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<document id="2E919CB110F915D4DD4DB563E7A762C9" ID-DOI="10.11646/zootaxa.5375.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10200036" ID-ZooBank="27D02F09-01B7-457A-8A99-D8644B7B6ADE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1700728652862" checkinUser="plazi" docAuthor="Arimoto, Kôichi" docDate="2023" docId="03AE8A2DFFBECF7FFF47E994FD2AF888" docLanguage="en" docName="zootaxa.5375.3.1.pdf" docOrigin="Zootaxa 5375 (3)" docSource="https://mapress.com/zt/article/download/zootaxa.5375.3.1/52332" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Penia takasago Kishii 1997" docType="treatment" docVersion="4" lastPageNumber="328" masterDocId="FF97F255FFA6CF64FFD0EB1BFFFFFFED" masterDocTitle="Taxonomic revision of the genus Penia Laporte (Coleoptera, Elateridae, Dendrometrinae) from Taiwan" masterLastPageNumber="335" masterPageNumber="301" pageNumber="325" updateTime="1700830679711" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="4A224002E0431636CB7F00151EF9377B">Taxonomic revision of the genus Penia Laporte (Coleoptera, Elateridae, Dendrometrinae) from Taiwan</mods:title>
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<mods:date id="B7210E137F9E1105BA14B0728B3BEEAB">2023</mods:date>
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<treatment id="03AE8A2DFFBECF7FFF47E994FD2AF888" ID-DOI="http://doi.org/10.5281/zenodo.10201492" ID-Zenodo-Dep="10201492" LSID="urn:lsid:plazi:treatment:03AE8A2DFFBECF7FFF47E994FD2AF888" httpUri="http://treatment.plazi.org/id/03AE8A2DFFBECF7FFF47E994FD2AF888" lastPageId="27" lastPageNumber="328" pageId="24" pageNumber="325">
<subSubSection id="C31D68B0FFBECF7CFF47E994FE13FD45" box="[151,492,654,681]" pageId="24" pageNumber="325" type="nomenclature">
<paragraph id="8BB83B3BFFBECF7CFF47E994FE13FD45" blockId="24.[151,492,654,717]" box="[151,492,654,681]" pageId="24" pageNumber="325">
<heading id="D0F08C57FFBECF7CFF47E994FE13FD45" bold="true" box="[151,492,654,681]" fontSize="11" level="1" pageId="24" pageNumber="325" reason="1">
<taxonomicName id="4C0740B8FFBECF7CFF47E994FE13FD45" authority="Kishii, 1997" authorityName="Kishii" authorityYear="1997" box="[151,492,654,681]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBECF7CFF47E994FE13FD45" bold="true" box="[151,492,654,681]" pageId="24" pageNumber="325">
<emphasis id="B973E729FFBECF7CFF47E994FEAEFD44" bold="true" box="[151,337,654,681]" italics="true" pageId="24" pageNumber="325">Penia takasago</emphasis>
<bibRefCitation id="EF9646CAFFBECF7CFE87E995FE13FD45" author="Kishii, T." box="[343,492,654,680]" pageId="24" pageNumber="325" pagination="9 - 14" refId="ref23002" refString="Kishii, T. (1997) A study on the elaterid-beetles of Shibata collection from Taiwan. V. (Coleoptera: Elateridae) On the subfamily Denticollinae: tribes Denticollini, Hemicrepidiini and Dimini. The Entomological Review of Japan, 52, 9 - 14." type="journal article" year="1997">Kishii, 1997</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFBECF7CFF47E9A8FEB4FD20" box="[151,331,691,717]" pageId="24" pageNumber="325" type="description">
<paragraph id="8BB83B3BFFBECF7CFF47E9A8FEB4FD20" blockId="24.[151,492,654,717]" box="[151,331,691,717]" pageId="24" pageNumber="325">
(
<figureCitation id="133C27BEFFBECF7CFF4FE9A8FEE5FD20" box="[159,282,691,717]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="24" pageNumber="325">Figures 16</figureCitation>
,
<figureCitation id="133C27BEFFBECF7CFEF6E9A8FEBCFD20" box="[294,323,691,717]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="24" pageNumber="325">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFBECF7CFF47E9E3FBE9FCC3" pageId="24" pageNumber="325" type="reference_group">
<paragraph id="8BB83B3BFFBECF7CFF47E9E3FBE9FCC3" blockId="24.[151,1437,759,814]" pageId="24" pageNumber="325">
<treatmentCitationGroup id="AB171C15FFBECF7CFF47E9E3FBE9FCC3" pageId="24" pageNumber="325">
<treatmentCitation id="0AA61D2AFFBECF7CFF47E9E3FE25FCE2" author="Kishii, T." box="[151,474,759,783]" page="12" pageId="24" pageNumber="325" year="1997">
<taxonomicName id="4C0740B8FFBECF7CFF47E9E3FE25FCE2" authority="Kishii, 1997: 12" authorityName="Kishii" authorityPageNumber="12" authorityYear="1997" box="[151,474,759,783]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBECF7CFF47E9E3FECCFCE2" box="[151,307,759,783]" italics="true" pageId="24" pageNumber="325">Penia takasago</emphasis>
<bibRefCitation id="EF9646CAFFBECF7CFEE9E9ECFE25FCE2" author="Kishii, T." box="[313,474,759,783]" pageId="24" pageNumber="325" pagination="9 - 14" refId="ref23002" refString="Kishii, T. (1997) A study on the elaterid-beetles of Shibata collection from Taiwan. V. (Coleoptera: Elateridae) On the subfamily Denticollinae: tribes Denticollini, Hemicrepidiini and Dimini. The Entomological Review of Japan, 52, 9 - 14." type="journal article" year="1997">Kishii, 1997: 12</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description;
<typeStatus id="54BC8599FFBECF7CFD6AE9E3FD19FCFD" box="[698,742,760,784]" pageId="24" pageNumber="325">type</typeStatus>
locality:
<collectingCountry id="F3107BABFFBECF7CFC93E9ECFC6FFCE2" box="[835,912,759,783]" name="Taiwan" pageId="24" pageNumber="325">Taiwan</collectingCountry>
,
<collectingRegion id="49C3F5D9FFBECF7CFC49E9E3FBCAFCE2" box="[921,1077,759,783]" country="Taiwan" name="Nantou" pageId="24" pageNumber="325">Nantou County</collectingRegion>
, Sungkang) [partim];
<treatmentCitation id="0AA61D2AFFBECF7CFAC0E9ECFF12FCC3" author="Suzuki, W." page="121" pageId="24" pageNumber="325" year="1999">
<bibRefCitation id="EF9646CAFFBECF7CFAC0E9ECFF12FCC3" author="Suzuki, W." pageId="24" pageNumber="325" pagination="1 - 348" refId="ref23700" refString="Suzuki, W. (1999) Catalogue of the family Elateridae (Coleoptera) of Taiwan. Miscellaneous Reports of the Hiwa Museum for Natural History, 38, 1 - 348." type="journal article" year="1999">Suzuki, 1999: 121</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AA61D2AFFBECF7CFEA3E80DFDEBFCC3" author="Cate, P. C." box="[371,532,790,814]" page="185" pageId="24" pageNumber="325" year="2007">
<bibRefCitation id="EF9646CAFFBECF7CFEA3E80DFDEBFCC3" author="Cate, P. C." box="[371,532,790,814]" pageId="24" pageNumber="325" refId="ref22578" refString="Cate, P. C. (2007) Family Elateridae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209." type="book" year="2007">Cate, 2007: 185</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AA61D2AFFBECF7CFD4AE80DFC68FCC3" ID-DOI="http://doi.org/10.5281/zenodo.5985900" ID-Zenodo-Dep="5985900" author="Kundrata, R. &amp; Musalkova, M. &amp; Kubaczkova, M." box="[666,919,790,814]" httpUri="http://treatment.plazi.org/id/A648DA7791759B0FFF5115B9FD9ADE86" page="56" pageId="24" pageNumber="325" year="2018">
<bibRefCitation id="EF9646CAFFBECF7CFD4AE80DFC68FCC3" author="Kundrata, R. &amp; Musalkova, M. &amp; Kubaczkova, M." box="[666,919,790,814]" pageId="24" pageNumber="325" pagination="1 - 75" refId="ref23051" refString="Kundrata, R., Musalkova, M. &amp; Kubaczkova, M. (2018) Annotated catalogue of the click-beetle tribe Dimini (Coleoptera: Elateridae: Dendrometrinae). Zootaxa, 4412 (1), 1 - 75. https: // doi. org / 10.11646 / zootaxa. 4412.1" type="journal article" year="2018">
Kundrata
<emphasis id="B973E729FFBECF7CFD2EE80CFCCEFCC3" box="[766,817,790,814]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
, 2018: 56
</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFBECF7CFF47E847FD0BF88A" pageId="24" pageNumber="325" type="materials_examined">
<paragraph id="8BB83B3BFFBECF7CFF47E847FB50FBEB" blockId="24.[151,1437,860,2039]" pageId="24" pageNumber="325">
<emphasis id="B973E729FFBECF7CFF47E847FEBDFC9B" bold="true" box="[151,322,860,886]" pageId="24" pageNumber="325">Type material.</emphasis>
<materialsCitation id="3B6F3166FFBECF7CFE97E847FBBDFC52" collectingDate="1969-07-21" collectingDateMax="1969-08-21" collectingDateMin="1969-07-21" collectionCode="OMNH, MAEDA, T" country="Taiwan" location="Nantou County" pageId="24" pageNumber="325" specimenCode="PTK01" specimenCount="2" stateProvince="Nantou" typeStatus="holotype">
<emphasis id="B973E729FFBECF7CFE97E847FE44FC9B" bold="true" box="[327,443,860,886]" pageId="24" pageNumber="325">
<typeStatus id="54BC8599FFBECF7CFE97E847FE49FC9B" box="[327,438,860,886]" pageId="24" pageNumber="325" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectorName id="26F25EEDFFBECF7CFE6FE847FDEBFC9B" box="[447,532,860,886]" pageId="24" pageNumber="325">Female</collectorName>
,
<collectorName id="26F25EEDFFBECF7CFDCCE847FD8EFC9B" box="[540,625,860,886]" pageId="24" pageNumber="325">Taiwan</collectorName>
,
<collectingRegion id="49C3F5D9FFBECF7CFDAAE846FCDAFC9B" box="[634,805,860,887]" country="Taiwan" name="Nantou" pageId="24" pageNumber="325">Nantou County</collectingRegion>
,
<collectorName id="26F25EEDFFBECF7CFCFDE847FC5FFC9B" box="[813,928,860,886]" pageId="24" pageNumber="325">Sungkang</collectorName>
,
<date id="FFB91DFBFFBECF7CFC78E847FBD1FC9B" box="[936,1070,860,887]" pageId="24" pageNumber="325" value="1969-07-21">
<collectingDate id="EFFDE413FFBECF7CFC78E847FBD1FC9B" box="[936,1070,860,887]" pageId="24" pageNumber="325" value="1969-07-21">21 VII 1969</collectingDate>
</date>
,
<collectorName id="26F25EEDFFBECF7CFBE6E846FB61FC9B" box="[1078,1182,860,887]" pageId="24" pageNumber="325">Y. Maeda</collectorName>
leg. [
<collectionCode id="ED16A3FEFFBECF7CFB06E847FACDFC9B" box="[1238,1330,860,886]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="24" pageNumber="325" type="Museum">OMNH</collectionCode>
;
<specimenCode id="DBA19340FFBECF7CFAEBE847FA6DFC9B" box="[1339,1426,860,886]" collectionCode="PTK" pageId="24" pageNumber="325">PTK01</specimenCode>
]. Verbatim label data (
<figureCitation id="133C27BEFFBECF7CFEAEE89BFE1AFC77" box="[382,485,896,922]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="24" pageNumber="325">Fig. 16A</figureCitation>
). “HOLOTYPE”; “SUNGKANG/
<collectingCountry id="F3107BABFFBECF7CFCBAE89BFC0EFC77" box="[874,1009,896,922]" name="Taiwan" pageId="24" pageNumber="325">FORMOSA</collectingCountry>
/
<date id="FFB91DFBFFBECF7CFBD0E89BFB62FC77" box="[1024,1181,896,923]" pageId="24" pageNumber="325" value="1969-08-21">
<collectingDate id="EFFDE413FFBECF7CFBD0E89BFB62FC77" box="[1024,1181,896,923]" pageId="24" pageNumber="325" value="1969-08-21">21. VIII. 1969</collectingDate>
</date>
/ Y.
<collectionCode id="ED16A3FEFFBECF7CFB19E89BFAC9FC77" box="[1225,1334,896,922]" pageId="24" pageNumber="325">MAEDA</collectionCode>
”: “
<taxonomicName id="4C0740B8FFBECF7CFA83E89BFA6CFC77" box="[1363,1427,896,922]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="335" phylum="Arthropoda" rank="genus">Penia</taxonomicName>
/ takasago/ KISHII,
<taxonomicNameLabel id="A2405A52FFBECF7CFEA0E8BEFE51FC53" box="[368,430,932,959]" pageId="24" pageNumber="325" rank="species">sp. n.</taxonomicNameLabel>
/ Det. KISHII, ’92”; “
<collectionCode id="ED16A3FEFFBECF7CFD7FE8BFFD3DFC53" box="[687,706,932,958]" name="Tavera, Department of Geology and Geophysics" pageId="24" pageNumber="325">T</collectionCode>
. KISHII Coll./
<collectionCode id="ED16A3FEFFBECF7CFCA7E8BFFC33FC53" box="[887,972,932,958]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="24" pageNumber="325" type="Museum">OMNH</collectionCode>
/ [15-16]”
</materialsCitation>
.
<materialsCitation id="3B6F3166FFBECF7CFB80E8BFFD99FC0E" collectingDate="1969-08-07" collectionCode="T" country="Taiwan" location="Musha" municipality="Taiwan" pageId="24" pageNumber="325" specimenCount="1" specimenCount-female="1" stateProvince="Nantou" typeStatus="paratype">
<emphasis id="B973E729FFBECF7CFB80E8BFFB3BFC53" bold="true" box="[1104,1220,932,958]" pageId="24" pageNumber="325">
<typeStatus id="54BC8599FFBECF7CFB80E8BFFB40FC53" box="[1104,1215,932,958]" pageId="24" pageNumber="325" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCount id="9D01F0B2FFBECF7CFB1EE8BFFACAFC53" box="[1230,1333,932,958]" count="1" pageId="24" pageNumber="325" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6BDCA141FFBECF7CFA93E8BFFA67FC53" box="[1347,1432,932,958]" pageId="24" pageNumber="325">Taiwan</collectingMunicipality>
,
<collectingRegion id="49C3F5D9FFBECF7CFF47E8D2FEB6FC0F" box="[151,329,968,995]" country="Taiwan" name="Nantou" pageId="24" pageNumber="325">Nantou County</collectingRegion>
,
<location id="8ED86DE0FFBECF7CFE88E8D3FE58FC0F" LSID="urn:lsid:plazi:treatment:03AE8A2DFFBECF7FFF47E994FD2AF888:8ED86DE0FFBECF7CFE88E8D3FE58FC0F" box="[344,423,968,994]" country="Taiwan" municipality="Taiwan" name="Musha" pageId="24" pageNumber="325" stateProvince="Nantou">Musha</location>
,
<date id="FFB91DFBFFBECF7CFE66E8D2FDBAFC0F" box="[438,581,968,995]" pageId="24" pageNumber="325" value="1969-08-07">
<collectingDate id="EFFDE413FFBECF7CFE66E8D2FDBAFC0F" box="[438,581,968,995]" pageId="24" pageNumber="325" value="1969-08-07">7 VIII 1969</collectingDate>
</date>
,
<collectionCode id="ED16A3FEFFBECF7CFD84E8D2FD99FC0E" box="[596,614,969,995]" name="Tavera, Department of Geology and Geophysics" pageId="24" pageNumber="325">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B6F3166FFBECF7CFDA5E8D3FC03FC0F" box="[629,1020,968,994]" collectionCode="OMNH" collectorName="Kobayashi" country="Taiwan" pageId="24" pageNumber="325" specimenCode="PTK02" specimenCount="1" stateProvince="Nantou" typeStatus="paratype">
<collectorName id="26F25EEDFFBECF7CFDA5E8D3FD12FC0F" box="[629,749,968,994]" pageId="24" pageNumber="325">Kobayashi</collectorName>
leg. [
<collectionCode id="ED16A3FEFFBECF7CFCE3E8D3FC70FC0F" box="[819,911,968,994]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="24" pageNumber="325" type="Museum">OMNH</collectionCode>
;
<specimenCode id="DBA19340FFBECF7CFC4EE8D3FC0AFC0F" box="[926,1013,968,994]" collectionCode="PTK" pageId="24" pageNumber="325">PTK02</specimenCode>
]
</materialsCitation>
;
<materialsCitation id="3B6F3166FFBECF7CFBDCE8D3FB54FBEB" collectingDate="1969-07-06" collectingDateMax="1969-08-06" collectingDateMin="1969-07-06" collectionCode="OMNH" collectorName="Y. Maeda" country="Taiwan" location="Sungkang" municipality="Taiwan" pageId="24" pageNumber="325" specimenCount="1" specimenCount-female="1" stateProvince="Nantou" typeStatus="paratype">
<specimenCount id="9D01F0B2FFBECF7CFBDCE8D3FB8BFC0F" box="[1036,1140,968,994]" count="1" pageId="24" pageNumber="325" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6BDCA141FFBECF7CFB53E8D3FB27FC0F" box="[1155,1240,968,994]" pageId="24" pageNumber="325">Taiwan</collectingMunicipality>
,
<collectingRegion id="49C3F5D9FFBECF7CFB37E8D2FA66FC0F" box="[1255,1433,968,995]" country="Taiwan" name="Nantou" pageId="24" pageNumber="325">Nantou County</collectingRegion>
,
<location id="8ED86DE0FFBECF7CFF47E8F7FEF5FBEB" LSID="urn:lsid:plazi:treatment:03AE8A2DFFBECF7FFF47E994FD2AF888:8ED86DE0FFBECF7CFF47E8F7FEF5FBEB" box="[151,266,1004,1030]" country="Taiwan" municipality="Taiwan" name="Sungkang" pageId="24" pageNumber="325" stateProvince="Nantou">Sungkang</location>
,
<date id="FFB91DFBFFBECF7CFEC5E8F7FE66FBEB" box="[277,409,1004,1031]" pageId="24" pageNumber="325" value="1969-08-06">
<collectingDate id="EFFDE413FFBECF7CFEC5E8F7FE66FBEB" box="[277,409,1004,1031]" pageId="24" pageNumber="325" value="1969-08-06">6 VIII 1969</collectingDate>
</date>
(as
<date id="FFB91DFBFFBECF7CFE18E8F7FDBDFBEB" box="[456,578,1004,1031]" pageId="24" pageNumber="325" value="1969-07-06">
<collectingDate id="EFFDE413FFBECF7CFE18E8F7FDBDFBEB" box="[456,578,1004,1031]" pageId="24" pageNumber="325" value="1969-07-06">
6 VII
<quantity id="4CFF96DEFFBECF7CFDDAE8F7FDBDFBEB" box="[522,578,1004,1030]" metricMagnitude="1" metricUnit="m" metricValue="5.00126" pageId="24" pageNumber="325" unit="in" value="1969.0">1969</quantity>
</collectingDate>
</date>
in original description),
<collectorName id="26F25EEDFFBECF7CFC86E8F6FC3EFBEB" box="[854,961,1004,1031]" pageId="24" pageNumber="325">Y. Maeda</collectorName>
leg. [
<collectionCode id="ED16A3FEFFBECF7CFC2FE8F7FBA4FBEB" box="[1023,1115,1004,1030]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="24" pageNumber="325" type="Museum">OMNH</collectionCode>
; 6348]
</materialsCitation>
.
</paragraph>
<paragraph id="8BB83B3BFFBECF7CFF17EF0BFABBFA7F" blockId="24.[151,1437,860,2039]" pageId="24" pageNumber="325">
<emphasis id="B973E729FFBECF7CFF17EF0BFE58FBC7" bold="true" box="[199,423,1040,1066]" pageId="24" pageNumber="325">Female. Diagnosis.</emphasis>
This species is characterized by the following features: eyes 0.4 x longer than interocular distance in dorsal view; antennae extending beyond pronotum posterior lateral apices by antennomere
<collectionCode id="ED16A3FEFFBECF7CFB2CEF2EFAE3FBA2" box="[1276,1308,1077,1103]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="24" pageNumber="325">VI</collectionCode>
, surpassing elytral half by antennomere XI; antennomeres
<collectionCode id="ED16A3FEFFBECF7CFD79EF42FD3AFB9E" box="[681,709,1113,1139]" pageId="24" pageNumber="325">III</collectionCode>
distinctly longer than
<collectionCode id="ED16A3FEFFBECF7CFC15EF42FC24FB9E" box="[965,987,1113,1139]" pageId="24" pageNumber="325">II</collectionCode>
;
<collectionCode id="ED16A3FEFFBECF7CFC38EF42FBFAFB9E" box="[1000,1029,1113,1139]" pageId="24" pageNumber="325">IV</collectionCode>
1.3–1.4 x longer than
<collectionCode id="ED16A3FEFFBECF7CFAD9EF42FAD7FB9E" box="[1289,1320,1113,1139]" pageId="24" pageNumber="325">III</collectionCode>
, 0.8–0.85 x longer than
<collectionCode id="ED16A3FEFFBECF7CFEEDEF66FEB0FB7A" box="[317,335,1149,1175]" pageId="24" pageNumber="325">II</collectionCode>
–
<collectionCode id="ED16A3FEFFBECF7CFE8EEF66FE85FB7A" box="[350,378,1149,1175]" pageId="24" pageNumber="325">III</collectionCode>
combined; apical maxillary palpomere 2.3–2.5 x longer than wide, shorter than maximum length of eye; pronotum strongly and straightly narrowed anterior to hind angles; posterior edge of pronotum with sublateral incisions; hind angles of pronotum acute, strongly protruding posterolaterad; hypomeron with moderate mesial projection; anterior angle of hypomeron rounded; hind angle of hypomeron claw-like shaped; scutellar shield almost as long as wide; mesosternal process between mesocoxae lower than mesocoxae, not visible in lateral view; posterior edge of mesosternal process 0.1–0.15 x wider than total width of mesosternum; elytron 3.4–3.6 x longer than wide, 2.9–3.0 x longer than pronotum length; abdominal ventrite
<collectionCode id="ED16A3FEFFBECF7CFC24EE4EFBF7FA82" box="[1012,1032,1365,1391]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="24" pageNumber="325" type="Museum">V</collectionCode>
curved triangular, rounded apically; spiculum ventrale 3.9–4.6 x longer than length of sternite VIII; ovipositor shorter than length of abdomen.
</paragraph>
<paragraph id="8BB83B3BFFBECF7CFF17EE86FD10F93B" blockId="24.[151,1437,860,2039]" pageId="24" pageNumber="325">
In
<collectingCountry id="F3107BABFFBECF7CFF34EE87FEC6FA5B" box="[228,313,1436,1462]" name="Taiwan" pageId="24" pageNumber="325">Taiwan</collectingCountry>
, five species with broad bodies are known:
<taxonomicName id="4C0740B8FFBECF7CFCCFEE86FC82FA5B" authorityName="Kishii" authorityYear="1994" box="[799,893,1436,1463]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B973E729FFBECF7CFCCFEE86FCCFFA5B" box="[799,816,1437,1462]" italics="true" pageId="24" pageNumber="325">P</emphasis>
.
<emphasis id="B973E729FFBECF7CFCEDEE87FC82FA5B" box="[829,893,1436,1462]" italics="true" pageId="24" pageNumber="325">babai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0740B8FFBECF7CFC5BEE86FBEBFA5B" authorityName="Arimoto" authorityYear="2023" box="[907,1044,1437,1463]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="inopinata">
<emphasis id="B973E729FFBECF7CFC5BEE86FC63FA5B" box="[907,924,1437,1462]" italics="true" pageId="24" pageNumber="325">P</emphasis>
.
<emphasis id="B973E729FFBECF7CFC79EE86FBEBFA5B" box="[937,1044,1437,1462]" italics="true" pageId="24" pageNumber="325">inopinata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0740B8FFBECF7CFBF2EE86FB84FA5B" authorityName="Arimoto" authorityYear="2023" box="[1058,1147,1436,1463]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="pulla">
<emphasis id="B973E729FFBECF7CFBF2EE86FBCCFA5B" box="[1058,1075,1437,1462]" italics="true" pageId="24" pageNumber="325">P</emphasis>
.
<emphasis id="B973E729FFBECF7CFB91EE87FB84FA5B" box="[1089,1147,1436,1462]" italics="true" pageId="24" pageNumber="325">pulla</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0740B8FFBECF7CFB58EE86FAF4FA5B" authorityName="Kishii" authorityYear="1997" box="[1160,1291,1436,1463]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBECF7CFB58EE86FB66FA5B" box="[1160,1177,1437,1462]" italics="true" pageId="24" pageNumber="325">P</emphasis>
.
<emphasis id="B973E729FFBECF7CFB76EE87FAF4FA5B" box="[1190,1291,1436,1462]" italics="true" pageId="24" pageNumber="325">takasago</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C0740B8FFBECF7CFA98EE86FA69FA5B" authorityName="Arimoto" authorityYear="2023" box="[1352,1430,1437,1463]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="tsou">
<emphasis id="B973E729FFBECF7CFA98EE86FAA6FA5B" box="[1352,1369,1437,1462]" italics="true" pageId="24" pageNumber="325">P</emphasis>
.
<emphasis id="B973E729FFBECF7CFAB7EE86FA69FA5B" box="[1383,1430,1437,1462]" italics="true" pageId="24" pageNumber="325">tsou</emphasis>
</taxonomicName>
. Their elytra are less than 4× their widths. They are separated from the elongate species, whose elytra are more than 4× the width. Among broad-bodied species,
<taxonomicName id="4C0740B8FFBECF7CFD56EEFEFCF6FA13" authorityName="Kishii" authorityYear="1997" box="[646,777,1508,1535]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBECF7CFD56EEFEFD68FA13" box="[646,663,1509,1534]" italics="true" pageId="24" pageNumber="325">P</emphasis>
.
<emphasis id="B973E729FFBECF7CFD74EEFFFCF6FA13" box="[676,777,1508,1534]" italics="true" pageId="24" pageNumber="325">takasago</emphasis>
</taxonomicName>
is distinguished from the other congeners by the following characters (the other species in parentheses): eyes 0.4 x longer than interocular distance in dorsal view (eyes 0.2–0.3 x longer than interocular distance in dorsal view); OI: 177–179 (OI: 141–162); pronotum strongly and straightly narrowed anterior to hind angles (pronotum slightly narrowed or slightly widening anterior to hind angles); hind angles of pronotum acute (hind angles of pronotum broad); spiculum ventrale less than 5 x longer than length of sternite VIII (spiculum ventrale more than 5 x longer than length of sternite VIII); ovipositor shorter than length of abdomen (ovipositor longer than length of abdomen).
</paragraph>
<paragraph id="8BB83B3BFFBECF7CFF17EDFAFD0BF88A" blockId="24.[151,1437,860,2039]" pageId="24" pageNumber="325">
<taxonomicName id="4C0740B8FFBECF7CFF17EDFAFE8DF917" authorityName="Kishii" authorityYear="1997" box="[199,370,1760,1786]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBECF7CFF17EDFAFE8DF917" box="[199,370,1760,1786]" italics="true" pageId="24" pageNumber="325">Penia takasago</emphasis>
</taxonomicName>
is remarkable also due to the many sclerotized spines inside the bursa copulatrix and the thick sac extending from around the apex of the bursa copulatrix and splitting in two; however, it is impossible to determine if these features are useful for species identification because the bursa copulatrix in the
<typeStatus id="54BC8599FFBECF7CFB87EC33FB46F8AF" box="[1111,1209,1832,1858]" pageId="24" pageNumber="325" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C0740B8FFBECF7CFB0FEC32FA96F8AF" authorityName="Arimoto" authorityYear="2023" box="[1247,1385,1833,1859]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="inopinata">
<emphasis id="B973E729FFBECF7CFB0FEC32FB0FF8AF" box="[1247,1264,1833,1858]" italics="true" pageId="24" pageNumber="325">P</emphasis>
.
<emphasis id="B973E729FFBECF7CFB2EEC32FA96F8AF" box="[1278,1385,1833,1858]" italics="true" pageId="24" pageNumber="325">inopinata</emphasis>
</taxonomicName>
was lost and could not be compared to that of
<taxonomicName id="4C0740B8FFBECF7CFDB9EC56FD12F88B" authorityName="Kishii" authorityYear="1997" box="[617,749,1868,1895]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBECF7CFDB9EC56FD85F88B" box="[617,634,1869,1894]" italics="true" pageId="24" pageNumber="325">P</emphasis>
.
<emphasis id="B973E729FFBECF7CFD58EC57FD12F88B" box="[648,749,1868,1894]" italics="true" pageId="24" pageNumber="325">takasago</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFBECF7FFF17EC6BFE77F9A8" lastPageId="27" lastPageNumber="328" pageId="24" pageNumber="325" type="description">
<paragraph id="8BB83B3BFFBECF7CFF17EC6BFD5EF81B" blockId="24.[151,1437,860,2039]" pageId="24" pageNumber="325">
<emphasis id="B973E729FFBECF7CFF17EC6BFE89F867" bold="true" box="[199,374,1904,1930]" pageId="24" pageNumber="325">Measurements</emphasis>
(n=3;
<typeStatus id="54BC8599FFBECF7CFE10EC6BFDDDF867" box="[448,546,1904,1930]" pageId="24" pageNumber="325" type="holotype">holotype</typeStatus>
in parentheses). BL: 8.61–8.79 (8.79), BW: 3.43–3.63 (3.58), MAE: 1.33–1.39 (1.39), MBE: 0.75–0.78 (0.78), OI: 177–179 (178), PL: 2.09–2.17 (2.17), PML: 1.63–1.71 (1.71), PW: 2.49–2.66 (2.62), PAW: 1.40–1.51 (1.50), PLI: 81.1–84.1 (83.0), PWI: 175–178 (175), EL: 6.25–6.45 (6.45), EW: 1.74–1.84 (1.82), EI: 341–364 (354), BI: 289–303 (297).
</paragraph>
<caption id="DF786BB3FFBFCF7DFF47ED6DFCFFF88B" ID-DOI="http://doi.org/10.5281/zenodo.10200069" ID-Zenodo-Dep="10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="25" pageNumber="326" startId="25.[151,250,1654,1678]" targetBox="[151,1436,191,1630]" targetPageId="25" targetType="figure">
<paragraph id="8BB83B3BFFBFCF7DFF47ED6DFCFFF88B" blockId="25.[151,1437,1654,1894]" pageId="25" pageNumber="326">
<emphasis id="B973E729FFBFCF7DFF47ED6DFEDEF963" bold="true" box="[151,289,1654,1678]" pageId="25" pageNumber="326">FIGURE 16.</emphasis>
<taxonomicName id="4C0740B8FFBFCF7DFEF8ED6CFE39F963" authorityName="Kishii" authorityYear="1997" box="[296,454,1654,1678]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBFCF7DFEF8ED6CFE39F963" box="[296,454,1654,1678]" italics="true" pageId="25" pageNumber="326">Penia takasago</emphasis>
</taxonomicName>
, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus.
</paragraph>
</caption>
<paragraph id="8BB83B3BFFBFCF7EFF17EC8FFC53FF14" blockId="25.[151,1436,1940,2039]" lastBlockId="26.[151,1437,151,1977]" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326">
<emphasis id="B973E729FFBFCF7DFF17EC8FFE8AF843" bold="true" box="[199,373,1940,1966]" pageId="25" pageNumber="326">Redescription.</emphasis>
Body broad, widest around elytral midlength (
<figureCitation id="133C27BEFFBFCF7DFCAEEC8FFC1BF843" box="[894,996,1940,1966]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="25" pageNumber="326">Fig. 16B</figureCitation>
); surface generally smooth; interspaces between punctures distinctly larger than fine puncture diameter (
<figureCitation id="133C27BEFFBFCF7DFC5AECA3FBE7F83E" box="[906,1048,1976,2003]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="25" pageNumber="326">Fig. 16B, C</figureCitation>
). Color. Body, antennae and legs reddish brown. Lateral margin of pronotum, and external margins of hypomeron and prosternum paler. Lateral margin of elytra yellow. External edge of mandible, lateral and posterior edges of pronotum and mesosternal process between mesocoxae, posterior edges of hypomera and prosternum, external margin of scutellum and anterior edge of elytra blackish (
<figureCitation id="133C27BEFFBCCF7EFEB9EBC4FE0FFF14" box="[361,496,223,249]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16B, C</figureCitation>
). Body covered with long yellow setae.
</paragraph>
<paragraph id="8BB83B3BFFBCCF7EFF17EA18FB43FDD0" blockId="26.[151,1437,151,1977]" pageId="26" pageNumber="327">
Head. Frons flatted medially (
<figureCitation id="133C27BEFFBCCF7EFDC9EA18FD5EFEF0" box="[537,673,259,285]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16D, E</figureCitation>
); frontal carina not complete (
<figureCitation id="133C27BEFFBCCF7EFC28EA18FB9EFEF0" box="[1016,1121,259,285]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16D</figureCitation>
); frontal margin rectangular but broadly rounded apically in dorsal view (
<figureCitation id="133C27BEFFBCCF7EFD70EA3CFCF7FEAC" box="[672,776,295,321]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16E</figureCitation>
); frontoclypeal region protruding beyond base of labrum. Eyes relatively normal in convexity, 0.4 x longer than interocular distance in dorsal view (
<figureCitation id="133C27BEFFBCCF7EFB64EA50FAE3FE88" box="[1204,1308,331,357]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16E</figureCitation>
). Antennae extending beyond pronotum posterior lateral apices by antennomere VI, surpassing elytral half by antennomere XI; antennomeres longer than wide; II obconical, shortest, 1.6 x longer than wide; III weakly serrated, 2.2–2.4 x longer than wide, 1.6 x longer than II; IV–XI filiform; IV 3.0–3.1 x longer than wide, 1.3–1.4 x longer than III, 0.8–0.85 x longer than II–III combined (
<figureCitation id="133C27BEFFBCCF7EFE0BEAC0FDC1FE18" box="[475,574,475,501]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16F</figureCitation>
); V 3.5–4.0 x longer than wide, 1.1–1.2 x longer than IV; XI 6.0–6.3 x longer than wide, 1.1–1.15 x longer than X. Mandible bidentate (
<figureCitation id="133C27BEFFBCCF7EFCF0EAE4FC78FDF4" box="[800,903,511,537]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16D</figureCitation>
). Apical maxillary palpomere semicircular (
<figureCitation id="133C27BEFFBCCF7EFABFEAE4FF34FDD0" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16G</figureCitation>
), 2.3–2.5 x longer than wide, shorter than maximum length of eye; anterior edge rounded.
</paragraph>
<paragraph id="8BB83B3BFFBCCF7EFF17E95CFC51F9C0" blockId="26.[151,1437,151,1977]" pageId="26" pageNumber="327">
Prothorax. Pronotum hexagonal, 0.8 x longer than wide, roundly widening anteriorly, widest around midlength, and then strongly and straightly narrowed anterior to hind angles (
<figureCitation id="133C27BEFFBCCF7EFC47E970FBFFFD68" box="[919,1024,619,645]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16B</figureCitation>
), tallest just behind midlength (
<figureCitation id="133C27BEFFBCCF7EFABFE970FF34FD44" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16H</figureCitation>
), without median longitudinal depression posteriorly; anterior edge weakly concave; anterior angles simple, nearly right angle; punctate lateral ridge extending from anterior angles to hind angles (
<figureCitation id="133C27BEFFBCCF7EFB47E9A8FAFFFD20" box="[1175,1280,691,717]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16B</figureCitation>
); hind angles simple, acute, strongly protruding posterolaterad; posterior edge with a moderate sublateral incision near each hind angle, without carinae next to sublateral incisions (
<figureCitation id="133C27BEFFBCCF7EFD30E9E0FCB7FCF8" box="[736,840,763,789]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16B</figureCitation>
). Hypomeron with moderate mesial projection (
<figureCitation id="133C27BEFFBCCF7EFABFE9E0FF3EFCD4" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16I</figureCitation>
: arrow); anterior angle rounded; mesial edge weakly and broadly rounded; mesial and posterior margins with impunctate ridge (
<figureCitation id="133C27BEFFBCCF7EFEB0E858FE44FCB0" box="[352,443,835,861]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16I</figureCitation>
); posterior margin with triangular projection between two large emarginations; hind angle abruptly narrowed to apex, claw-like shaped. Prosternum weakly to strongly incurved ventrally in lateral view (
<figureCitation id="133C27BEFFBCCF7EFABFE87CFF34FC48" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16H</figureCitation>
); anterior lobe distinctly protruding beyond prosternal ventral line in lateral view (
<figureCitation id="133C27BEFFBCCF7EFB55E890FB0FFC48" box="[1157,1264,907,933]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16H</figureCitation>
); anterior edge broadly rounded in ventral view (
<figureCitation id="133C27BEFFBCCF7EFDDBE8B4FD8EFC24" box="[523,625,943,969]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16C</figureCitation>
). Prosternal process broad, 1.8 x longer than procoxal cavity length (
<figureCitation id="133C27BEFFBCCF7EFABFE8B4FF3EFC00" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16J</figureCitation>
), concave between procoxae, strongly curved dorsad from anterior edge of procoxal cavities in lateral view (
<figureCitation id="133C27BEFFBCCF7EFABFE8C8FF34FBFC" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16H</figureCitation>
), without subapical tooth; dorsal lobe roundly expanded anterior to apex in ventral view (
<figureCitation id="133C27BEFFBCCF7EFB77E8ECFAFBFBFC" box="[1191,1284,1015,1041]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16J</figureCitation>
); ventral lobe strongly roundly expanded near base and then abruptly narrowed posterad in ventral view (
<figureCitation id="133C27BEFFBCCF7EFB58EF00FB1AFBD8" box="[1160,1253,1051,1077]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16J</figureCitation>
); ventral margin weakly and roundly expanded medially in lateral view (
<figureCitation id="133C27BEFFBCCF7EFCDEEF24FC88FBB4" box="[782,887,1087,1113]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16H</figureCitation>
); apex rounded in lateral and ventral views (
<figureCitation id="133C27BEFFBCCF7EFABFEF24FF1BFB90" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16H, J</figureCitation>
). Pronotosternal sutures not grooved (
<figureCitation id="133C27BEFFBCCF7EFD47EF78FD0AFB90" box="[663,757,1123,1149]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16I</figureCitation>
), nearly straight in ventral view (
<figureCitation id="133C27BEFFBCCF7EFBA2EF78FB26FB90" box="[1138,1241,1123,1149]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16C</figureCitation>
), slightly opened anteriorly. Scutellar shield tongue-shaped (
<figureCitation id="133C27BEFFBCCF7EFD58EF9CFD0BFB4C" box="[648,756,1159,1185]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16K</figureCitation>
), 0.95–1.00 x longer than wide, widest anteriorly, weakly constricted anteriorly and narrowed posteriad but parallel-sided in
<typeStatus id="54BC8599FFBCCF7EFC55EFB0FC17FB28" box="[901,1000,1195,1221]" pageId="26" pageNumber="327" type="paratype">paratype</typeStatus>
, flat, inclined anterior-downwards, not visible in lateral view (
<figureCitation id="133C27BEFFBCCF7EFE78EFD4FDEBFB04" box="[424,532,1231,1257]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16H</figureCitation>
); anterior edge broadly rounded but slightly protruding medially in
<typeStatus id="54BC8599FFBCCF7EFAE2EFD4FA67FB04" box="[1330,1432,1231,1257]" pageId="26" pageNumber="327" type="holotype">holotype</typeStatus>
; posterior edge rounded. Mesosternum: borders of mesosternal cavity slightly rounded and then obtusely curved in lateral view (
<figureCitation id="133C27BEFFBCCF7EFEFAEE0CFE6DFADC" box="[298,402,1303,1329]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16H</figureCitation>
); mesosternal process between mesocoxae lower than mesocoxae, not visible in lateral view (
<figureCitation id="133C27BEFFBCCF7EFF4FEE20FEF7FAB8" box="[159,264,1339,1365]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16H</figureCitation>
); posterior edge 0.1–0.15 x wider than total width of mesosternum, weakly to strongly emarginate (
<figureCitation id="133C27BEFFBCCF7EFABFEE20FF37FA94" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16L</figureCitation>
). Mesepisternum reaching mesocoxal cavity (
<figureCitation id="133C27BEFFBCCF7EFD6FEE44FCDDFA94" box="[703,802,1375,1401]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16L</figureCitation>
). Metasternum sulcate medially and anterior to metacoxal cavities (
<figureCitation id="133C27BEFFBCCF7EFF2CEE98FE9DFA70" box="[252,354,1411,1437]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16L</figureCitation>
). Metacoxal plate narrowed toward outer side, becoming like a parallel-sided bar at its outer 1/
<quantity id="4CFF96DEFFBCCF7EFA5EEE98FF52FA2C" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="26" pageNumber="327" unit="in" value="8.0">8 in</quantity>
ventral view (
<figureCitation id="133C27BEFFBCCF7EFE9CEEBCFE4DFA2C" box="[332,434,1447,1473]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16C</figureCitation>
). Elytron broadly strongly convex, but with outer margin widely depressed, widest behind midlength, 3.4–3.6 x longer than wide, 2.9–3.0 x longer than pronotum length; apex rounded; elytral striae defined by lines of elongated punctures. Hind wings fully developed. Tibiae with paired spurs; relative tarsomere lengths: IV&lt;III II&lt;V&lt;I; tarsomeres III and IV with lobe ventrally (
<figureCitation id="133C27BEFFBCCF7EFCE4ED08FC5EF9C0" box="[820,929,1555,1581]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16M</figureCitation>
).
</paragraph>
<paragraph id="8BB83B3BFFBCCF7EFF17ED2CFC1EF854" blockId="26.[151,1437,151,1977]" pageId="26" pageNumber="327">
Abdomen. Ventrite V curved triangular, rounded apically (
<figureCitation id="133C27BEFFBCCF7EFC87ED2CFC41F9BC" box="[855,958,1591,1617]" captionStart="FIGURE 16" captionStartId="25.[151,250,1654,1678]" captionTargetBox="[151,1436,191,1630]" captionTargetId="figure-16@25.[151,1436,181,1630]" captionTargetPageId="25" captionText="FIGURE 16. Penia takasago, holotype, female (PTK01). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of right antenna; G: right maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with moderate mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 16H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 18H); M: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200069" httpUri="https://zenodo.org/record/10200069/files/figure.png" pageId="26" pageNumber="327">Fig. 16C</figureCitation>
), 0.45–0.5 x longer than wide. Tergite and sternite VIII yellow. Terigite VIII curved triangular, rounded apically, 0.8–0.9 x longer than wide (
<figureCitation id="133C27BEFFBCCF7EFB2AED40FA9CF998" box="[1274,1379,1627,1653]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17A</figureCitation>
), 0.7 x longer than wide in
<typeStatus id="54BC8599FFBCCF7EFE42ED64FE0DF974" box="[402,498,1663,1689]" pageId="26" pageNumber="327" type="paratype">paratype</typeStatus>
(6348); sternite VIII (between base of spiculum ventrale and apex) hexagonal, 0.8 x longer than wide, 0.7 x longer than wide in
<typeStatus id="54BC8599FFBCCF7EFD73EDB8FCFCF950" box="[675,771,1699,1725]" pageId="26" pageNumber="327" type="paratype">paratype</typeStatus>
(6348) (
<figureCitation id="133C27BEFFBCCF7EFCB7EDB8FC31F950" box="[871,974,1699,1725]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17B</figureCitation>
; the subgenital segments of the
<typeStatus id="54BC8599FFBCCF7EFAECEDB8FA63F950" box="[1340,1436,1699,1725]" pageId="26" pageNumber="327" type="paratype">paratype</typeStatus>
(6348) had been mounted in balsam on slides and were distorted by the pressure exerted by the coverslip,
<figureCitation id="133C27BEFFBCCF7EFAE4EDDCFF51F8E8" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17C, D</figureCitation>
); spiculum ventrale 3.9–4.6 x longer than length of sternite VIII (
<figureCitation id="133C27BEFFBCCF7EFC4BEDF0FBFFF8E8" box="[923,1024,1771,1797]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17E</figureCitation>
), 3.4 x longer than length of sternite VIII in
<typeStatus id="54BC8599FFBCCF7EFF3DEC14FEB2F8C4" box="[237,333,1807,1833]" pageId="26" pageNumber="327" type="paratype">paratype</typeStatus>
(6348) (
<figureCitation id="133C27BEFFBCCF7EFE60EC14FDE5F8C4" box="[432,538,1807,1833]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17D</figureCitation>
). Ovipositor 0.85–0.9 x longer than length of abdomen; coxites two segmented at ventral side (
<figureCitation id="133C27BEFFBCCF7EFE97EC28FE54F8A0" box="[327,427,1843,1869]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17F</figureCitation>
), with several setae each dorsally, ventrally, and apically; stylus with a seta around apex. Vagina short (
<figureCitation id="133C27BEFFBCCF7EFEE2EC4CFE68F89C" box="[306,407,1879,1905]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17E</figureCitation>
); bursa copulatrix elongated spheroid, posteriorly with a short sac (
<figureCitation id="133C27BEFFBCCF7EFB54EC4CFB13F89C" box="[1156,1260,1879,1905]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17G</figureCitation>
: arrow 1); thick sac extending from around apex of bursa copulatrix (
<figureCitation id="133C27BEFFBCCF7EFD22EC60FCA3F878" box="[754,860,1915,1941]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17G</figureCitation>
: arrow 2), splitting in two (
<figureCitation id="133C27BEFFBCCF7EFB45EC60FB00F878" box="[1173,1279,1915,1941]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17G</figureCitation>
: arrows 3, 4); many sclerotized spines in various sizes inside bursa copulatrix (
<figureCitation id="133C27BEFFBCCF7EFCBCEC84FC2BF854" box="[876,980,1951,1977]" captionStart="FIGURE 17" captionStartId="27.[151,250,1365,1389]" captionTargetBox="[151,1436,181,1341]" captionTargetId="figure-16@27.[151,1436,181,1341]" captionTargetPageId="27" captionText="FIGURE 17. Penia takasago, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix." figureDoi="http://doi.org/10.5281/zenodo.10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="26" pageNumber="327">Fig. 17H</figureCitation>
).
</paragraph>
<caption id="DF786BB3FFBDCF7FFF47EE4EFC13FA34" ID-DOI="http://doi.org/10.5281/zenodo.10200071" ID-Zenodo-Dep="10200071" httpUri="https://zenodo.org/record/10200071/files/figure.png" pageId="27" pageNumber="328" startId="27.[151,250,1365,1389]" targetBox="[151,1436,181,1341]" targetPageId="27" targetType="figure">
<paragraph id="8BB83B3BFFBDCF7FFF47EE4EFC13FA34" blockId="27.[151,1437,1365,1497]" pageId="27" pageNumber="328">
<emphasis id="B973E729FFBDCF7FFF47EE4EFEDFFA80" bold="true" box="[151,288,1365,1389]" pageId="27" pageNumber="328">FIGURE 17.</emphasis>
<taxonomicName id="4C0740B8FFBDCF7FFEF7EE4DFE3CFA80" authorityName="Kishii" authorityYear="1997" box="[295,451,1365,1389]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBDCF7FFEF7EE4DFE3CFA80" box="[295,451,1365,1389]" italics="true" pageId="27" pageNumber="328">Penia takasago</emphasis>
</taxonomicName>
, females. A, B, E–G: holotype (PTK01); C, D: paratype (6348); H: paratype (PTK02). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac; arrow 2: thick sac extending from around apex of bursa copulatrix; arrows 3, 4: thick sac splitting in two); H: sclerotized spines inside bursa copulatrix.
</paragraph>
</caption>
<paragraph id="8BB83B3BFFBDCF7FFF17ED30FE77F9A8" blockId="27.[151,1437,1579,1894]" box="[199,392,1579,1605]" pageId="27" pageNumber="328">
<emphasis id="B973E729FFBDCF7FFF17ED30FEF4F9A8" bold="true" box="[199,267,1579,1605]" pageId="27" pageNumber="328">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFBDCF7FFF17ED54FEA7F8AC" pageId="27" pageNumber="328" type="discussion">
<paragraph id="8BB83B3BFFBDCF7FFF17ED54FD0FF960" blockId="27.[151,1437,1579,1894]" pageId="27" pageNumber="328">
<emphasis id="B973E729FFBDCF7FFF17ED54FEB3F984" bold="true" box="[199,332,1615,1641]" pageId="27" pageNumber="328">Discussion.</emphasis>
A female
<typeStatus id="54BC8599FFBDCF7FFE14ED4BFDDBF987" box="[452,548,1616,1642]" pageId="27" pageNumber="328" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C0740B8FFBDCF7FFD9DED4BFD03F984" authorityName="Kishii" authorityYear="1997" box="[589,764,1615,1641]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBDCF7FFD9DED4BFD03F984" box="[589,764,1615,1641]" italics="true" pageId="27" pageNumber="328">Penia takasago</emphasis>
</taxonomicName>
(6349) does not belong to
<taxonomicName id="4C0740B8FFBDCF7FFBE5ED4BFB45F984" authorityName="Kishii" authorityYear="1997" box="[1077,1210,1615,1642]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBDCF7FFBE5ED4BFBB9F984" box="[1077,1094,1616,1641]" italics="true" pageId="27" pageNumber="328">P</emphasis>
.
<emphasis id="B973E729FFBDCF7FFB85ED54FB45F984" box="[1109,1210,1615,1641]" italics="true" pageId="27" pageNumber="328">takasago</emphasis>
</taxonomicName>
but rather to a new species,
<taxonomicName id="4C0740B8FFBDCF7FFF26ED6FFE7FF960" authorityName="Arimoto" authorityYear="2023" box="[246,384,1652,1678]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="inopinata">
<emphasis id="B973E729FFBDCF7FFF26ED6FFEF8F960" box="[246,263,1652,1677]" italics="true" pageId="27" pageNumber="328">P</emphasis>
.
<emphasis id="B973E729FFBDCF7FFEC5ED6FFE7FF960" box="[277,384,1652,1677]" italics="true" pageId="27" pageNumber="328">inopinata</emphasis>
</taxonomicName>
(see discussion of
<taxonomicName id="4C0740B8FFBDCF7FFD86ED6FFD1FF960" authorityName="Arimoto" authorityYear="2023" box="[598,736,1652,1678]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="inopinata">
<emphasis id="B973E729FFBDCF7FFD86ED6FFD98F960" box="[598,615,1652,1677]" italics="true" pageId="27" pageNumber="328">P</emphasis>
.
<emphasis id="B973E729FFBDCF7FFDA5ED6FFD1FF960" box="[629,736,1652,1677]" italics="true" pageId="27" pageNumber="328">inopinata</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BB83B3BFFBDCF7FFF17ED8CFEA7F8AC" blockId="27.[151,1437,1579,1894]" pageId="27" pageNumber="328">
<bibRefCitation id="EF9646CAFFBDCF7FFF17ED8CFDF4F95C" author="Platia, G. &amp; Schimmel, R." box="[199,523,1687,1713]" pageId="27" pageNumber="328" pagination="49 - 91" refId="ref23269" refString="Platia, G. &amp; Schimmel, R. (2007) Click beetles of Taiwan collected by the expeditions of the Hungarian Natural History Museum in the years 1995 to 2003 (Coleoptera: Elateridae). Annales Historico - Naturales Musei Nationalis Hungarici, 99, 49 - 91." type="journal article" year="2007">Platia and Schimmel (2007)</bibRefCitation>
reported a male of
<taxonomicName id="4C0740B8FFBDCF7FFD27ED83FC57F95C" authorityName="Kishii" authorityYear="1997" box="[759,936,1687,1713]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBDCF7FFD27ED83FC57F95C" box="[759,936,1687,1713]" italics="true" pageId="27" pageNumber="328">Penia takasago</emphasis>
</taxonomicName>
with a short description as the first record of a male but that was based on misidentification because the specimen differs from
<taxonomicName id="4C0740B8FFBDCF7FFBB4EDA7FB15F938" authorityName="Kishii" authorityYear="1997" box="[1124,1258,1723,1750]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBDCF7FFBB4EDA7FB8AF938" box="[1124,1141,1724,1749]" italics="true" pageId="27" pageNumber="328">P</emphasis>
.
<emphasis id="B973E729FFBDCF7FFB55EDA0FB15F938" box="[1157,1258,1723,1749]" italics="true" pageId="27" pageNumber="328">takasago</emphasis>
</taxonomicName>
in the shape of the pronotum and the ratio of the elytron length to pronotum length. Therefore, the male of
<taxonomicName id="4C0740B8FFBDCF7FFB65EDFBFAC4F914" authorityName="Kishii" authorityYear="1997" box="[1205,1339,1759,1786]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFBDCF7FFB65EDFBFB39F914" box="[1205,1222,1760,1785]" italics="true" pageId="27" pageNumber="328">P</emphasis>
.
<emphasis id="B973E729FFBDCF7FFB06EDC4FAC4F914" box="[1238,1339,1759,1785]" italics="true" pageId="27" pageNumber="328">takasago</emphasis>
</taxonomicName>
remains unknown. In this study, the male specimen, which is deposited at the Hungarian Natural History Museum, could not be examined.
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFBDCF7FFF17EC50FD2AF888" box="[199,725,1867,1894]" pageId="27" pageNumber="328" type="distribution">
<paragraph id="8BB83B3BFFBDCF7FFF17EC50FD2AF888" blockId="27.[151,1437,1579,1894]" box="[199,725,1867,1894]" pageId="27" pageNumber="328">
<emphasis id="B973E729FFBDCF7FFF17EC50FE9EF888" bold="true" box="[199,353,1867,1893]" pageId="27" pageNumber="328">Distribution.</emphasis>
<collectingCountry id="F3107BABFFBDCF7FFEB7EC50FE42F888" box="[359,445,1867,1893]" name="Taiwan" pageId="27" pageNumber="328">Taiwan</collectingCountry>
:
<collectingRegion id="49C3F5D9FFBDCF7FFE18EC57FD8CF888" box="[456,627,1867,1894]" country="Taiwan" name="Nantou" pageId="27" pageNumber="328">Nantou County</collectingRegion>
(
<figureCitation id="133C27BEFFBDCF7FFD53EC50FD3BF888" box="[643,708,1867,1893]" captionStart="FIGURE 1" captionStartId="3.[151,250,902,926]" captionTargetBox="[158,1429,188,872]" captionTargetId="figure-16@3.[151,1436,181,879]" captionTargetPageId="3" captionText="FIGURE 1. Map of Taiwan. A: location of Taiwan; B: the collection sites of the material used in this study." figureDoi="http://doi.org/10.5281/zenodo.10200038" httpUri="https://zenodo.org/record/10200038/files/figure.png" pageId="27" pageNumber="328">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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