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<document id="94307DDF6228DC775FE21E6C700DD06B" ID="10.11646/zootaxa.4204.1.1" ID-DOI="10.11646/zootaxa.4204.1.1" ID-GBIF-Dataset="3f8866d2-529e-43ad-b971-29fc52a13858" ID-ISSN="1175-5326" ID-Zenodo-Dep="192302" ID-ZooBank="C050058A-774D-49C0-93F9-7A055B51C2A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1480667017343" checkinUser="plazi" docAuthor="Williams, Paul H., Huang, Jiaxing, Rasmont, Pierre &amp; An, Jiandong" docDate="2016" docId="03AE67547C5B3334B090A5F8A727FDE4" docLanguage="en" docName="zootaxa.4204.1.1.pdf" docOrigin="Zootaxa 4204 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Bombus mendax Gerstaecker" docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="FF971F2C7C6D330CB007A424A336FFA4" masterDocTitle="Early-diverging bumblebees from across the roof of the world: the high-mountain subgenus Mendacibombus revised from species gene coalescents and morphology (Hymenoptera, Apidae)" masterLastPageNumber="72" masterPageNumber="1" pageNumber="55" updateTime="1698395689147" updateUser="plazi">
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<mods:title id="C570DD5470713688A0C5171C7AF52199">Early-diverging bumblebees from across the roof of the world: the high-mountain subgenus Mendacibombus revised from species gene coalescents and morphology (Hymenoptera, Apidae)</mods:title>
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<mods:namePart id="6069DC72C289A13AC1471E3C455D2084">Williams, Paul H.</mods:namePart>
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<mods:namePart id="5341C7698887A1930BC240B194DBC3A8">Huang, Jiaxing</mods:namePart>
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<mods:namePart id="ABE9CAE7B0A5F0C327451D7FA061C791">Rasmont, Pierre</mods:namePart>
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<mods:namePart id="F3B563E91BBB14D2D98E2D16BA4FE310">An, Jiandong</mods:namePart>
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<mods:date id="BD85D5D8D64A10C411F378808F77FD08">2016</mods:date>
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<mods:number id="1A50CA5CEBFB5BE5C8122A74AC957C55">4204</mods:number>
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12.
<taxonomicName id="4C07ADC17C5B333AB0C4A5F8A118FE52" ID-CoL="MFPJ" authority="Gerstaecker" authorityName="Gerstaecker" box="[195,558,476,502]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB0C4A5F8A2BBFE52" bold="true" box="[195,397,476,502]" italics="true" pageId="54" pageNumber="55">Bombus mendax</emphasis>
Gerstaecker
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</emphasis>
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<subSubSection id="C31D85C97C5B333AB090A5DAA074FD73" pageId="54" pageNumber="55" type="description">
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(
<figureCitation id="133CCAC77C5B333AB098A5DAA3C3FDB3" box="[159,245,510,535]" captionStart="FIGURES 24 35" captionStartId="26.[151,264,1867,1889]" captionTargetBox="[233,1387,193,1789]" captionTargetId="figure@26.[233,1390,193,1894]" captionTargetPageId="26" captionText="FIGURES 24 35. Morphology of the male genitalia for species of the subgenus Mendacibombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 24, B. superbus, China MD # 339; 25, B. waltoni, China MD # 216; 26, B. convexus, China MD # 215; 27, B. makarjini, Kazakhstan MD # 1240; 28, B. marussinus, Pakistan MD # 408; 29, B. avinoviellus, India MD # 404; 30, B. himalayanus, India MD # 426; 31, B. turkestanicus, China MD # 351; 32, B. defector, Kyrgyzstan MD # 367; 33, B. margreiteri, China MD # 294; 34, B. handlirschianus, Russia MD # 835; 35, B. mendax, Spain MD # 689." httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="54" pageNumber="55">Figs 35</figureCitation>
,
<figureCitation id="133CCAC77C5B333AB106A5DBA2ABFDB3" box="[257,413,511,535]" captionStart="FIGURES 36 55" captionStartId="29.[151,264,1875,1897]" captionTargetBox="[151,1415,299,1829]" captionTargetId="figure@29.[151,1436,193,1853]" captionTargetPageId="29" captionText="FIGURES 36 55. Morphology of parts of the male genitalia from the dorsal aspect of the left side, anterior to the left of the image, posterior to the right, white bars indicating morphological characters referred to in the key to males: 36, B. waltoni; 37, B. mendax; 38, B. superbus; 39, B. convexus; 40, B. waltoni; 41, B. convexus; 42, B. defector; 43, B. mendax; 44, B. mendax; 45, B. turkestanicus; 46, B. avinoviellus; 47, B. mendax; 48, B. mendax; 49, B. makarjini; 50, B. handlirschianus; 51, B. himalayanus; 52, B. turkestanicus; 53, B. himalayanus; 54, B. himalayanus; 55, B. marussinus." httpUri="https://zenodo.org/record/192314/files/figure.png" pageId="54" pageNumber="55">37, 43, 44, 48</figureCitation>
,
<figureCitation id="133CCAC77C5B333AB1AEA5DAA2F0FDB3" box="[425,454,510,535]" captionStart="FIGURES 56 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="54" pageNumber="55">67</figureCitation>
)
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<emphasis id="B9730A507C5B333AB090A662A3EFFDF8" box="[151,217,582,604]" italics="true" pageId="54" pageNumber="55">Bomb.</emphasis>
&lt;
<emphasis id="B9730A507C5B333AB0E1A66CA3CAFDF9" box="[230,252,584,605]" italics="true" pageId="54" pageNumber="55">us</emphasis>
&gt;
<taxonomicName id="4C07ADC17C5B333AB114A662A17BFDF8" authority="Gerstaecker 1869:323" authorityName="Gerstaecker" authorityPageNumber="323" authorityYear="1869" box="[275,589,582,604]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB114A662A254FDF8" box="[275,354,582,604]" italics="true" pageId="54" pageNumber="55">mendax</emphasis>
<bibRefCitation id="EF96ABB37C5B333AB16DA662A117FDF8" author="Gerstaecker" box="[362,545,582,604]" pageId="54" pageNumber="55" refString="Gerstaecker, A. (1869) Beitrage zur naheren Kenntniss einiger Bienen - Gattungen. Stettiner entomologische Zeitung, 30, 315 - 367." type="journal article" year="1869">Gerstaecker 1869</bibRefCitation>
:323
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, type-locality citation Königsalpen bei Kreuth (4000 Fuss).
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<typeStatus id="54BC68E07C5B333AB4CCA662A600FDF8" box="[1227,1334,582,604]" pageId="54" pageNumber="55" type="lectotype">Lectotype</typeStatus>
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queen by present designation
<collectionCode id="ED164E877C5B333AB196A641A2F3FDDF" box="[401,453,613,635]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="54" pageNumber="55">ZISP</collectionCode>
examined (PR),
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4000 (
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,
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).
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1.
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<treatmentCitationGroup id="AB17F16C7C5B333AB090A6A1A1CAFD1C" pageId="54" pageNumber="55">
&lt;
<taxonomicName id="4C07ADC17C5B333AB0A2A6A0A27AFD3E" ID-CoL="MFPJ" box="[165,332,644,666]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB0A2A6A0A27AFD3E" box="[165,332,644,666]" italics="true" pageId="54" pageNumber="55">Bombus mendax</emphasis>
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&gt;
<emphasis id="B9730A507C5B333AB167A6A0A2D7FD3E" box="[352,481,644,666]" italics="true" pageId="54" pageNumber="55">latofasciatus</emphasis>
<treatmentCitation id="0AA6F0537C5B333AB1EFA6A1A145FD3E" author="Vogt" box="[488,627,644,666]" page="50" pageId="54" pageNumber="55" year="1909">
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:50
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, type-locality citation pyrenaeischen.
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</emphasis>
queen by present designation
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examined,
<location id="8ED880997C5B333AB18BA686A2D7FD1C" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5B333AB18BA686A2D7FD1C" box="[396,481,674,696]" country="France" county="Note" municipality="Pyrenees" name="Gedres'" pageId="54" pageNumber="55">Gèdres</location>
(
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,
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).
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2.
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<treatmentCitationGroup id="AB17F16C7C5B333AB090A6E6A074FD73" box="[151,834,705,727]" pageId="54" pageNumber="55">
[
<taxonomicName id="4C07ADC17C5B333AB098A6E5A270FD73" ID-CoL="MFPJ" box="[159,326,705,727]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB098A6E5A270FD73" box="[159,326,705,727]" italics="true" pageId="54" pageNumber="55">Bombus mendax</emphasis>
</taxonomicName>
ab.
<emphasis id="B9730A507C5B333AB176A6E5A2C0FD73" box="[369,502,705,727]" italics="true" pageId="54" pageNumber="55">atrocaudatus</emphasis>
<treatmentCitation id="0AA6F0537C5B333AB1FBA6E6A1B1FD73" author="Vogt" box="[508,647,705,727]" page="50" pageId="54" pageNumber="55" year="1909">
<bibRefCitation id="EF96ABB37C5B333AB1FBA6E6A151FD73" author="Vogt" box="[508,615,705,727]" pageId="54" pageNumber="55" refString="Vogt, O. (1909) Studien uber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1909, 28 - 84." type="journal article" year="1909">Vogt 1909</bibRefCitation>
:50
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, infrasubspecific.]
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</paragraph>
</subSubSection>
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<treatmentCitationGroup id="AB17F16C7C5B333AB090A6C5A781FD52" box="[151,1207,736,758]" pageId="54" pageNumber="55">
[
<taxonomicName id="4C07ADC17C5B333AB098A6C4A2EFFD52" ID-CoL="5FYXH" box="[159,473,736,758]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="variety" species="mendax" variety="bizonatus">
<emphasis id="B9730A507C5B333AB098A6C4A270FD52" box="[159,326,736,758]" italics="true" pageId="54" pageNumber="55">Bombus mendax</emphasis>
var.
<emphasis id="B9730A507C5B333AB17FA6C4A2EFFD52" box="[376,473,736,758]" italics="true" pageId="54" pageNumber="55">bizonatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C07ADC17C5B333AB1E2A6C5A147FD52" box="[485,625,736,758]" pageId="54" pageNumber="55" rank="variety" variety="friseanus">
var.
<emphasis id="B9730A507C5B333AB216A6C4A147FD52" box="[529,625,736,758]" italics="true" pageId="54" pageNumber="55">friseanus</emphasis>
</taxonomicName>
<emphasis id="B9730A507C5B333AB276A6C4A143FD52" box="[625,629,736,758]" italics="true" pageId="54" pageNumber="55">,</emphasis>
<taxonomicName id="4C07ADC17C5B333AB27CA6C5A0FAFD52" authority="Skorikov 1910b:328" authorityName="Skorikov" authorityPageNumber="328" authorityYear="1910" box="[635,972,736,758]" pageId="54" pageNumber="55" rank="variety" variety="perfuga">
var.
<emphasis id="B9730A507C5B333AB2A1A6C4A1C3FD52" box="[678,757,736,758]" italics="true" pageId="54" pageNumber="55">perfuga</emphasis>
<treatmentCitation id="0AA6F0537C5B333AB2FCA6C4A0FAFD52" author="Skorikov" box="[763,972,736,758]" page="328" pageId="54" pageNumber="55" year="1910">
<bibRefCitation id="EF96ABB37C5B333AB2FCA6C4A097FD52" author="Skorikov" box="[763,929,736,758]" pageId="54" pageNumber="55" refString="Skorikov, A. S. (1910 b) [Intraspecific forms of Bombus mendax Gerst.] (Hymenoptera, Bombidae). Russkoe entomologicheskoe Obozrenie, 9 (1909), 328 - 330." type="journal article" year="1910" yearSuffix="b">Skorikov 1910b</bibRefCitation>
:328
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-329, infrasubspecific.]
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</paragraph>
<paragraph id="8BB8D6427C5B333AB090A6DAA01BFC97" blockId="54.[151,1436,582,1064]" pageId="54" pageNumber="55">
<materialsCitation id="3B6FDC1F7C5B333AB090A6DAA01BFC97" ID-GBIF-Occurrence="1413110431" collectionCode="MNHU" country="Switzerland" location="Alps" municipality="Note" pageId="54" pageNumber="55" specimenCount="1" typeStatus="lectotype">
<treatmentCitationGroup id="AB17F16C7C5B333AB090A6DAA01BFC97" pageId="54" pageNumber="55">
<taxonomicName id="4C07ADC17C5B333AB090A6DAA195FCB0" ID-CoL="5FYXF" authority="Friese 1911b:572" authorityName="Friese" authorityPageNumber="572" authorityYear="1911" box="[151,675,766,789]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="variety" species="mendax" variety="anonymus">
<emphasis id="B9730A507C5B333AB090A6DAA274FCB0" box="[151,322,766,788]" italics="true" pageId="54" pageNumber="55">Bombus mendax</emphasis>
var.
<emphasis id="B9730A507C5B333AB17CA724A2D7FCB1" box="[379,481,768,789]" italics="true" pageId="54" pageNumber="55">anonymus</emphasis>
<treatmentCitation id="0AA6F0537C5B333AB1EBA6DAA195FCB0" author="Friese" box="[492,675,766,788]" page="572" pageId="54" pageNumber="55" year="1911">
<bibRefCitation id="EF96ABB37C5B333AB1EBA6DAA143FCB0" author="Friese" box="[492,629,766,788]" pageId="54" pageNumber="55" refString="Friese, H. (1911 b) Neue Varietaten von Bombus, III. Deutsche Entomologische Zeitschrift, 1911, 571 - 572." type="journal article" year="1911" yearSuffix="b">Friese 1911b</bibRefCitation>
:572
</treatmentCitation>
</taxonomicName>
, type-locality citation Alpen.
<emphasis id="B9730A507C5B333AB3F4A6DAA768FCB0" bold="true" box="[1011,1118,766,788]" pageId="54" pageNumber="55">
<typeStatus id="54BC68E07C5B333AB3F4A6DAA768FCB0" box="[1011,1118,766,788]" pageId="54" pageNumber="55" type="lectotype">Lectotype</typeStatus>
</emphasis>
queen by present designation
<collectionCode id="ED164E877C5B333AB0C0A73AA222FC97" box="[199,276,798,819]" pageId="54" pageNumber="55">MNHU</collectionCode>
examined, Sierre Alp. (
<location id="8ED880997C5B333AB210A739A17CFC97" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5B333AB210A739A17CFC97" box="[535,586,797,819]" country="Switzerland" municipality="Note" name="Alps" pageId="54" pageNumber="55">Alps</location>
,
<collectingCountry id="F31096D27C5B333AB252A739A1E4FC97" box="[597,722,797,819]" name="Switzerland" pageId="54" pageNumber="55">Switzerland</collectingCountry>
).
<collectingMunicipality id="6BDC4C387C5B333AB2E5A73AA025FC97" box="[738,787,798,819]" pageId="54" pageNumber="55">Note</collectingMunicipality>
3.
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
<paragraph id="8BB8D6427C5B333AB090A718A1CCFCF6" blockId="54.[151,1436,582,1064]" box="[151,762,828,850]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="AB17F16C7C5B333AB090A718A1CCFCF6" box="[151,762,828,850]" pageId="54" pageNumber="55">
<taxonomicName id="4C07ADC17C5B333AB090A718A110FCF6" ID-CoL="5JMRJ" authority="Gerstaecker" authorityName="Gerstaecker" box="[151,550,828,850]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB090A718A2A2FCF6" box="[151,404,828,850]" italics="true" pageId="54" pageNumber="55">Mendacibombus mendax</emphasis>
(Gerstaecker)
</taxonomicName>
;
<treatmentCitation id="0AA6F0537C5B333AB237A718A1C0FCF6" author="Skorikov" box="[560,758,828,850]" page="149" pageId="54" pageNumber="55" year="1923">
<bibRefCitation id="EF96ABB37C5B333AB237A718A1FCFCF6" author="Skorikov" box="[560,714,828,850]" pageId="54" pageNumber="55" refString="Skorikov, A. S. (1923) [Palaearctic bumblebees. Part I. General biology (including zoogeography)]. Izvestiya Severnoi oblastnoi stantsii zashchity rastenii ot vreditelei, 4 (1922), 1 - 160." type="journal article" year="1923">Skorikov 1923</bibRefCitation>
:149
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31D85C97C5B333AB090A77FA7CDFC0A" pageId="54" pageNumber="55" type="description">
<paragraph id="8BB8D6427C5B333AB090A77FA24EFC2B" blockId="54.[151,1436,582,1064]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="AB17F16C7C5B333AB090A77FA24EFC2B" pageId="54" pageNumber="55">
[
<taxonomicName id="4C07ADC17C5B333AB098A77EA271FCD4" ID-CoL="MFPJ" box="[159,327,858,880]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB098A77EA271FCD4" box="[159,327,858,880]" italics="true" pageId="54" pageNumber="55">Bombus mendax</emphasis>
</taxonomicName>
morpha. nov.
<emphasis id="B9730A507C5B333AB1D9A77EA169FCD4" box="[478,607,858,880]" italics="true" pageId="54" pageNumber="55">subglacialis,</emphasis>
ab. nov.
<emphasis id="B9730A507C5B333AB2B8A77EA034FCD4" box="[703,770,858,880]" italics="true" pageId="54" pageNumber="55">flavior</emphasis>
, ab. nov.
<emphasis id="B9730A507C5B333AB36EA77EA0E1FCD4" box="[873,983,858,880]" italics="true" pageId="54" pageNumber="55">feretypicus</emphasis>
, ab. nov.
<emphasis id="B9730A507C5B333AB439A77EA794FCD4" box="[1086,1186,858,880]" italics="true" pageId="54" pageNumber="55">basizonus</emphasis>
<treatmentCitation id="0AA6F0537C5B333AB4ADA77EA65EFCD4" author="Pittioni" box="[1194,1384,858,880]" page="121" pageId="54" pageNumber="55" year="1937">
<bibRefCitation id="EF96ABB37C5B333AB4ADA77EA608FCD4" author="Pittioni" box="[1194,1342,858,880]" pageId="54" pageNumber="55" refString="Pittioni, B. (1937 a) Die Hummelfauna des Kalsbachtales in Ost-Tirol. Ein Beitrag zur Okologie und Systematik der Hummeln Mitteleuropas. Festschrift zum 60. Geburtstag von Professor Dr. Embrik Strand, 3, 64 - 122." type="journal article" year="1937" yearSuffix="a">Pittioni 1937a</bibRefCitation>
:121
</treatmentCitation>
-122, infrasubspecific.]
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB8D6427C5B333AB090A7BDA7CDFC0A" blockId="54.[151,1436,582,1064]" box="[151,1275,920,942]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="AB17F16C7C5B333AB090A7BDA7CDFC0A" box="[151,1275,920,942]" pageId="54" pageNumber="55">
[
<taxonomicName id="4C07ADC17C5B333AB098A7BCA132FC0A" ID-CoL="MFPJ" box="[159,516,920,942]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax" subGenus="Mendacibombus">
<emphasis id="B9730A507C5B333AB098A7BCA132FC0A" box="[159,516,920,942]" italics="true" pageId="54" pageNumber="55">Bombus (Mendacibombus) mendax</emphasis>
</taxonomicName>
f.
<emphasis id="B9730A507C5B333AB227A7BCA19AFC0A" box="[544,684,920,942]" italics="true" pageId="54" pageNumber="55">flavopleuralis</emphasis>
, f.
<emphasis id="B9730A507C5B333AB2C9A7BCA0B0FC0A" box="[718,902,920,942]" italics="true" pageId="54" pageNumber="55">quasilatofasciatus</emphasis>
<treatmentCitation id="0AA6F0537C5B333AB38BA7BCA709FC0A" author="Pittioni" box="[908,1087,920,942]" page="103" pageId="54" pageNumber="55" year="1939">
<bibRefCitation id="EF96ABB37C5B333AB38BA7BCA722FC0A" author="Pittioni" box="[908,1044,920,942]" pageId="54" pageNumber="55" refString="Pittioni, B. (1939) Die Hummeln und Schmarotzerhummeln der Balkan-Halbinsel. II. Spezieller Teil. Izvestiya na Tsarskite prirodonauchni instituti v Sofiya, 12, 49 - 115." type="journal article" year="1939">Pittioni 1939</bibRefCitation>
:103
</treatmentCitation>
, infrasubspecific.]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31D85C97C5B333AB090A793A06CFBAE" pageId="54" pageNumber="55" type="reference_group">
<paragraph id="8BB8D6427C5B333AB090A793A224FC4F" blockId="54.[151,1436,582,1064]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="AB17F16C7C5B333AB090A793A224FC4F" pageId="54" pageNumber="55">
&lt;
<taxonomicName id="4C07ADC17C5B333AB0A2A792A295FC68" ID-CoL="5JMRJ" box="[165,419,950,972]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB0A2A792A295FC68" box="[165,419,950,972]" italics="true" pageId="54" pageNumber="55">Mendacibombus mendax</emphasis>
</taxonomicName>
&gt; ssp.
<emphasis id="B9730A507C5B333AB1ECA79CA10FFC69" box="[491,569,952,973]" italics="true" pageId="54" pageNumber="55">pyrenes</emphasis>
<treatmentCitation id="0AA6F0537C5B333AB245A792A1C5FC68" author="Tkalcu" box="[578,755,950,972]" page="173" pageId="54" pageNumber="55" year="1975">
<bibRefCitation id="EF96ABB37C5B333AB245A792A1FEFC68" author="Tkalcu" box="[578,712,950,972]" pageId="54" pageNumber="55" refString="Tkalcu, B. (1975) Beitrag zur Kenntnis der Hummelfauna der franzosischen Basses - Alpes (Hymenoptera, Apoidea, Bombinae). Sbornik Slovenskeho Narodneho Muzea, 20 (1974), 167 - 186." type="journal article" year="1975">Tkalců 1975</bibRefCitation>
:173
</treatmentCitation>
, unnecessary replacement name for
<emphasis id="B9730A507C5B333AB468A792A7C7FC68" box="[1135,1265,950,972]" italics="true" pageId="54" pageNumber="55">latofasciatus</emphasis>
<bibRefCitation id="EF96ABB37C5B333AB4FEA793A6AFFC68" author="Vogt" box="[1273,1433,950,972]" pageId="54" pageNumber="55" refString="Vogt, O. (1909) Studien uber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1909, 28 - 84." type="journal article" year="1909">Vogt (1909:50)</bibRefCitation>
. Note 4.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB8D6427C5B333AB090A7D0A06CFBAE" blockId="54.[151,1436,582,1064]" box="[151,858,1012,1034]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="AB17F16C7C5B333AB090A7D0A06CFBAE" box="[151,858,1012,1034]" pageId="54" pageNumber="55">
<taxonomicName id="4C07ADC17C5B333AB090A7D0A1A0FBAE" authority="Gerstaecker" authorityName="Gerstaecker" box="[151,662,1012,1034]" class="Magnoliopsida" family="Scrophulariaceae" genus="Bombias" kingdom="Plantae" order="Lamiales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="mendax" subGenus="Mendacibombus">
<emphasis id="B9730A507C5B333AB090A7D0A135FBAE" box="[151,515,1012,1034]" italics="true" pageId="54" pageNumber="55">Bombias (Mendacibombus) mendax</emphasis>
(Gerstaecker)
</taxonomicName>
;
<treatmentCitation id="0AA6F0537C5B333AB2A7A7D1A061FBAE" author="Rasmont" box="[672,855,1012,1034]" page="10" pageId="54" pageNumber="55" year="1983">
<bibRefCitation id="EF96ABB37C5B333AB2A7A7D1A00FFBAE" author="Rasmont" box="[672,825,1012,1034]" pageId="54" pageNumber="55" refString="Rasmont, P. (1983) Catalogue commente des bourdons de la region ouest - palearctique (Hymenoptera, Apoidea, Apidae). Notes Fauniques de Gembloux, 7, 1 - 71." type="journal article" year="1983">Rasmont 1983</bibRefCitation>
:10
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31D85C97C5B333BB090A036A7BFFDBC" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="description">
<paragraph id="8BB8D6427C5B333AB090A036A043FB8C" blockId="54.[151,1436,582,1064]" box="[151,885,1042,1064]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="AB17F16C7C5B333AB090A036A043FB8C" box="[151,885,1042,1064]" pageId="54" pageNumber="55">
<taxonomicName id="4C07ADC17C5B333AB090A036A148FB8C" ID-CoL="MFPJ" authority="Gerstaecker" authorityName="Gerstaecker" box="[151,638,1042,1064]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax" subGenus="Mendacibombus">
<emphasis id="B9730A507C5B333AB090A036A2CAFB8C" box="[151,508,1042,1064]" italics="true" pageId="54" pageNumber="55">Bombus (Mendacibombus) mendax</emphasis>
Gerstaecker
</taxonomicName>
;
<treatmentCitation id="0AA6F0537C5B333AB28FA037A047FB8C" author="Williams" box="[648,881,1042,1064]" page="99" pageId="54" pageNumber="55" year="1998">
<bibRefCitation id="EF96ABB37C5B333AB28FA037A064FB8C" author="Williams" box="[648,850,1042,1064]" pageId="54" pageNumber="55" refString="Williams, P. H. (1998) An annotated checklist of bumble bees with an analysis of patterns of description (Hymenoptera: Apidae, Bombini). Bulletin of The Natural History Museum (Entomology), 67, 79 - 152. [updated at www. nhm. ac. uk / bombus / accessed 2015]." type="journal article" year="1998">P.H. Williams 1998</bibRefCitation>
:99
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB8D6427C5B333AB090A072A104FAE1" blockId="54.[151,1436,1109,2034]" pageId="54" pageNumber="55">
Note 1 (
<taxonomicName id="4C07ADC17C5B333AB0F5A071A27FFBCA" box="[242,329,1109,1134]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB0F5A071A27FFBCA" box="[242,329,1109,1134]" italics="true" pageId="54" pageNumber="55">mendax</emphasis>
</taxonomicName>
). Gerstaeckers original description of the taxon
<taxonomicName id="4C07ADC17C5B333AB36BA071A0F5FBCA" box="[876,963,1109,1134]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB36BA071A0F5FBCA" box="[876,963,1109,1134]" italics="true" pageId="54" pageNumber="55">mendax</emphasis>
</taxonomicName>
cites the
<typeStatus id="54BC68E07C5B333AB42AA072A76BFBCA" box="[1069,1117,1110,1134]" pageId="54" pageNumber="55">type</typeStatus>
locality as Königsalpen bei Kreuth (4000 Fuss). The ZISP collection contains material studied by Gerstaecker and includes a queen that agrees with the original description and carries the labels: (1) white, handwritten Kreuth 4000 ; (2) white, printed Osnovnaya kollektsiya; (3) white, handwritten
<taxonomicName id="4C07ADC17C5B333AB2B9A0E5A09EFB7E" authority="Gerst. Typ." authorityName="Gerst. Typ." box="[702,936,1217,1242]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">mendax Gerst. Typ.</taxonomicName>
; (4) white with a black border, hand-written
<taxonomicName id="4C07ADC17C5B333AB09CA0C1A241FB5A" authority="Gerstack." authorityName="Gerstack." box="[155,375,1253,1278]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">mendax Gerstäck.</taxonomicName>
; (5) red, handwritten Type. No labels were added for this project. This queen, which is complete, is regarded as a
<typeStatus id="54BC68E07C5B333AB1C8A12EA110FA86" box="[463,550,1290,1314]" pageId="54" pageNumber="55" type="syntype">syntype</typeStatus>
and is designated here as the
<typeStatus id="54BC68E07C5B333AB38EA12DA0D9FA86" box="[905,1007,1289,1314]" pageId="54" pageNumber="55" type="lectotype">lectotype</typeStatus>
in order to reduce uncertainty in the identity and application of the name.
</paragraph>
<paragraph id="8BB8D6427C5B333AB0C0A176A709F976" blockId="54.[151,1436,1109,2034]" pageId="54" pageNumber="55">
Note 2 (
<emphasis id="B9730A507C5B333AB122A175A282FACE" box="[293,436,1361,1386]" italics="true" pageId="54" pageNumber="55">latofasciatus</emphasis>
). The name
<emphasis id="B9730A507C5B333AB241A175A1E3FACE" box="[582,725,1361,1386]" italics="true" pageId="54" pageNumber="55">latofasciatus</emphasis>
was used by
<bibRefCitation id="EF96ABB37C5B333AB376A176A7F5FACE" author="Vogt" box="[881,1219,1361,1386]" pageId="54" pageNumber="55" refString="Vogt, O. (1909) Studien uber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1909, 28 - 84." type="journal article" year="1909">Vogt (1909: pages 42, 49, 50)</bibRefCitation>
for colour-patternbased taxa within four different species (
<emphasis id="B9730A507C5B333AB25BA152A0E1FA2A" box="[604,983,1397,1422]" italics="true" pageId="54" pageNumber="55">
<taxonomicName id="4C07ADC17C5B333AB25BA152A1ECFA2A" box="[604,730,1397,1422]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="lucorum">B. lucorum</taxonomicName>
,
<taxonomicName id="4C07ADC17C5B333AB2E3A152A06BFA2A" box="[740,861,1397,1422]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">B. mendax</taxonomicName>
,
<taxonomicName id="4C07ADC17C5B333AB360A152A0E1FA2A" box="[871,983,1397,1422]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="sichelii">B. sichelii</taxonomicName>
</emphasis>
[as
<taxonomicName id="4C07ADC17C5B333AB403A152A75AFA2A" box="[1028,1132,1397,1422]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="sicheli">
<emphasis id="B9730A507C5B333AB403A152A75AFA2A" box="[1028,1132,1397,1422]" italics="true" pageId="54" pageNumber="55">B. sicheli</emphasis>
</taxonomicName>
], and
<taxonomicName id="4C07ADC17C5B333AB4B5A152A600FA29" box="[1202,1334,1398,1421]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="terrestris">
<emphasis id="B9730A507C5B333AB4B5A152A600FA29" box="[1202,1334,1398,1421]" italics="true" pageId="54" pageNumber="55">B. terrestris</emphasis>
</taxonomicName>
). For the names
<emphasis id="B9730A507C5B333AB0EFA1BDA241FA16" box="[232,375,1433,1458]" italics="true" pageId="54" pageNumber="55">latofasciatus</emphasis>
that were used for parts of
<taxonomicName id="4C07ADC17C5B333AB2BAA1BEA00AFA16" box="[701,828,1433,1458]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="lucorum">
<emphasis id="B9730A507C5B333AB2BAA1BEA00AFA16" box="[701,828,1433,1458]" italics="true" pageId="54" pageNumber="55">B. lucorum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C07ADC17C5B333AB37FA1BEA772FA16" authority="Vogt" authorityName="Vogt" box="[888,1092,1434,1458]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="terrestris">
<emphasis id="B9730A507C5B333AB37FA1BEA0C9FA15" box="[888,1023,1434,1457]" italics="true" pageId="54" pageNumber="55">B. terrestris</emphasis>
, Vogt
</taxonomicName>
explicitly referred to his new names using the term ab., so these taxa must remain permanently of infrasubspecific rank (
<bibRefCitation id="EF96ABB37C5B333AB4BDA19AA3D4FA5E" author="ICZN" pageId="54" pageNumber="55" refString="ICZN (1999) International code of zoological nomenclature (4 ed.). London, International Commission on Zoological Nomenclature." type="book chapter" year="1999">ICZN 1999: Article 45.6.2</bibRefCitation>
). In the case of the name
<emphasis id="B9730A507C5B333AB20AA1C5A1AAFA5E" box="[525,668,1505,1530]" italics="true" pageId="54" pageNumber="55">latofasciatus</emphasis>
that was used for a part of
<taxonomicName id="4C07ADC17C5B333AB3D8A1C6A767FA5E" box="[991,1105,1505,1530]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="sichelii">
<emphasis id="B9730A507C5B333AB3D8A1C6A767FA5E" box="[991,1105,1505,1530]" italics="true" pageId="54" pageNumber="55">B. sichelii</emphasis>
</taxonomicName>
, he related it to one of three subsets (Grade) within the part of the population with yellow bands known from the Alps and Pyrenees (referred to by him as [subspecies]
<emphasis id="B9730A507C5B333AB1B2A20DA13CF9E6" box="[437,522,1577,1602]" italics="true" pageId="54" pageNumber="55">alticola</emphasis>
), thereby implying infrasubspecific rank for this
<emphasis id="B9730A507C5B333AB429A20DA78BF9E6" box="[1070,1213,1577,1602]" italics="true" pageId="54" pageNumber="55">latofasciatus</emphasis>
as well. In contrast, Vogt gave no explicit rank to the taxon
<emphasis id="B9730A507C5B333AB287A269A026F9C2" box="[640,784,1613,1638]" italics="true" pageId="54" pageNumber="55">latofasciatus</emphasis>
of
<emphasis id="B9730A507C5B333AB346A26AA0F2F9C2" box="[833,964,1613,1638]" italics="true" pageId="54" pageNumber="55">
<taxonomicName id="4C07ADC17C5B333AB346A26AA0F6F9C2" box="[833,960,1613,1638]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">B. mendax</taxonomicName>
,
</emphasis>
but did relate it to Die pyrenaeischen Exemplare, contrasting them in colour pattern with specimens from the Alps, so this characterisation by geographic distribution and colour-pattern (without a higher subspecific taxon) should be considered to imply subspecific rank as the original intention of the author (ICZN 1 999: Article 45.6.1).
</paragraph>
<paragraph id="8BB8D6427C5B333AB0C0A2FAA0FCF855" blockId="54.[151,1436,1109,2034]" pageId="54" pageNumber="55">
<materialsCitation id="3B6FDC1F7C5B333AB0C0A2FAA635F80E" ID-GBIF-Occurrence="1413110425" collectionCode="RMNH, ZMAN, HYME" collectorName="C. et O. Vogt &amp; Vogt" county="Vogt" determinerName="PH Williams" location="Vogt's" municipality="Collectie" pageId="54" pageNumber="55" specimenCount="3" typeStatus="lectotype">
<location id="8ED880997C5B333AB0C0A2FAA238F952" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5B333AB0C0A2FAA238F952" box="[199,270,1758,1782]" county="Vogt" municipality="Collectie" name="Vogt's" pageId="54" pageNumber="55">Vogts</location>
original description of the taxon
<emphasis id="B9730A507C5B333AB293A2F9A012F952" box="[660,804,1757,1782]" italics="true" pageId="54" pageNumber="55">latofasciatus</emphasis>
cites the
<typeStatus id="54BC68E07C5B333AB391A2FAA0F0F952" box="[918,966,1758,1782]" pageId="54" pageNumber="55">type</typeStatus>
locality as pyrenaeischen.
<location id="8ED880997C5B333AB515A2F9A608F952" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5B333AB515A2F9A608F952" box="[1298,1342,1757,1782]" county="Vogt" municipality="Collectie" name="The" pageId="54" pageNumber="55">The</location>
<collectionCode id="ED164E877C5B333AB54FA2FAA6AAF951" box="[1352,1436,1758,1781]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="54" pageNumber="55">RMNH</collectionCode>
collection studied by
<location id="8ED880997C5B333AB181A326A28CF8BE" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5B333AB181A326A28CF8BE" box="[390,442,1794,1818]" county="Vogt" municipality="Collectie" name="Vogt" pageId="54" pageNumber="55">Vogt</location>
contains a queen that agrees with the original description and carries the labels (1) green, printed in red
<location id="8ED880997C5B333AB142A301A290F89A" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5B333AB142A301A290F89A" box="[325,422,1829,1854]" county="Vogt" municipality="Collectie" name="Gedres'" pageId="54" pageNumber="55">Gèdres</location>
; (2) white, printed
<collectingMunicipality id="6BDC4C387C5B333AB288A301A1CFF89A" box="[655,761,1829,1854]" pageId="54" pageNumber="55">Collectie</collectingMunicipality>
/
<collectorName id="26F2B3947C5B333AB310A302A083F89A" box="[791,949,1830,1854]" pageId="54" pageNumber="55">C. et O. Vogt</collectorName>
/ Acq. 1960; (3) red, printed
<taxonomicName id="4C07ADC17C5B333AB520A301A6BFF89A" box="[1319,1417,1829,1854]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Bombus</taxonomicName>
/
<taxonomicName id="4C07ADC17C5B333AB090A36DA3C6F8C6" box="[151,240,1865,1890]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">mendax</taxonomicName>
latofasciatus /
<bibRefCitation id="EF96ABB37C5B333AB19CA36EA126F8C6" author="Vogt" box="[411,528,1866,1890]" pageId="54" pageNumber="55" refString="Vogt, O. (1909) Studien uber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1909, 28 - 84." type="journal article" year="1909">
<collectingCounty id="62D9AECE7C5B333AB19CA36EA2F9F8C6" box="[411,463,1866,1890]" pageId="54" pageNumber="55">Vogt</collectingCounty>
1909
</bibRefCitation>
/
<collectionCode id="ED164E877C5B333AB220A36EA14FF8C6" box="[551,633,1866,1890]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="54" pageNumber="55">ZMAN</collectionCode>
<typeStatus id="54BC68E07C5B333AB285A36EA184F8C6" box="[642,690,1866,1890]" pageId="54" pageNumber="55">type</typeStatus>
<collectionCode id="ED164E877C5B333AB2BDA36DA025F8C6" box="[698,787,1865,1890]" pageId="54" pageNumber="55">HYME</collectionCode>
.0146.9; (4) green, printed
<taxonomicName id="4C07ADC17C5B333AB457A36DA63FF8C6" box="[1104,1289,1865,1890]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B9730A507C5B333AB457A36DA63FF8C6" box="[1104,1289,1865,1890]" italics="true" pageId="54" pageNumber="55">Mendacibombus</emphasis>
</taxonomicName>
/ MD# 1239 det. PHW; (5) red, printed
<typeStatus id="54BC68E07C5B333AB1EEA34AA1B8F822" box="[489,654,1902,1926]" pageId="54" pageNumber="55" type="lectotype">LECTOTYPE</typeStatus>
[female] /
<taxonomicName id="4C07ADC17C5B333AB31EA349A0E0F822" box="[793,982,1901,1926]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB31EA349A0E0F822" box="[793,982,1901,1926]" italics="true" pageId="54" pageNumber="55">Bombus mendax</emphasis>
</taxonomicName>
/
<emphasis id="B9730A507C5B333AB3F2A349A7B2F822" box="[1013,1156,1901,1926]" italics="true" pageId="54" pageNumber="55">latofasciatus</emphasis>
/
<bibRefCitation id="EF96ABB37C5B333AB4A3A34AA612F822" author="Vogt" box="[1188,1316,1902,1926]" pageId="54" pageNumber="55" refString="Vogt, O. (1909) Studien uber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1909, 28 - 84." type="journal article" year="1909">
<collectorName id="26F2B3947C5B333AB4A3A34AA7EAF822" box="[1188,1244,1902,1926]" pageId="54" pageNumber="55">Vogt</collectorName>
, 1909
</bibRefCitation>
/ det.
<determinerName id="7E643A867C5B333AB57FA34AA3CBF80E" pageId="54" pageNumber="55">PH Williams</determinerName>
2012; (6) white, printed [female]
<taxonomicName id="4C07ADC17C5B333AB28EA3B5A1D5F80E" box="[649,739,1937,1962]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B9730A507C5B333AB28EA3B5A1D5F80E" box="[649,739,1937,1962]" italics="true" pageId="54" pageNumber="55">Bombus</emphasis>
</taxonomicName>
/ (
<taxonomicName id="4C07ADC17C5B333AB307A3B5A08FF80E" box="[768,953,1937,1962]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B9730A507C5B333AB307A3B5A08FF80E" box="[768,953,1937,1962]" italics="true" pageId="54" pageNumber="55">Mendacibombus</emphasis>
</taxonomicName>
) /
<taxonomicName id="4C07ADC17C5B333AB3D0A3B5A718F80E" box="[983,1070,1937,1962]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5B333AB3D0A3B5A718F80E" box="[983,1070,1937,1962]" italics="true" pageId="54" pageNumber="55">mendax</emphasis>
</taxonomicName>
/ Gerstaecker / det
</materialsCitation>
.
<materialsCitation id="3B6FDC1F7C5B333AB50BA3B6A0FCF855" ID-GBIF-Occurrence="1413110409" country="Philippines" location="This" municipality="Vogt's" pageId="54" pageNumber="55" specimenCount="2" typeStatus="syntype">
<collectingCountry id="F31096D27C5B333AB50BA3B6A606F80D" box="[1292,1328,1938,1961]" name="Philippines" pageId="54" pageNumber="55">PH</collectingCountry>
Williams 2012.
<location id="8ED880997C5B333AB0E1A391A22EF86A" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5B333AB0E1A391A22EF86A" box="[230,280,1973,1998]" country="Philippines" municipality="Vogt's" name="This" pageId="54" pageNumber="55">This</location>
queen, which is complete, is regarded as one of
<collectingMunicipality id="6BDC4C387C5B333AB33FA392A049F86A" box="[824,895,1974,1998]" pageId="54" pageNumber="55">Vogts</collectingMunicipality>
<typeStatus id="54BC68E07C5B333AB381A392A0DEF86A" box="[902,1000,1974,1998]" pageId="54" pageNumber="55" type="syntype">syntypes</typeStatus>
and is designated here as the
<typeStatus id="54BC68E07C5B333AB532A391A6AAF86A" box="[1333,1436,1973,1998]" pageId="54" pageNumber="55" type="lectotype">lectotype</typeStatus>
in order to reduce uncertainty in the identity and application of the name.
</materialsCitation>
</paragraph>
<paragraph id="8BB8D6427C5A333BB0C0A4BCA104FE6B" blockId="55.[151,1437,151,2034]" pageId="55" pageNumber="56">
<materialsCitation id="3B6FDC1F7C5A333BB0C0A4BCA013FE2C" ID-GBIF-Occurrence="1413110411" collectingDate="1984-06-19" collectionCode="MNHU" collectorName="Friese &amp; Gerstaecker" county="Berlin'" determinerName="PH Williams" location="Note" municipality="Type'" pageId="55" pageNumber="56" specimenCount="2" stateProvince="Mus" typeStatus="lectotype">
<location id="8ED880997C5A333BB0C0A4BCA3CBFF0B" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5A333BB0C0A4BCA3CBFF0B" box="[199,253,152,175]" county="Berlin'" municipality="Type'" name="Note" pageId="55" pageNumber="56" stateProvince="Mus">Note</location>
3 (
<taxonomicName id="4C07ADC17C5A333BB124A4BDA2A3FF14" box="[291,405,153,176]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="variety" species="mendax" variety="anonymus">
<emphasis id="B9730A507C5A333BB124A4BDA2A3FF14" box="[291,405,153,176]" italics="true" pageId="55" pageNumber="56">anonymus</emphasis>
</taxonomicName>
).
<location id="8ED880997C5A333BB1ABA4B3A135FF14" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5A333BB1ABA4B3A135FF14" box="[428,515,151,176]" county="Berlin'" municipality="Type'" name="Friese's" pageId="55" pageNumber="56" stateProvince="Mus">Frieses</location>
original description of the taxon
<taxonomicName id="4C07ADC17C5A333BB37CA4BDA0DBFF14" box="[891,1005,153,176]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="variety" species="mendax" variety="anonymus">
<emphasis id="B9730A507C5A333BB37CA4BDA0DBFF14" box="[891,1005,153,176]" italics="true" pageId="55" pageNumber="56">anonymus</emphasis>
</taxonomicName>
cites the
<typeStatus id="54BC68E07C5A333BB45FA4BCA7BEFF14" box="[1112,1160,152,176]" pageId="55" pageNumber="56">type</typeStatus>
locality as
<location id="8ED880997C5A333BB509A4B3A657FF14" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5A333BB509A4B3A657FF14" box="[1294,1377,151,176]" county="Berlin'" municipality="Type'" name="Alpen'" pageId="55" pageNumber="56" stateProvince="Mus">Alpen</location>
for a queen, worker, and male.
<location id="8ED880997C5A333BB1B8A498A2DDFF71" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5A333BB1B8A498A2DDFF71" box="[447,491,188,213]" county="Berlin'" municipality="Type'" name="The" pageId="55" pageNumber="56" stateProvince="Mus">The</location>
<collectionCode id="ED164E877C5A333BB1F3A499A17FFF70" box="[500,585,189,212]" pageId="55" pageNumber="56">MNHU</collectionCode>
collection studied by
<location id="8ED880997C5A333BB34FA498A0BAFF71" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5A333BB34FA498A0BAFF71" box="[840,908,188,213]" county="Berlin'" municipality="Type'" name="Friese" pageId="55" pageNumber="56" stateProvince="Mus">Friese</location>
contains a queen that agrees with the original description and carries the labels (1) white, printed Sierre Alp. /
<collectingDate id="EFFD096A7C5A333BB394A4C4A0CDFF5C" box="[915,1019,223,248]" pageId="55" pageNumber="56" value="1984-06-19">
<date id="FFB9F0827C5A333BB394A4C4A0CDFF5C" box="[915,1019,223,248]" pageId="55" pageNumber="56" value="1984-06-19">19. 6. 84</date>
</collectingDate>
/
<location id="8ED880997C5A333BB411A4FBA751FF5C" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5A333BB411A4FBA751FF5C" box="[1046,1127,223,248]" county="Berlin'" municipality="Type'" name="Friese'" pageId="55" pageNumber="56" stateProvince="Mus">Friese</location>
; (2) white, handwritten by
<location id="8ED880997C5A333BB090A520A3EDFEB9" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5A333BB090A520A3EDFEB9" box="[151,219,260,285]" county="Berlin'" municipality="Type'" name="Friese" pageId="55" pageNumber="56" stateProvince="Mus">Friese</location>
<emphasis id="B9730A507C5A333BB0EBA521A25DFEB9" box="[236,363,260,285]" italics="true" pageId="55" pageNumber="56">
<taxonomicName id="4C07ADC17C5A333BB0EBA521A251FEB9" box="[236,359,260,285]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mendax">B. mendax</taxonomicName>
.
</emphasis>
/
<emphasis id="B9730A507C5A333BB185A521A2A2FEB8" box="[386,404,261,284]" italics="true" pageId="55" pageNumber="56">v.</emphasis>
/
<taxonomicName id="4C07ADC17C5A333BB1ACA521A12BFEB8" box="[427,541,261,284]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="variety" species="mendax" variety="anonymus">
<emphasis id="B9730A507C5A333BB1ACA521A12BFEB8" box="[427,541,261,284]" italics="true" pageId="55" pageNumber="56">anonymus</emphasis>
</taxonomicName>
/ [female] 1909 Friese
<emphasis id="B9730A507C5A333BB320A521A074FEB8" box="[807,834,261,284]" italics="true" pageId="55" pageNumber="56">Fr</emphasis>
. Det.; (3) maroon, printed
<collectingMunicipality id="6BDC4C387C5A333BB485A520A7FDFEB8" box="[1154,1227,260,284]" pageId="55" pageNumber="56">Type</collectingMunicipality>
; (4) white, printed Zool.
<collectingRegion id="49C318A07C5A333BB0E3A50CA22FFE9B" box="[228,281,296,319]" country="Turkey" name="Mus" pageId="55" pageNumber="56">Mus</collectingRegion>
. /
<collectingCounty id="62D9AECE7C5A333BB134A503A2B0FEE4" box="[307,390,295,320]" pageId="55" pageNumber="56">Berlin</collectingCounty>
; (5) green, printed
<taxonomicName id="4C07ADC17C5A333BB260A503A016FEE4" box="[615,800,295,320]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B9730A507C5A333BB260A503A016FEE4" box="[615,800,295,320]" italics="true" pageId="55" pageNumber="56">Mendacibombus</emphasis>
</taxonomicName>
/ MD# 3540 det. PHW; (6) red, printed
<typeStatus id="54BC68E07C5A333BB4F0A50CA6AAFEE4" box="[1271,1436,296,320]" pageId="55" pageNumber="56" type="lectotype">LECTOTYPE</typeStatus>
[female] /
<taxonomicName id="4C07ADC17C5A333BB108A568A2FFFEC1" box="[271,457,332,357]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5A333BB108A568A2FFFEC1" box="[271,457,332,357]" italics="true" pageId="55" pageNumber="56">Bombus mendax</emphasis>
</taxonomicName>
var. /
<taxonomicName id="4C07ADC17C5A333BB212A569A1B1FEC0" box="[533,647,333,356]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="variety" species="mendax" variety="anonymus">
<emphasis id="B9730A507C5A333BB212A569A1B1FEC0" box="[533,647,333,356]" italics="true" pageId="55" pageNumber="56">anonymus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EF96ABB37C5A333BB2A7A568A01DFEC0" author="Friese" box="[672,811,332,357]" pageId="55" pageNumber="56" refString="Friese, H. (1911 b) Neue Varietaten von Bombus, III. Deutsche Entomologische Zeitschrift, 1911, 571 - 572." type="journal article" year="1911">
<collectorName id="26F2B3947C5A333BB2A7A568A1D1FEC1" box="[672,743,332,357]" pageId="55" pageNumber="56">Friese</collectorName>
, 1911
</bibRefCitation>
/ det.
<determinerName id="7E643A867C5A333BB370A569A73FFEC1" box="[887,1033,332,357]" pageId="55" pageNumber="56">PH Williams</determinerName>
2012; (7) white, printed [female]
<taxonomicName id="4C07ADC17C5A333BB090A54BA3C4FE2C" box="[151,242,367,392]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B9730A507C5A333BB090A54BA3C4FE2C" box="[151,242,367,392]" italics="true" pageId="55" pageNumber="56">Bombus</emphasis>
</taxonomicName>
/ (
<taxonomicName id="4C07ADC17C5A333BB113A54BA2FBFE2C" box="[276,461,367,392]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B9730A507C5A333BB113A54BA2FBFE2C" box="[276,461,367,392]" italics="true" pageId="55" pageNumber="56">Mendacibombus</emphasis>
</taxonomicName>
) /
<taxonomicName id="4C07ADC17C5A333BB1E8A54BA170FE2C" box="[495,582,367,392]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5A333BB1E8A54BA170FE2C" box="[495,582,367,392]" italics="true" pageId="55" pageNumber="56">mendax</emphasis>
</taxonomicName>
/
<collectorName id="26F2B3947C5A333BB267A54BA1D3FE2C" box="[608,741,367,392]" pageId="55" pageNumber="56">Gerstaecker</collectorName>
/ det
</materialsCitation>
.
<materialsCitation id="3B6FDC1F7C5A333BB336A554A104FE6B" ID-GBIF-Occurrence="1413110419" country="Philippines" location="This" municipality="Friese's" pageId="55" pageNumber="56" specimenCount="2" typeStatus="syntype">
<collectingCountry id="F31096D27C5A333BB336A554A063FE23" box="[817,853,368,391]" name="Philippines" pageId="55" pageNumber="56">PH</collectingCountry>
Williams 2012.
<location id="8ED880997C5A333BB41AA54BA779FE2C" LSID="urn:lsid:plazi:treatment:03AE67547C5B3334B090A5F8A727FDE4:8ED880997C5A333BB41AA54BA779FE2C" box="[1053,1103,367,392]" country="Philippines" municipality="Friese's" name="This" pageId="55" pageNumber="56">This</location>
queen, which is complete, is regarded as one of
<collectingMunicipality id="6BDC4C387C5A333BB17AA5B0A2E2FE09" box="[381,468,404,429]" pageId="55" pageNumber="56">Frieses</collectingMunicipality>
<typeStatus id="54BC68E07C5A333BB1D8A5B1A177FE09" box="[479,577,405,429]" pageId="55" pageNumber="56" type="syntype">syntypes</typeStatus>
and is designated here as
<typeStatus id="54BC68E07C5A333BB378A5B0A0D0FE09" box="[895,998,404,429]" pageId="55" pageNumber="56" type="lectotype">lectotype</typeStatus>
in order to reduce uncertainty in the identity and application of the name.
</materialsCitation>
</paragraph>
<paragraph id="8BB8D6427C5A333BB0C0A5F9A7BFFDBC" blockId="55.[151,1437,151,2034]" pageId="55" pageNumber="56">
Note 4 (
<emphasis id="B9730A507C5A333BB122A5F9A24DFE50" box="[293,379,477,500]" italics="true" pageId="55" pageNumber="56">pyrenes</emphasis>
). The absence of homonyms for the name
<emphasis id="B9730A507C5A333BB360A5F8A0C0FE51" box="[871,1014,476,501]" italics="true" pageId="55" pageNumber="56">latofasciatus</emphasis>
<bibRefCitation id="EF96ABB37C5A333BB3F9A5F9A7D9FE51" author="Vogt" box="[1022,1263,476,501]" pageId="55" pageNumber="56" refString="Vogt, O. (1909) Studien uber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1909, 28 - 84." type="journal article" year="1909">Vogt (1909: page 50)</bibRefCitation>
available from the same date or older (see note 2) makes Tkalcůs (1975) replacement name unnecessary.
</paragraph>
</subSubSection>
<subSubSection id="C31D85C97C5A333BB0C0A600A1AFF8C9" pageId="55" pageNumber="56" type="etymology">
<paragraph id="8BB8D6427C5A333BB0C0A600A787FDC4" blockId="55.[151,1437,151,2034]" pageId="55" pageNumber="56">
<emphasis id="B9730A507C5A333BB0C0A600A278FD99" bold="true" box="[199,334,548,573]" pageId="55" pageNumber="56">Etymology.</emphasis>
The species is named from the Latin
<taxonomicName id="4C07ADC17C5A333BB32EA600A0B6FD99" box="[809,896,548,573]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5A333BB32EA600A0B6FD99" box="[809,896,548,573]" italics="true" pageId="55" pageNumber="56">mendax</emphasis>
</taxonomicName>
for liar, presumably a reference to its close resemblance to other orange-tailed bumblebee species in the mountains of southern
<collectingCountry id="F31096D27C5A333BB444A66CA79BFDC4" box="[1091,1197,584,608]" name="Germany" pageId="55" pageNumber="56">Germany</collectingCountry>
.
</paragraph>
<paragraph id="8BB8D6427C5A333BB0C0A648A3FEFD69" blockId="55.[151,1437,151,2034]" pageId="55" pageNumber="56">
<emphasis id="B9730A507C5A333BB0C0A648A2C0FD21" bold="true" box="[199,502,620,645]" pageId="55" pageNumber="56">Taxonomy and variation.</emphasis>
The interpretation of this species here is based on DNA and the form of the male genitalia. It agrees with earlier interpretations, which include the substantial variation in the colour pattern of the hair.
</paragraph>
<paragraph id="8BB8D6427C5A333BB0C0A6F3A02CFC63" blockId="55.[151,1437,151,2034]" pageId="55" pageNumber="56">
This species shows unbanded (taxon
<emphasis id="B9730A507C5A333BB28CA6F3A01BFD54" box="[651,813,727,752]" italics="true" pageId="55" pageNumber="56">
<taxonomicName id="4C07ADC17C5A333BB28CA6F3A1D4FD54" box="[651,738,727,752]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" sensu="stricto" species="mendax">mendax</taxonomicName>
<taxonomicNameLabel id="A240B72B7C5A333BB2F6A6FDA01BFD54" box="[753,813,729,752]" pageId="55" pageNumber="56" sensu="stricto">s. str</taxonomicNameLabel>
</emphasis>
.) or narrowly yellow-grey-banded (taxon
<taxonomicName id="4C07ADC17C5A333BB52DA6FDA6AAFD54" box="[1322,1436,729,752]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="variety" species="mendax" variety="anonymus">
<emphasis id="B9730A507C5A333BB52DA6FDA6AAFD54" box="[1322,1436,729,752]" italics="true" pageId="55" pageNumber="56">anonymus</emphasis>
</taxonomicName>
MD#3540) colour patterns in the Alps (
<bibRefCitation id="EF96ABB37C5A333BB25AA6D8A034FCB0" author="Pittioni" box="[605,770,764,789]" pageId="55" pageNumber="56" refString="Pittioni, B. (1937 a) Die Hummelfauna des Kalsbachtales in Ost-Tirol. Ein Beitrag zur Okologie und Systematik der Hummeln Mitteleuropas. Festschrift zum 60. Geburtstag von Professor Dr. Embrik Strand, 3, 64 - 122." type="journal article" year="1937" yearSuffix="a">Pittioni 1937a</bibRefCitation>
), with more broadly yellow-banded colour patterns (taxon
<emphasis id="B9730A507C5A333BB090A73BA210FC9C" box="[151,294,799,824]" italics="true" pageId="55" pageNumber="56">latofasciatus</emphasis>
MD#1239) further west in the Pyrenees and in Santander in northern
<collectingCountry id="F31096D27C5A333BB457A73BA7A6FC9C" box="[1104,1168,799,824]" name="Spain" pageId="55" pageNumber="56">Spain</collectingCountry>
(
<bibRefCitation id="EF96ABB37C5A333BB4A7A704A62DFC9C" author="Vogt" box="[1184,1307,800,824]" pageId="55" pageNumber="56" refString="Vogt, O. (1909) Studien uber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1909, 28 - 84." type="journal article" year="1909">Vogt 1909</bibRefCitation>
;
<bibRefCitation id="EF96ABB37C5A333BB52EA704A3E5FCF9" author="Kruseman" pageId="55" pageNumber="56" refString="Kruseman, G. (1958) Notes sur les bourdons pyreneens du genre Bombus dans les collections neerlandaises. Beaufortia, 6, 161 - 170." type="journal article" year="1958">Kruseman 1958</bibRefCitation>
: his fig. 6). Our COI tree shows that specimens with these colour patterns can be indistinguishable in the available sequences and are interpreted as conspecific (
<figureCitation id="133CCAC77C5A333BB30AA743A055FC24" box="[781,867,871,896]" captionStart="FIGURE 13" captionStartId="16.[151,250,1733,1755]" captionTargetBox="[314,1260,194,1721]" captionTargetId="figure@16.[314,1261,193,1722]" captionTargetPageId="16" captionText="FIGURE 13. Assigning names to species from an estimate of phylogeny for 130 COI sequences of the subgenus Mendacibombus. Based on a BEAST analysis of the 30 unique haplotypes (Fig. 11), with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is added after the first matching sequence, with the addition of the two short-sequence yellow unbanded specimens from Pakistan, MD # 4058, 4059). The sequence labels follow the format used in Figs 11, 12. Lineages accepted as species from the GMYC models (Fig. 11) are shown as thick branches, the most recent common ancestors of each species are shown with a black spot, and branches within species are shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names (Table 1). The proxy sample for the type specimen for the oldest available name (the valid name) for each species (from Fig. 11) is marked in bold. The interpretations of the species with their valid names are shown in grey to the right." httpUri="https://zenodo.org/record/192309/files/figure.png" pageId="55" pageNumber="56">Fig. 13</figureCitation>
: the unbanded taxon
<emphasis id="B9730A507C5A333BB450A743A7DBFC24" box="[1111,1261,871,896]" italics="true" pageId="55" pageNumber="56">
<taxonomicName id="4C07ADC17C5A333BB450A743A798FC24" box="[1111,1198,871,896]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" sensu="stricto" species="mendax">mendax</taxonomicName>
<taxonomicNameLabel id="A240B72B7C5A333BB4B0A74DA7DBFC24" box="[1207,1261,873,896]" pageId="55" pageNumber="56" sensu="stricto">s. str</taxonomicNameLabel>
</emphasis>
. from the Alps MD#301, the narrow-banded taxon
<taxonomicName id="4C07ADC17C5A333BB237A7A9A197FC00" box="[560,673,909,932]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="variety" species="mendax" variety="anonymus">
<emphasis id="B9730A507C5A333BB237A7A9A197FC00" box="[560,673,909,932]" italics="true" pageId="55" pageNumber="56">anonymus</emphasis>
</taxonomicName>
from the Alps MD#303, and the broad-banded taxon
<emphasis id="B9730A507C5A333BB50AA7A8A6AAFC01" box="[1293,1436,908,933]" italics="true" pageId="55" pageNumber="56">latofasciatus</emphasis>
from the Pyrenees MD#3857), as parts of
<emphasis id="B9730A507C5A333BB269A795A025FC6C" box="[622,787,943,968]" italics="true" pageId="55" pageNumber="56">
<taxonomicName id="4C07ADC17C5A333BB269A795A1D2FC6C" box="[622,740,943,968]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" sensu="lato" species="mendax">B. mendax</taxonomicName>
<taxonomicNameLabel id="A240B72B7C5A333BB2ECA795A025FC6C" box="[747,787,943,968]" pageId="55" pageNumber="56" sensu="lato">s. l.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
<paragraph id="8BB8D6427C5A333BB0C0A7F0A64BFBB4" blockId="55.[151,1437,151,2034]" pageId="55" pageNumber="56">
The distribution of this species is interrupted by two relatively small disjunctions (
<emphasis id="B9730A507C5A333BB462A7F0A610FC48" box="[1125,1318,980,1004]" italics="true" pageId="55" pageNumber="56">
cf.
<taxonomicName id="4C07ADC17C5A333BB48FA7F1A610FC48" box="[1160,1318,981,1004]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="margreiteri">B. margreiteri</taxonomicName>
</emphasis>
): between the Alps and Pyrenees (&gt;
<quantity id="4CFF7BA77C5A333BB1B0A7D3A127FBB4" box="[439,529,1015,1040]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="55" pageNumber="56" unit="km" value="500.0">500 km</quantity>
); and between the Pyrenees and the Picos de Europa in Santander (&gt;
<quantity id="4CFF7BA77C5A333BB510A7DCA646FBB4" box="[1303,1392,1015,1040]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="55" pageNumber="56" unit="km" value="300.0">300 km</quantity>
).
</paragraph>
<paragraph id="8BB8D6427C5A333BB0C0A038A0D6FAF0" blockId="55.[151,1437,151,2034]" pageId="55" pageNumber="56">
The RMNH collection includes at least two queens each with a label printed with Tunkun / Sayan and a handwritten label
<taxonomicName id="4C07ADC17C5A333BB16EA01BA2F6FBFC" box="[361,448,1087,1112]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5A333BB16EA01BA2F6FBFC" box="[361,448,1087,1112]" italics="true" pageId="55" pageNumber="56">mendax</emphasis>
</taxonomicName>
. One is a queen of
<taxonomicName id="4C07ADC17C5A333BB2B5A065A065FBFC" box="[690,851,1089,1112]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="margreiteri">
<emphasis id="B9730A507C5A333BB2B5A065A065FBFC" box="[690,851,1089,1112]" italics="true" pageId="55" pageNumber="56">B. margreiteri</emphasis>
</taxonomicName>
(MD#1144), for which the Tunkin mountains are consistent with its known distribution (
<figureCitation id="133CCAC77C5A333BB27DA040A1E0FBD9" box="[634,726,1124,1149]" captionStart="FIGURES 56 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="55" pageNumber="56">Fig. 65</figureCitation>
). However, the other queen (MD#1280) appears from its morphology and colour pattern to be a genuine queen of
<taxonomicName id="4C07ADC17C5A333BB32AA0ADA090FB04" box="[813,934,1159,1184]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5A333BB32AA0ADA090FB04" box="[813,934,1159,1184]" italics="true" pageId="55" pageNumber="56">B. mendax</emphasis>
</taxonomicName>
. This specimen has the hair of the thoracic dorsum anteriorly with a narrow pale greyish band with black intermixed, whereas the side of the thorax, the scutellum, T2, and the foreleg and midleg basitarsi are predominantly black, like many
<taxonomicName id="4C07ADC17C5A333BB47AA0F5A7C3FB4C" box="[1149,1269,1231,1256]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5A333BB47AA0F5A7C3FB4C" box="[1149,1269,1231,1256]" italics="true" pageId="55" pageNumber="56">B. mendax</emphasis>
</taxonomicName>
from the Alps (
<bibRefCitation id="EF96ABB37C5A333BB099A0D0A272FAA8" author="Pittioni" box="[158,324,1268,1293]" pageId="55" pageNumber="56" refString="Pittioni, B. (1937 a) Die Hummelfauna des Kalsbachtales in Ost-Tirol. Ein Beitrag zur Okologie und Systematik der Hummeln Mitteleuropas. Festschrift zum 60. Geburtstag von Professor Dr. Embrik Strand, 3, 64 - 122." type="journal article" year="1937" yearSuffix="a">Pittioni 1937a</bibRefCitation>
). Without COI data we cannot at present make a definitive determination. Therefore the apparent Tunkin location is likely to be the result of mislabeling and this record is not included on the map for
<taxonomicName id="4C07ADC17C5A333BB523A13DA6AAFA94" box="[1316,1436,1303,1328]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B9730A507C5A333BB523A13DA6AAFA94" box="[1316,1436,1303,1328]" italics="true" pageId="55" pageNumber="56">B. mendax</emphasis>
</taxonomicName>
(mislabelling is not unknown: see the note on the
<typeStatus id="54BC68E07C5A333BB2C2A118A011FAF1" box="[709,807,1340,1365]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C07ADC17C5A333BB34BA119A0E6FAF1" box="[844,976,1340,1365]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="superbus">
<emphasis id="B9730A507C5A333BB34BA119A0E6FAF1" box="[844,976,1340,1365]" italics="true" pageId="55" pageNumber="56">B. superbus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BB8D6427C5A333BB0C0A17BA1AFF8C9" blockId="55.[151,1437,151,2034]" pageId="55" pageNumber="56">
<emphasis id="B9730A507C5A333BB0C0A17BA2EFFADC" bold="true" box="[199,473,1375,1400]" pageId="55" pageNumber="56">Diagnostic description.</emphasis>
Wings nearly clear.
<emphasis id="B9730A507C5A333BB2B9A17BA0CFFADC" box="[702,1017,1375,1400]" italics="true" pageId="55" pageNumber="56">Female hair colour pattern:</emphasis>
generally black, but with yellow hair varying from completely absent from the entire body to present over most of the face and in a small patch on the vertex of the head, a transverse band anteriorly on the thoracic dorsum and extending laterally and ventrally almost to the midleg base, in a transverse band posteriorly on the thoracic dorsum (scutellum; so the thoracic dorsum between the wing bases has the hair entirely black, sometimes the yellow bands with black hairs intermixed), often on T1, sometimes on T2 pale in the anterior half, always orange hair on T3 as a posterior fringe and throughout T46, except T6 medially with black hair. Hindleg tibia with the corbicular fringes extensively orange.
<emphasis id="B9730A507C5A333BB54FA203A211F9C6" italics="true" pageId="55" pageNumber="56">Female morphology:</emphasis>
labrum with the basal depression broad, the transverse ridge in the median third narrowed but not interrupted or subsiding, or subsiding only slightly, with only a few scattered punctures, the lateral tubercles with few punctures.
<emphasis id="B9730A507C5A333BB143A2A8A13BF901" box="[324,525,1676,1701]" italics="true" pageId="55" pageNumber="56">Male morphology</emphasis>
: genitalia (
<figureCitation id="133CCAC77C5A333BB289A2A8A1D6F901" box="[654,736,1676,1701]" captionStart="FIGURES 24 35" captionStartId="26.[151,264,1867,1889]" captionTargetBox="[233,1387,193,1789]" captionTargetId="figure@26.[233,1390,193,1894]" captionTargetPageId="26" captionText="FIGURES 24 35. Morphology of the male genitalia for species of the subgenus Mendacibombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 24, B. superbus, China MD # 339; 25, B. waltoni, China MD # 216; 26, B. convexus, China MD # 215; 27, B. makarjini, Kazakhstan MD # 1240; 28, B. marussinus, Pakistan MD # 408; 29, B. avinoviellus, India MD # 404; 30, B. himalayanus, India MD # 426; 31, B. turkestanicus, China MD # 351; 32, B. defector, Kyrgyzstan MD # 367; 33, B. margreiteri, China MD # 294; 34, B. handlirschianus, Russia MD # 835; 35, B. mendax, Spain MD # 689." httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="55" pageNumber="56">Fig. 35</figureCitation>
) with the volsella distally rounded (finger-shaped) and curled back dorsally but not anteriorly; volsella at its broadest near the midpoint of its length, the dorsal surface just distal to this point without a raised curved ridge just inside the inner margin. Gonostylus distally thick, rounded in section, and finger-like; gonostylus with the inner distal corner almost a right angle from both the inner and dorsal aspects. Penis-valve inner shoulder located at Ĺ 0.5× the length of the penis valve from the distal end to the broadest point of the spatha; penis valve proximal to the outer shoulder&gt; 2× as broad as the penis-valve head; penis-valve head dorso-ventrally compressed.
</paragraph>
</subSubSection>
<subSubSection id="C31D85C97C5A3334B0C0A351A6AAFDBB" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="8BB8D6427C5A333BB0C0A351A7BFF856" blockId="55.[151,1437,151,2034]" pageId="55" pageNumber="56">
<emphasis id="B9730A507C5A333BB0C0A351A286F82A" bold="true" box="[199,432,1909,1934]" pageId="55" pageNumber="56">Material examined.</emphasis>
383 queens 1249 workers
<specimenCount id="9D011DCB7C5A333BB2DDA352A07AF82A" box="[730,844,1909,1934]" pageId="55" pageNumber="56" type="male">142 males</specimenCount>
(plus 6 with sex/caste undetermined), from
<collectingRegion id="49C318A07C5A333BB53FA351A6AEF82A" box="[1336,1432,1909,1934]" country="Andorra" name="Andorra la Vella" pageId="55" pageNumber="56">
<collectingCountry id="F31096D27C5A333BB53FA351A6AEF82A" box="[1336,1432,1909,1934]" name="Andorra" pageId="55" pageNumber="56">Andorra</collectingCountry>
</collectingRegion>
,
<collectingCountry id="F31096D27C5A333BB090A3B2A3DBF80B" box="[151,237,1942,1967]" name="Austria" pageId="55" pageNumber="56">Austria</collectingCountry>
,
<collectingCountry id="F31096D27C5A333BB0FEA3B3A27EF80A" box="[249,328,1943,1966]" name="France" pageId="55" pageNumber="56">France</collectingCountry>
,
<collectingCountry id="F31096D27C5A333BB152A3B3A288F80B" box="[341,446,1943,1967]" name="Germany" pageId="55" pageNumber="56">Germany</collectingCountry>
,
<collectingCountry id="F31096D27C5A333BB1CCA3B2A2C9F80B" box="[459,511,1942,1967]" name="Italy" pageId="55" pageNumber="56">Italy</collectingCountry>
,
<collectingCountry id="F31096D27C5A333BB20AA3B2A166F80B" box="[525,592,1942,1967]" name="Spain" pageId="55" pageNumber="56">Spain</collectingCountry>
, and
<collectingCountry id="F31096D27C5A333BB289A3B2A022F80B" box="[654,788,1942,1967]" name="Switzerland" pageId="55" pageNumber="56">Switzerland</collectingCountry>
(
<figureCitation id="133CCAC77C5A333BB323A3B2A04DF80B" box="[804,891,1942,1967]" captionStart="FIGURES 56 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="55" pageNumber="56">Fig. 67</figureCitation>
: AA,
<collectingCountry id="F31096D27C5A333BB3C7A3B3A0D0F80B" box="[960,998,1943,1967]" name="American Samoa" pageId="55" pageNumber="56">AS</collectingCountry>
,
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, LC,
<collectionCode id="ED164E877C5A333BB48EA3B3A7D7F80A" box="[1161,1249,1943,1966]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="55" pageNumber="56">MNHN</collectionCode>
,
<collectionCode id="ED164E877C5A333BB4E9A3B3A671F80A" box="[1262,1351,1943,1966]" pageId="55" pageNumber="56">MNHU</collectionCode>
,
<collectionCode id="ED164E877C5A333BB553A3B3A6AEF80A" box="[1364,1432,1943,1966]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="55" pageNumber="56">NHM</collectionCode>
, PW,
<collectionCode id="ED164E877C5A333BB0D7A39DA220F874" box="[208,278,1977,2000]" pageId="55" pageNumber="56">RBIN</collectionCode>
,
<collectionCode id="ED164E877C5A333BB123A39DA24DF874" box="[292,379,1977,2000]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="55" pageNumber="56">RMNH</collectionCode>
,
<collectionCode id="ED164E877C5A333BB18EA39CA2C7F874" box="[393,497,1976,2000]" pageId="55" pageNumber="56">UMONS</collectionCode>
,
<collectionCode id="ED164E877C5A333BB207A39DA172F874" box="[512,580,1977,2000]" pageId="55" pageNumber="56">ZIUB</collectionCode>
,
<collectionCode id="ED164E877C5A333BB254A39CA1A6F874" box="[595,656,1976,2000]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="55" pageNumber="56">ZSM</collectionCode>
; specimens in the
<collectionCode id="ED164E877C5A333BB36DA39CA0F9F874" box="[874,975,1976,2000]" pageId="55" pageNumber="56">UMONS</collectionCode>
database have not had individual MD# labels added), with
<specimenCount id="9D011DCB7C5A333BB175A3FEA2CAF856" box="[370,508,2009,2034]" pageId="55" pageNumber="56" type="generic">7 specimens</specimenCount>
sequenced (interpretable sequences listed in
<figureCitation id="133CCAC77C5A333BB3F2A3FDA74AF856" box="[1013,1148,2009,2034]" captionStart-0="FIGURES 11 12" captionStart-1="FIGURE 13" captionStartId-0="15.[151,264,1630,1652]" captionStartId-1="16.[151,250,1733,1755]" captionTargetBox-0="[300,1121,202,1610]" captionTargetBox-1="[314,1260,194,1721]" captionTargetId-0="figure@15.[292,1266,193,1625]" captionTargetId-1="figure@16.[314,1261,193,1722]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 11 12. Recognising species from an estimate of phylogeny for 30 unique COI haplotypes of the subgenus Mendacibombus by BEAST, using GMYC models and the PTP procedure. From the GMYC models (11), the single threshold T is shown by the vertical grey bar so that the intersecting lineages are interpreted as indicating 12 separate species (black spots at coalescent nodes; outgroups not shown). Values below the nodes of the maximum clade-credibility tree are Bayesian posterior probabilities showing branch support (groups with values &lt;0.8 are considered unreliable) from the BEAST analysis with two billion MCMC generations and a 1 % burn-in. The x - scale axis is calibrated in substitutions per nucleotide site. From the PTP procedure (12), the Bayesian PTP solution with highest support also indicates 12 species. Values above the nodes are PTP Bayesian support values that all daughter haplotypes are parts of a single species; values below the nodes are Bayesian posterior probabilities showing branch support. The scale bar is calibrated in substitutions per nucleotide site. Each unique haplotype is represented by one of the longest available sample sequences, labelled with: the sequence length in number of base pairs, a taxon name or name for an unpublished colour pattern (light / dark / yellow / white / pale), and then a code that consists of a specimen identifier (MD #) from the project database and (after the hyphen) a sample identifier from BOLD or from GenBank, followed with its geographic origin." captionText-1="FIGURE 13. Assigning names to species from an estimate of phylogeny for 130 COI sequences of the subgenus Mendacibombus. Based on a BEAST analysis of the 30 unique haplotypes (Fig. 11), with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is added after the first matching sequence, with the addition of the two short-sequence yellow unbanded specimens from Pakistan, MD # 4058, 4059). The sequence labels follow the format used in Figs 11, 12. Lineages accepted as species from the GMYC models (Fig. 11) are shown as thick branches, the most recent common ancestors of each species are shown with a black spot, and branches within species are shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names (Table 1). The proxy sample for the type specimen for the oldest available name (the valid name) for each species (from Fig. 11) is marked in bold. The interpretations of the species with their valid names are shown in grey to the right." httpUri-0="https://zenodo.org/record/192308/files/figure.png" httpUri-1="https://zenodo.org/record/192309/files/figure.png" pageId="55" pageNumber="56">Figs. 1113</figureCitation>
).
</paragraph>
<paragraph id="8BB8D6427C553334B0C0A4B3A3D6FE92" blockId="56.[151,1437,151,576]" pageId="56" pageNumber="57">
<emphasis id="B9730A507C553334B0C0A4B3A2C7FF14" bold="true" box="[199,497,151,176]" pageId="56" pageNumber="57">Habitat and distribution.</emphasis>
Flower-rich alpine and subalpine grassland, at elevations 655(2111)
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a.s.l.. A species distributed in three principal disjunct centres, in the Alps, the Pyrenees, and the Cantabrian mountains. There is no overlap in distribution with any other
<taxonomicName id="4C07ADC17C553334B2E4A4FEA0AAFF57" box="[739,924,218,243]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B9730A507C553334B2E4A4FEA0AAFF57" box="[739,924,218,243]" italics="true" pageId="56" pageNumber="57">Mendacibombus</emphasis>
</taxonomicName>
species. Distribution maps are available for
<collectingCountry id="F31096D27C553334B090A4D9A3D5FEB0" box="[151,227,253,276]" name="France" pageId="56" pageNumber="57">France</collectingCountry>
(
<bibRefCitation id="EF96ABB37C553334B0F2A4D8A296FEB0" author="Rasmont" box="[245,416,252,276]" pageId="56" pageNumber="57" refString="Rasmont, P. (1988) Monographie ecologique et zoogeographique des bourdons de France et de Belgique (Hymenoptera, Apidae, Bombinae). In: Faculte des Sciences agronomiques de l'Etat, Gembloux, p. 309 + lxi." type="book chapter" year="1988">Rasmont 1988</bibRefCitation>
),
<collectingCountry id="F31096D27C553334B1B0A4D8A2C1FEB1" box="[439,503,252,277]" name="Spain" pageId="56" pageNumber="57">Spain</collectingCountry>
(
<bibRefCitation id="EF96ABB37C553334B20EA4D8A022FEB0" author="Ornosa" box="[521,788,252,276]" pageId="56" pageNumber="57" refString="Ornosa, C. &amp; Torres, F. (2010) Mendacibombus (Mendacibombus) mendax (Gerstaecker, 1869). In: Atlas de Invertebrados Amenazados de Espana, pp. 160 - 164." type="book chapter" year="2010">Ornosa &amp; Torres 2010</bibRefCitation>
), and Europe (
<bibRefCitation id="EF96ABB37C553334B3C7A4D8A7D5FEB0" author="Rasmont" box="[960,1251,252,277]" pageId="56" pageNumber="57" refString="Rasmont, P. &amp; Iserbyt, S. (2012) Atlas of the European bees: genus Bombus In: STEP Project, Atlas Hymenoptera, Mons, Gembloux." type="book" year="2012">Rasmont &amp; Iserbyt 2012</bibRefCitation>
;
<bibRefCitation id="EF96ABB37C553334B4F4A4D9A3E2FE92" author="Rasmont" pageId="56" pageNumber="57" refString="Rasmont, P., Franzen, M., Lecocq, T., Harpke, A., Roberts, S. P. M., Biesmeijer, J. C., et al. (2015) Climatic risk and distribution atlas of European bumblebees, Pensoft." type="book" year="2015">
Rasmont
<emphasis id="B9730A507C553334B567A4D9A6AAFEB1" box="[1376,1436,252,277]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
<paragraph id="8BB8D6427C553334B0C0A51BA708FE5A" blockId="56.[151,1437,151,576]" pageId="56" pageNumber="57">
So far, this is the only species of the subgenus
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<emphasis id="B9730A507C553334B2C8A51AA0BEFEF3" box="[719,904,318,343]" italics="true" pageId="56" pageNumber="57">Mendacibombus</emphasis>
</taxonomicName>
to have been assessed for Red List threat status using the
<collectionCode id="ED164E877C553334B10FA544A27AFEDC" box="[264,332,352,376]" pageId="56" pageNumber="57">IUCN</collectionCode>
criteria (2001).
<bibRefCitation id="EF96ABB37C553334B20FA57BA024FEDC" box="[520,786,351,376]" pageId="56" pageNumber="57" refString="Cederberg, B., Michez, D., Nieto, A., Radchenko, V., Rasmont, P. &amp; Roberts, S. P. M. (2013) Bombus mendax. In: IUCN Red List of Threatened Species. IUCN. Avaliable from: http: // www. iucnredlist. org (Accessed 1 Dec. 2016)" type="journal article">
Cederberg
<emphasis id="B9730A507C553334B282A545A181FEDC" box="[645,695,351,376]" italics="true" pageId="56" pageNumber="57">et al</emphasis>
. (2013)
</bibRefCitation>
listed it as Near Threatened because its known area of occupancy (AOO) is small (
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²), the species distribution is considered severely fragmented, and they believe that the number of individuals has been declining due to climate change, putting the species close to qualifying for the
<collectionCode id="ED164E877C553334B166A5E0A293FE78" box="[353,421,452,476]" pageId="56" pageNumber="57">IUCN</collectionCode>
category Vulnerable (Criterion B2).
<bibRefCitation id="EF96ABB37C553334B352A5E1A77FFE79" box="[853,1097,452,477]" pageId="56" pageNumber="57" refString="Rasmont, P., Franzen, M., Lecocq, T., Harpke, A., Roberts, S. P. M., Biesmeijer, J. C., et al. (2015) Climatic risk and distribution atlas of European bumblebees, Pensoft." type="book">
Rasmont
<emphasis id="B9730A507C553334B3C7A5E1A0C6FE79" box="[960,1008,452,477]" italics="true" pageId="56" pageNumber="57">et al</emphasis>
. (2015)
</bibRefCitation>
use climate-change models to assess this species as having a high risk of extinction from climate change by 2100.
</paragraph>
<paragraph id="8BB8D6427C553334B0C0A622A6AAFDBB" blockId="56.[151,1437,151,576]" box="[199,1436,518,543]" pageId="56" pageNumber="57">
<emphasis id="B9730A507C553334B0C0A622A26CFDBB" bold="true" box="[199,346,518,543]" pageId="56" pageNumber="57">Food plants.</emphasis>
<bibRefCitation id="EF96ABB37C553334B167A622A125FDBB" author="Pittioni" box="[352,531,518,543]" pageId="56" pageNumber="57" refString="Pittioni, B. (1937 a) Die Hummelfauna des Kalsbachtales in Ost-Tirol. Ein Beitrag zur Okologie und Systematik der Hummeln Mitteleuropas. Festschrift zum 60. Geburtstag von Professor Dr. Embrik Strand, 3, 64 - 122." type="journal article" year="1937" yearSuffix="a">Pittioni (1937a)</bibRefCitation>
,
<bibRefCitation id="EF96ABB37C553334B219A622A1EDFDBB" author="Aichhorn" box="[542,731,518,543]" pageId="56" pageNumber="57" refString="Aichhorn, A. (1976) Beitrag zur Hummelzucht zur Biologie von Bombus mendax. Berichte aus dem Haus der Natur Salzburg, 7, 13 - 31." type="journal article" year="1976">Aichhorn (1976)</bibRefCitation>
,
<bibRefCitation id="EF96ABB37C553334B2E1A623A047FDBB" author="Haas" box="[742,881,518,543]" pageId="56" pageNumber="57" refString="Haas, A. (1976) Paarungsverhalten und Nestbau der alpinen Hummelart Bombus mendax (Hymenoptera: Apidae). Entomologica Germanica, 3, 248 - 259." type="journal article" year="1976">Haas (1976)</bibRefCitation>
,
<bibRefCitation id="EF96ABB37C553334B37BA623A705FDBB" author="Rasmont" box="[892,1075,519,543]" pageId="56" pageNumber="57" refString="Rasmont, P. (1988) Monographie ecologique et zoogeographique des bourdons de France et de Belgique (Hymenoptera, Apidae, Bombinae). In: Faculte des Sciences agronomiques de l'Etat, Gembloux, p. 309 + lxi." type="book chapter" year="1988">Rasmont (1988)</bibRefCitation>
, von
<bibRefCitation id="EF96ABB37C553334B468A623A6AEFDBB" author="Hagen" box="[1135,1432,518,543]" pageId="56" pageNumber="57" refString="Hagen, E. v. &amp; Aichhorn, A. (2003) Hummeln: Bestimmen, Ansiedeln, Vermehren, Schutzen (2 ed.). Nottuln, Fauna - Verlag." type="book chapter" year="2003">Hagen &amp; Aichhorn (2003)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D85C97C553334B0C0A603A727FDE4" box="[199,1041,551,576]" pageId="56" pageNumber="57" type="description">
<paragraph id="8BB8D6427C553334B0C0A603A727FDE4" blockId="56.[151,1437,151,576]" box="[199,1041,551,576]" pageId="56" pageNumber="57">
<emphasis id="B9730A507C553334B0C0A603A27EFDE4" bold="true" box="[199,328,551,576]" pageId="56" pageNumber="57">Behaviour.</emphasis>
<bibRefCitation id="EF96ABB37C553334B148A603A13AFDE4" author="Aichhorn" box="[335,524,551,576]" pageId="56" pageNumber="57" refString="Aichhorn, A. (1976) Beitrag zur Hummelzucht zur Biologie von Bombus mendax. Berichte aus dem Haus der Natur Salzburg, 7, 13 - 31." type="journal article" year="1976">Aichhorn (1976)</bibRefCitation>
,
<bibRefCitation id="EF96ABB37C553334B21FA60CA192FDE4" author="Haas" box="[536,676,551,576]" pageId="56" pageNumber="57" refString="Haas, A. (1976) Paarungsverhalten und Nestbau der alpinen Hummelart Bombus mendax (Hymenoptera: Apidae). Entomologica Germanica, 3, 248 - 259." type="journal article" year="1976">Haas (1976)</bibRefCitation>
, von
<bibRefCitation id="EF96ABB37C553334B2E6A60CA73AFDE4" author="Hagen" box="[737,1036,551,576]" pageId="56" pageNumber="57" refString="Hagen, E. v. &amp; Aichhorn, A. (2003) Hummeln: Bestimmen, Ansiedeln, Vermehren, Schutzen (2 ed.). Nottuln, Fauna - Verlag." type="book chapter" year="2003">Hagen &amp; Aichhorn (2003)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>