pageNumber="9" type="generic">5 W</specimenCount>
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.
<date id="FFAF106DBA0BFFBEFE072E75FE5DFC8E" box="[413,497,844,870]" pageId="8" pageNumber="9" value="1921-05-07">V­7/21</date>
­1996, J.D. Pinto. MT Coastal Sage Scrub. METU11, UNBURNED (USNM). 1 ɗ
<typeStatus id="54AA880FBA0BFFBEFE2A2E48FDB9FC63" box="[432,533,881,907]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
Riverside Co. Lake Skinner (NE End) 33°
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117°
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<date id="FFAF106DBA0BFFBEFB6D2E48FE89FC59" pageId="8" pageNumber="9" value="1921-05-07">V­7/ 21</date>
­1996, J.D. Pinto. MT Coastal Sage Scrub (UCRC). 1 ɗ
<typeStatus id="54AA880FBA0BFFBEFC4E2EAEFB95FC59" box="[980,1081,919,945]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
Riverside Co. Lake Skinner (NE End) 33°
<geoCoordinate id="EE25506ABA0BFFBEFD8A2E85FDC5FC3E" box="[528,617,956,982]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="36.116665">36 7 N</geoCoordinate>
117°
<geoCoordinate id="EE25506ABA0BFFBEFD332E85FD54FC3E" box="[681,760,956,982]" direction="west" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-2.0833333">2 5 W</geoCoordinate>
ca 1580
<date id="FFAF106DBA0BFFBEFCE92E85FC3FFC3E" box="[883,915,956,982]" pageId="8" pageNumber="9">v.2</date>
/vi.4.1998 MT coastal sage scrub, J.D. Pinto (UCDC).
</paragraph>
<paragraph id="8BAE36ADBA0BFFBEFE922A37FB5BF8A1" blockId="8.[264,1323,1806,1865]" pageId="8" pageNumber="9">
<emphasis id="B965EABFBA0BFFBEFE922A37FED6F8CE" bold="true" box="[264,378,1806,1830]" pageId="8" pageNumber="9">PLATE 1.</emphasis>
Figures 13.
<taxonomicName id="4C114D2EBA0BFFBEFD8A2A36FD76F8CE" box="[528,730,1807,1830]" class="Insecta" family="Pteromalidae" genus="Chlorocytus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="analis" status="comb. nov.">
<emphasis id="B965EABFBA0BFFBEFD8A2A36FD76F8CE" box="[528,730,1807,1830]" italics="true" pageId="8" pageNumber="9">Chlorocytus analis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657C4BA0BFFBEFD782A36FCEDF8CE" box="[738,833,1807,1830]" pageId="8" pageNumber="9">n. comb.</taxonomicNameLabel>
<typeStatus id="54AA880FBA0BFFBEFCDD2A36FC05F8CE" box="[839,937,1807,1830]" pageId="8" pageNumber="9" type="lectotype">lectotype</typeStatus>
female. Figure 1. Head and mesosoma dorsal view. Figure 2. Head and mesosoma lateral view. Figure 3. Metasoma dorsal view.
</paragraph>
<paragraph id="8BAE36ADBA0AFFBFFE922BEAFBFCF882" blockId="9.[264,1323,1747,1898]" pageId="9" pageNumber="10">
<emphasis id="B965EABFBA0AFFBFFE922BEAFED5F903" bold="true" box="[264,377,1747,1771]" pageId="9" pageNumber="10">PLATE 2.</emphasis>
Figure 4.
<taxonomicName id="4C114D2EBA0AFFBFFE7D2BEAFD02F902" box="[487,686,1747,1770]" class="Insecta" family="Pteromalidae" genus="Meximalus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="heratyi">
<emphasis id="B965EABFBA0AFFBFFE7D2BEAFD02F902" box="[487,686,1747,1770]" italics="true" pageId="9" pageNumber="10">Meximalus heratyi</emphasis>
</taxonomicName>
<typeStatus id="54AA880FBA0AFFBFFD2E2BEAFCBDF902" box="[692,785,1747,1770]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
female, head from dorsal view. Figures 68.
<taxonomicName id="4C114D2EBA0AFFBFFB6E2BEAFE70F8E2" class="Insecta" family="Pteromalidae" genus="Meximalus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="skinnerensis" status="sp. nov.">
<emphasis id="B965EABFBA0AFFBFFB6E2BEAFE70F8E2" italics="true" pageId="9" pageNumber="10">Meximalus skinnerensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657C4BA0AFFBFFE7F2BCAFD8EF8E2" box="[485,546,1779,1802]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
allotype male. Figure 6. Antenna. Figure 7. Lateral habitus. Figure 8. Head and mesosoma dorsal view. Figures 910.
<taxonomicName id="4C114D2EBA0AFFBFFCB92A2AFB88F8C2" box="[803,1060,1811,1834]" class="Insecta" family="Pteromalidae" genus="Meximalus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="skinnerensis" status="sp. nov.">
<emphasis id="B965EABFBA0AFFBFFCB92A2AFB88F8C2" box="[803,1060,1811,1834]" italics="true" pageId="9" pageNumber="10">Meximalus skinnerensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657C4BA0AFFBFFBB42A2AFBC7F8C2" box="[1070,1131,1811,1834]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
<typeStatus id="54AA880FBA0AFFBFFBEE2A2AFB7DF8C2" box="[1140,1233,1811,1834]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
female. Figure 9. Face and antennae. Figure 10. Lateral habitus. Figures 1112.
<taxonomicName id="4C114D2EBA0AFFBFFB9A2A0AFB16F8A2" box="[1024,1210,1843,1866]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="basalis" status="sp. nov.">
<emphasis id="B965EABFBA0AFFBFFB9A2A0AFB16F8A2" box="[1024,1210,1843,1866]" italics="true" pageId="9" pageNumber="10">Euteloida basalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657C4BA0AFFBFFB5A2A0AFB54F8A2" box="[1216,1272,1843,1866]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
Figure 11. Allotype male antenna. Figure 12.
<typeStatus id="54AA880FBA0AFFBFFD512A6AFC84F882" box="[715,808,1875,1898]" pageId="9" pageNumber="10" type="paratype">Paratype</typeStatus>
female thorax, dorsal view.
</paragraph>
<paragraph id="8BAE36ADBA09FFBCFE9228A6FBECFA13" blockId="10.[264,1324,1439,1532]" pageId="10" pageNumber="11">
<emphasis id="B965EABFBA09FFBCFE9228A6FED5FA5F" bold="true" box="[264,377,1439,1463]" pageId="10" pageNumber="11">PLATE 3.</emphasis>
Figures 1315.
<taxonomicName id="4C114D2EBA09FFBCFDBD28A6FCB4FA5E" box="[551,792,1439,1462]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="skinnerensis" status="sp. nov.">
<emphasis id="B965EABFBA09FFBCFDBD28A6FCB4FA5E" box="[551,792,1439,1462]" italics="true" pageId="10" pageNumber="11">Euteloida skinnerensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657C4BA09FFBCFC8428A6FCF4FA5E" box="[798,856,1439,1462]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
female. Figure 13. Propodeum of
<typeStatus id="54AA880FBA09FFBCFB5D28A6FA84FA5E" box="[1223,1320,1439,1462]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
; bcn = basal carina of nucha. Figure 14. Antennae of
<typeStatus id="54AA880FBA09FFBCFCA328FBFC34FA31" box="[825,920,1474,1497]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
, ventral view. Figure 15. Forewing of
<typeStatus id="54AA880FBA09FFBCFE9228DDFEC5FA13" box="[264,361,1508,1531]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
. Figure 16.
<taxonomicName id="4C114D2EBA09FFBCFE7D28DCFD18FA14" box="[487,692,1509,1532]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="subnigra">
<emphasis id="B965EABFBA09FFBCFE7D28DCFD18FA14" box="[487,692,1509,1532]" italics="true" pageId="10" pageNumber="11">Euteloida subnigra</emphasis>
</taxonomicName>
<typeStatus id="54AA880FBA09FFBCFD2128DDFCB4FA13" box="[699,792,1508,1531]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
female forewing basal cell.
</paragraph>
<paragraph id="8BAE36ADBA08FFBDFE9228B6FBDDFA04" blockId="11.[264,1323,1423,1516]" pageId="11" pageNumber="12">
<emphasis id="B965EABFBA08FFBDFE9228B6FED5FA4F" bold="true" box="[264,377,1423,1447]" pageId="11" pageNumber="12">PLATE 4.</emphasis>
Figures 1721.
<taxonomicName id="4C114D2EBA08FFBDFDBD28A9FCB0FA4F" box="[551,796,1424,1447]" class="Insecta" family="Torymidae" genus="Pseuderimerus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="burgeri" status="sp. nov.">
<emphasis id="B965EABFBA08FFBDFDBD28A9FCB0FA4F" box="[551,796,1424,1447]" italics="true" pageId="11" pageNumber="12">Pseuderimerus burgeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657C4BA08FFBDFCB928A9FCF0FA4F" box="[803,860,1424,1447]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
Figure 17.
<typeStatus id="54AA880FBA08FFBDFC4C28A9FB95FA4F" box="[982,1081,1424,1447]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
female dorsal habitus. Figure 18.
<typeStatus id="54AA880FBA08FFBDFE19288AFE4AFA22" box="[387,486,1459,1482]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
female lateral habitus. Figure 19. Allotype male lateral habitus. Figure 20. Allotype male genitalia from ventral view. Figure 21. Melanistic
<typeStatus id="54AA880FBA08FFBDFC2528ECFBB7FA04" box="[959,1051,1493,1516]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
female.
</paragraph>
</subSubSection>
<subSubSection id="C30B6526BA08FFBAFEA22B5BFDB7FD3E" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BAE36ADBA08FFBDFEA22B5BFD53F8FA" blockId="11.[264,1323,1634,1810]" pageId="11" pageNumber="12">
<emphasis id="B965EABFBA08FFBDFEA22B5BFE6FF994" bold="true" box="[312,451,1634,1660]" pageId="11" pageNumber="12">Discussion.</emphasis>
The differences between this species and
<taxonomicName id="4C114D2EBA08FFBDFC372B5AFB8BF994" box="[941,1063,1635,1660]" class="Insecta" family="Pteromalidae" genus="Meximalus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="heratyi">
<emphasis id="B965EABFBA08FFBDFC372B5AFB8BF994" box="[941,1063,1635,1660]" italics="true" pageId="11" pageNumber="12">M. heratyi</emphasis>
</taxonomicName>
are few, but sufficient to recognize them as different species. The differences in femoral length/width between the two species are more than that exhibited by most pteromaline genera, where pro­ and/ or metafemoral differences of this magnitude constitute a relatively useful character helping to distinguish certain groups of genera.
</paragraph>
<paragraph id="8BAE36ADBA0FFFBAFE922C25FE4EFEDD" blockId="12.[264,482,283,310]" box="[264,482,283,310]" pageId="12" pageNumber="13">
<taxonomicName id="4C114D2EBA0FFFBAFE922C25FE14FEDD" authority="Bou" authorityName="Bou" box="[264,440,283,310]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFE922C25FE14FEDD" bold="true" box="[264,440,283,310]" pageId="12" pageNumber="13">
<emphasis id="B965EABFBA0FFFBAFE922C25FED1FEDE" bold="true" box="[264,381,284,310]" italics="true" pageId="12" pageNumber="13">Euteloida</emphasis>
Bou
</emphasis>
</taxonomicName>
č
<emphasis id="B965EABFBA0FFFBAFE5E2C22FE4EFEDD" bold="true" box="[452,482,283,309]" pageId="12" pageNumber="13">ek</emphasis>
</paragraph>
<paragraph id="8BAE36ADBA0FFFBAFE922C5EFEE3FE1B" blockId="12.[264,1323,359,1900]" pageId="12" pageNumber="13">
We have found several specimens, of this previously monospecific genus, from Southern California, where it appears to be relatively common. This constitutes the first discovery of males of the genus and the first reports of the genus aside from the
<typeStatus id="54AA880FBA0FFFBAFBDD2C8AFB02FE25" box="[1095,1198,435,461]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C114D2EBA0FFFBAFB4D2C8AFEEBFE1A" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subnigra">
<emphasis id="B965EABFBA0FFFBAFB4D2C8AFEEBFE1A" italics="true" pageId="12" pageNumber="13">E. subnigra</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAE36ADBA0FFFBAFEA22CC7FC05FD8C" blockId="12.[264,1323,359,1900]" pageId="12" pageNumber="13">
Bouček (1993) described
<taxonomicName id="4C114D2EBA0FFFBAFDF72CC6FD73FDF0" box="[621,735,511,536]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFDF72CC6FD73FDF0" box="[621,735,511,536]" italics="true" pageId="12" pageNumber="13">Euteloida</emphasis>
</taxonomicName>
with little comment on its placement and differences from other genera. With the discovery of a new species, the list of diagnostic characters of the genus relative to similar genera can be refined.
</paragraph>
<paragraph id="8BAE36ADBA0FFFBAFEA22F49FDB7FD3E" blockId="12.[264,1323,359,1900]" pageId="12" pageNumber="13">
<emphasis id="B965EABFBA0FFFBAFEA22F49FE26FD62" bold="true" box="[312,394,624,650]" pageId="12" pageNumber="13">Males.</emphasis>
Flagellum (Fig. 11) with few, irregularly placed longitudinal sensilla but with many curved flagellar setae. Club not acuminate, with symmetrical sutures and no conspicuous sensory areas.
</paragraph>
</subSubSection>
<subSubSection id="C30B6526BA0FFFBAFEA22FD8FDEDFBED" pageId="12" pageNumber="13" type="description">
<paragraph id="8BAE36ADBA0FFFBAFEA22FD8FDEDFBED" blockId="12.[264,1323,359,1900]" pageId="12" pageNumber="13">
<emphasis id="B965EABFBA0FFFBAFEA22FD8FE69FD13" bold="true" box="[312,453,737,763]" pageId="12" pageNumber="13">Both sexes.</emphasis>
The deep lateral gap between the pronotal collar and mesoscutum (Fig. 12) was cited by Bouček as a generic character, but we have found this character to be only occasionally useful and frequently misleading because it can be affected by the position of the body at death. Additionally, the extreme reduction or lack of a posterior smooth strip of the pronotal collar seemed to be a significant character, but the smooth strip is conspicuous and relatively long in males and small females. Nucha (Fig. 13) with a weak basal crosscarina merging smoothly with posterior remnants of plicae; variably emarginate posteriorly, not always deeply so.
</paragraph>
</subSubSection>
<subSubSection id="C30B6526BA0FFFBBFEA22929FBDDFDD7" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BAE36ADBA0FFFBBFEA22929FC5EFEB4" blockId="12.[264,1323,359,1900]" lastBlockId="13.[264,1323,284,575]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B965EABFBA0FFFBAFEA22929FE6FFBC2" bold="true" box="[312,451,1040,1066]" pageId="12" pageNumber="13">Discussion.</emphasis>
<taxonomicName id="4C114D2EBA0FFFBAFE542928FDECFBC2" box="[462,576,1041,1066]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFE542928FDECFBC2" box="[462,576,1041,1066]" italics="true" pageId="12" pageNumber="13">Euteloida</emphasis>
</taxonomicName>
is similar to some other pteromaline genera having the propodeal plicae smoothly merging with an isolated basal cross­carina of the nucha but lacking a distinct median carina, such as the Palearctic
<taxonomicName id="4C114D2EBA0FFFBAFCEA2964FB1BFB9E" authority="Boucek. However" authorityName="Boucek. However" box="[880,1207,1116,1142]" class="Insecta" family="Pteromalidae" genus="Lyubana" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFCEA2964FC7AFB9E" box="[880,982,1117,1142]" italics="true" pageId="12" pageNumber="13">Lyubana</emphasis>
Bouček. However
</taxonomicName>
,
<taxonomicName id="4C114D2EBA0FFFBAFB5F2964FA87FB9E" box="[1221,1323,1117,1142]" class="Insecta" family="Pteromalidae" genus="Lyubana" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFB5F2964FA87FB9E" box="[1221,1323,1117,1142]" italics="true" pageId="12" pageNumber="13">Lyubana</emphasis>
</taxonomicName>
has a genal carina, a narrowly and deeply incised clypeal margin with a pair of sharp and narrow denticles, and flagellum with 3 anelli and 5 funicular segments. Each of these characters can be variable between closely related pteromaline genera and even within some genera, such that it is difficult to discern a clear phylogenetic pattern using them. The Australasian genus
<taxonomicName id="4C114D2EBA0FFFBAFE522823FD3EFADC" authority="Boucek" authorityName="Boucek" box="[456,658,1306,1332]" class="Insecta" family="Pteromalidae" genus="Nuchata" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFE522823FD80FADB" box="[456,556,1306,1331]" italics="true" pageId="12" pageNumber="13">Nuchata</emphasis>
Bouček
</taxonomicName>
is similar to
<taxonomicName id="4C114D2EBA0FFFBAFCA42823FC1CFADB" box="[830,944,1306,1331]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFCA42823FC1CFADB" box="[830,944,1306,1331]" italics="true" pageId="12" pageNumber="13">Euteloida</emphasis>
</taxonomicName>
in claval and propodeal form, including the bump­like callar elevations.
<taxonomicName id="4C114D2EBA0FFFBAFD6F2879FCF4FAB1" box="[757,856,1344,1369]" class="Insecta" family="Pteromalidae" genus="Nuchata" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFD6F2879FCF4FAB1" box="[757,856,1344,1369]" italics="true" pageId="12" pageNumber="13">Nuchata</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C114D2EBA0FFFBAFC692879FBC9FAB1" box="[1011,1125,1344,1369]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFC692879FBC9FAB1" box="[1011,1125,1344,1369]" italics="true" pageId="12" pageNumber="13">Euteloida</emphasis>
</taxonomicName>
in having a faint occipital carina, a strong genal carina, and a longer nucha relative to the rest of the propodeum.
<taxonomicName id="4C114D2EBA0FFFBAFEC628B5FDC8FA4D" authority="Graham" authorityName="Graham" box="[348,612,1419,1445]" class="Insecta" family="Pteromalidae" genus="Sceptrothelys" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFEC628B5FE55FA4D" box="[348,505,1420,1445]" italics="true" pageId="12" pageNumber="13">Sceptrothelys</emphasis>
Graham
</taxonomicName>
is also similar in claval form and in general reduction to
<taxonomicName id="4C114D2EBA0FFFBAFE922888FED6FA22" box="[264,378,1457,1482]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFE922888FED6FA22" box="[264,378,1457,1482]" italics="true" pageId="12" pageNumber="13">Euteloida</emphasis>
</taxonomicName>
, but has a much different propodeum with a long, cylindrical nucha, strong plicae extending from anterior plical pits to near nuchal apex, and no nuchal cross­carina.
<taxonomicName id="4C114D2EBA0FFFBAFE9228C4FE76F9FF" authority="Walker" authorityName="Walker" box="[264,474,1533,1559]" class="Insecta" family="Pteromalidae" genus="Norbanus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFE9228C4FED0F9FE" box="[264,380,1533,1558]" italics="true" pageId="12" pageNumber="13">Norbanus</emphasis>
Walker
</taxonomicName>
and some species groups of
<taxonomicName id="4C114D2EBA0FFFBAFCA028C4FBECF9FF" authority="Thomson" authorityName="Thomson" box="[826,1088,1533,1559]" class="Insecta" family="Pteromalidae" genus="Homoporus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFCA028C4FC69F9FE" box="[826,965,1533,1558]" italics="true" pageId="12" pageNumber="13">Homoporus</emphasis>
Thomson
</taxonomicName>
, along with several related genera, are similar to
<taxonomicName id="4C114D2EBA0FFFBAFDFE2B1AFD7AF9D4" box="[612,726,1571,1596]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFDFE2B1AFD7AF9D4" box="[612,726,1571,1596]" italics="true" pageId="12" pageNumber="13">Euteloida</emphasis>
</taxonomicName>
in having a relatively acarinate propodeum that is uniformly alveolate, but they have a distinct claval spicule in females and differ in a number of other characters, especially in having a propodeal costula and lacking posterior plicae (when propodeal carinae are discernable), such that they would not seem to be especially close to
<taxonomicName id="4C114D2EBA0FFFBAFE732B82FDF7F93C" box="[489,603,1723,1748]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFE732B82FDF7F93C" box="[489,603,1723,1748]" italics="true" pageId="12" pageNumber="13">Euteloida</emphasis>
</taxonomicName>
. We suspect that the basal cross­carina of the nucha in
<taxonomicName id="4C114D2EBA0FFFBAFB712B82FEE8F912" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFB712B82FEE8F912" italics="true" pageId="12" pageNumber="13">Euteloida</emphasis>
</taxonomicName>
is homologous with the more distinct nuchal carina found in most pteromaline genera such as
<taxonomicName id="4C114D2EBA0FFFBAFE142A3FFD86F8F7" box="[398,554,1798,1823]" class="Insecta" family="Pteromalidae" genus="Mesopolobus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0FFFBAFE142A3FFD86F8F7" box="[398,554,1798,1823]" italics="true" pageId="12" pageNumber="13">Mesopolobus</emphasis>
</taxonomicName>
, and that the apparent lack of separation between it and the plicae is due to the presence of these carinae only across the base of a convex nucha and their lack of connection to any other carinae or to either propodeal margin. This state constitutes a distinction between
<taxonomicName id="4C114D2EBA0EFFBBFD8E2C25FD2AFEDD" box="[532,646,284,309]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0EFFBBFD8E2C25FD2AFEDD" box="[532,646,284,309]" italics="true" pageId="13" pageNumber="14">Euteloida</emphasis>
</taxonomicName>
and nearly all other pteromaline genera. Unfortunately, this carina is not easily visible in many specimens of
<taxonomicName id="4C114D2EBA0EFFBBFCE32C7BFC47FEB3" box="[889,1003,322,347]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0EFFBBFCE32C7BFC47FEB3" box="[889,1003,322,347]" italics="true" pageId="13" pageNumber="14">Euteloida</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAE36ADBA0EFFBBFEA22C51FBDDFDD7" blockId="13.[264,1323,284,575]" pageId="13" pageNumber="14">
The revised diagnosis of
<taxonomicName id="4C114D2EBA0EFFBBFDF32C51FD77FE69" box="[617,731,360,385]" class="Insecta" family="Pteromalidae" genus="Euteloida" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B965EABFBA0EFFBBFDF32C51FD77FE69" box="[617,731,360,385]" italics="true" pageId="13" pageNumber="14">Euteloida</emphasis>
</taxonomicName>
can be summarized as follows: flagellum with 2 typical anelli; club with asymmetrical sutures in females and apical claval segment with two distinct ventral micropilose areas; genal carina absent; clypeal margin very shallowly emarginate; left mandible with 3 denticles, the right with 4; pronotal collar not carinate; notauli incomplete; median carina and plicae incomplete; nucha convex but short, hardly projecting beyond supracoxal flange at metacoxal bases; 1 metatibial spur.
</paragraph>
</subSubSection>
</treatment>
</document>