191 lines
18 KiB
XML
191 lines
18 KiB
XML
<document id="115C186B0646104826BA0378D16B4300" ID-DOI="10.11646/palaeoentomology.6.3.8" ID-ISSN="2624-2834" ID-Zenodo-Dep="8073095" ID-ZooBank="43173F45-5BC4-40F6-9BCA-DD8A876F7FF4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1687503302260" checkinUser="plazi" docAuthor="BASHKUEV, ALEXEY S. & JARZEMBOWSKI, EDMUND A." docDate="2023" docId="03A3EE79EC01FFFEFF79FF586835B584" docLanguage="en" docName="palaeoentomology.6.3.8.pdf" docOrigin="Palaeoentomology 6 (3)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.3.8" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Typhothauma agari BASHKUEV & JARZEMBOWSKI 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="258" masterDocId="FF9A9601EC00FFFDFFF1F815691CB758" masterDocTitle="The first British Cretaceous eomeropid scorpionfly (Mecoptera: Eomeropidae)" masterLastPageNumber="259" masterPageNumber="255" pageNumber="256" updateTime="1687503664218" updateUser="ExternalLinkService">
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<mods:mods id="FF2039CAEAADB660ED6E87F6D6F055F8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7760D025CF145FB742F6DB579E5BC53A">
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<mods:title id="FABA7056C83E23045FCBD5A7845C4552">The first British Cretaceous eomeropid scorpionfly (Mecoptera: Eomeropidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="6EDA33667FC1E98422D9300BFAE05FAF" type="personal">
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<mods:role id="35F412CC5D1DA7AEC3821728285C7890">
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<mods:roleTerm id="DC2109DFF2B67E45BC7CD030B074C10B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2FB2E126C72E317BC3C4B5CEA2866478">BASHKUEV, ALEXEY S.</mods:namePart>
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</mods:name>
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<mods:name id="69C1B3717DA4A13B381A7193D8C211F1" type="personal">
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<mods:role id="A4A08FE451D292AD48B29D9DE8EFBBAE">
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<mods:roleTerm id="3DED175CD71F28924013E98126070760">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EA6396D6EC0FDD19C74E74D4183973E9">JARZEMBOWSKI, EDMUND A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="31B8EBD4DDA3AD34AB0350C33F12A9F8">text</mods:typeOfResource>
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<mods:relatedItem id="C11672CAC8A86AA3A7EC01A0E7C7C682" type="host">
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<mods:titleInfo id="830D5B1860F4D72E08D268535FEEAF0F">
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<mods:title id="B401FF36691340CF6C2AB311D318A951">Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part id="3A714AEC4EECD3B3DBCAD3CAB0278095">
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<mods:date id="62A0CCD99454BC697D64823889407CDF">2023</mods:date>
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<mods:detail id="BD80CB99E4A381A286B7FE74FFBD50DA" type="pubDate">
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<mods:number id="4AE09A4AFAB9B240CCB7A07E7389ECED">2023-06-19</mods:number>
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</mods:detail>
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<mods:detail id="47397351C50521B53904C026DBB2D714" type="volume">
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<mods:number id="EDAF537E878471EDAD80A83264364050">6</mods:number>
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</mods:detail>
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<mods:detail id="3F08AB1315C5972E61DD16C22B4291CD" type="issue">
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<mods:number id="66C33D3FA16A57BD93634441BE6F951D">3</mods:number>
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</mods:detail>
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<mods:extent id="A78F31FF03B3862C735A896AC6BDAC15" unit="page">
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<mods:start id="D0B0319CDB04DAA29CD1842CA849C631">255</mods:start>
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<mods:end id="1E3F5C194F80DC83FEA80A75B905E294">259</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location id="0E36124BD41D433E131A513117F6F702">
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<mods:url id="24AED186C3D7923772396D081CB00568">http://dx.doi.org/10.11646/palaeoentomology.6.3.8</mods:url>
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</mods:location>
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<mods:classification id="D1DED5FB4CE0790248FEB99ECD1D02CE">journal article</mods:classification>
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<mods:identifier id="ACACD33FB5E9AE7C02F9C36CC5CDF3D8" type="DOI">10.11646/palaeoentomology.6.3.8</mods:identifier>
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<mods:identifier id="FCD7D59FABF7925D80BE0767874D4C17" type="ISSN">2624-2834</mods:identifier>
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<mods:identifier id="20274A2EF153DE5A89853B1A29502EE3" type="Zenodo-Dep">8073095</mods:identifier>
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<mods:identifier id="AE123EAD78821504C2BAD6E30A9EA979" type="ZooBank">43173F45-5BC4-40F6-9BCA-DD8A876F7FF4</mods:identifier>
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</mods:mods>
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<treatment id="03A3EE79EC01FFFEFF79FF586835B584" LSID="urn:lsid:plazi:treatment:03A3EE79EC01FFFEFF79FF586835B584" httpUri="http://treatment.plazi.org/id/03A3EE79EC01FFFEFF79FF586835B584" lastPageId="3" lastPageNumber="258" pageId="1" pageNumber="256">
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<subSubSection id="C3100CE4EC01FFFCFF79FF5868C7B030" box="[136,475,1869,1896]" pageId="1" pageNumber="256" type="nomenclature">
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<paragraph id="8BB55F6FEC01FFFCFF79FF5868C7B030" blockId="1.[136,475,1869,1935]" box="[136,475,1869,1896]" pageId="1" pageNumber="256">
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<heading id="D0FDE803EC01FFFCFF79FF5868C7B030" bold="true" box="[136,475,1869,1896]" fontSize="10" level="2" pageId="1" pageNumber="256" reason="3">
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<emphasis id="B97E837DEC01FFFCFF79FF5868C7B030" bold="true" box="[136,475,1869,1896]" pageId="1" pageNumber="256">
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?
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<taxonomicName id="4C0A24ECEC01FFFCFF67FF586867B03F" authority="BASHKUEV & JARZEMBOWSKI, 2023" authorityName="BASHKUEV & JARZEMBOWSKI" authorityYear="2023" box="[150,379,1869,1896]" class="Insecta" family="Eomeropidae" genus="Typhothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="agari" status="sp. nov.">
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<emphasis id="B97E837DEC01FFFCFF67FF586867B03F" bold="true" box="[150,379,1869,1896]" italics="true" pageId="1" pageNumber="256">Typhothauma agari</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24D3E06EC01FFFCFE73FF5B68C7B030" box="[386,475,1870,1896]" pageId="1" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3100CE4EC01FFFCFF79FF6269C4B0D7" box="[136,216,1911,1935]" pageId="1" pageNumber="256" type="description">
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<paragraph id="8BB55F6FEC01FFFCFF79FF6269C4B0D7" blockId="1.[136,475,1869,1935]" box="[136,216,1911,1935]" pageId="1" pageNumber="256">
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(
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<figureCitation id="133143EAEC01FFFCFF61FF6269CCB0D7" box="[144,208,1911,1935]" captionStart="FIGURE 2" captionStartId="2.[136,235,1951,1975]" captionTargetBox="[174,1415,189,1924]" captionTargetId="figure-29@2.[170,1418,189,1927]" captionTargetPageId="2" captionText="FIGURE 2.?Typhothauma agari sp. nov. A, B, Photographs of the holotype in different optical and illumination settings. C, Venation interpretation, with the hypothetical position of Sc3. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8073099" httpUri="https://zenodo.org/record/8073099/files/figure.png" pageId="1" pageNumber="256">Fig. 2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3100CE4EC01FFFCFF79FFD068B8BF74" pageId="1" pageNumber="256" type="materials_examined">
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<paragraph id="8BB55F6FEC01FFFCFF79FFD068B8BF74" blockId="1.[136,778,1989,2092]" pageId="1" pageNumber="256">
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<emphasis id="B97E837DEC01FFFCFF79FFD069EEB085" bold="true" box="[136,242,1989,2013]" pageId="1" pageNumber="256">Material.</emphasis>
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<materialsCitation id="3B625532EC01FFFCFF06FFD068B8BF74" collectionCode="NHMUK" pageId="1" pageNumber="256" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54B1E1CDEC01FFFCFF06FFD06846B085" box="[247,346,1989,2013]" pageId="1" pageNumber="256">Holotype</typeStatus>
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<collectionCode id="ED1BC7AAEC01FFFCFE91FFD368D9B085" box="[352,453,1990,2013]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="1" pageNumber="256" type="Museum">NHMUK</collectionCode>
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In 64761 [S 1164], impression (without counterpart) of forewing, deposited in the Natural History Museum, London.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3100CE4EC01FFFCFCAEFECD6AA3B04B" pageId="1" pageNumber="256" type="etymology">
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<paragraph id="8BB55F6FEC01FFFCFCAEFECD6CB0B1B7" blockId="1.[810,1452,1751,2092]" box="[863,1452,1751,1776]" pageId="1" pageNumber="256">
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<emphasis id="B97E837DEC01FFFCFCAEFECD6AFCB1A8" bold="true" box="[863,992,1752,1776]" pageId="1" pageNumber="256">Etymology.</emphasis>
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Named after amateur Wealden geologist
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</paragraph>
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<paragraph id="8BB55F6FEC01FFFCFCDBFEEE6AA3B04B" blockId="1.[810,1452,1751,2092]" box="[810,959,1787,1811]" pageId="1" pageNumber="256">Richard Agar.</paragraph>
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</subSubSection>
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<subSubSection id="C3100CE4EC01FFFCFCAEFF0B6ABEBF74" pageId="1" pageNumber="256" type="diagnosis">
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<paragraph id="8BB55F6FEC01FFFCFCAEFF0B6CB7B001" blockId="1.[810,1452,1751,2092]" pageId="1" pageNumber="256">
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<emphasis id="B97E837DEC01FFFCFCAEFF0B6AC8B06E" bold="true" box="[863,980,1822,1846]" pageId="1" pageNumber="256">Diagnosis.</emphasis>
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Differs from
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<taxonomicName id="4C0A24ECEC01FFFCFB92FF0A6DD1B06E" authorityName="Zhang, Shih & Ren" authorityYear="2012" box="[1123,1229,1822,1846]" class="Insecta" family="Eomeropidae" genus="Typhothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="excelsa">
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<emphasis id="B97E837DEC01FFFCFB92FF0A6D6EB06E" box="[1123,1138,1823,1846]" italics="true" pageId="1" pageNumber="256">T</emphasis>
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.
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<emphasis id="B97E837DEC01FFFCFB8EFF0B6DD1B06E" box="[1151,1229,1822,1846]" italics="true" pageId="1" pageNumber="256">excelsa</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C0A24ECEC01FFFCFB0EFF0A6C8DB06E" authorityName="Ren & Shih" authorityYear="2005" box="[1279,1425,1822,1846]" class="Insecta" family="Eomeropidae" genus="Typhothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="yixianensis">
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<emphasis id="B97E837DEC01FFFCFB0EFF0A6C12B06E" box="[1279,1294,1823,1846]" italics="true" pageId="1" pageNumber="256">T</emphasis>
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.
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<emphasis id="B97E837DEC01FFFCFAEBFF0A6C8DB06E" box="[1306,1425,1823,1846]" italics="true" pageId="1" pageNumber="256">yixianensis</emphasis>
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</taxonomicName>
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in smaller size of wing (
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<quantity id="4CF2F28AEC01FFFCFBE6FF546D65B001" box="[1047,1145,1857,1881]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="1" pageNumber="256" unit="mm" value="10.2">10.2 mm</quantity>
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<emphasis id="B97E837DEC01FFFCFB73FF576D84B001" box="[1154,1176,1858,1881]" italics="true" pageId="1" pageNumber="256">vs</emphasis>
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.
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<quantity id="4CF2F28AEC01FFFCFB56FF546C14B001" box="[1191,1288,1857,1881]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="1" pageNumber="256" unit="mm" value="13.3">13.3 mm</quantity>
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and>
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<quantity id="4CF2F28AEC01FFFCFAA1FF546C83B001" box="[1360,1439,1857,1881]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="256" unit="mm" value="18.0">18 mm</quantity>
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),
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</paragraph>
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<paragraph id="8BB55F6FEC01FFFCFCDBFF716D2BB0C7" blockId="1.[810,1452,1751,2092]" pageId="1" pageNumber="256">
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much shorter Sc
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<subScript id="178E5D2AEC01FFFCFC2AFF616AFFB0DA" attach="left" box="[987,995,1908,1922]" fontSize="6" pageId="1" pageNumber="256">1</subScript>
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, and longer Cu
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<subScript id="178E5D2AEC01FFFCFB7DFF616D88B0DA" attach="left" box="[1164,1172,1908,1922]" fontSize="6" pageId="1" pageNumber="256">1</subScript>
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, both reaching somewhat beyond wing mid-length.
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</paragraph>
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<paragraph id="8BB55F6FEC01FFFCFCAEFFBF6CB7B09B" blockId="1.[810,1452,1751,2092]" box="[863,1451,1962,1987]" pageId="1" pageNumber="256">
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<emphasis id="B97E837DEC01FFFCFCAEFFBF6D4CB09A" bold="true" box="[863,1104,1962,1987]" pageId="1" pageNumber="256">Locality and horizon.</emphasis>
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Smokejacks brickworks,
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<collectingRegion id="49CE918DEC01FFFCFAAEFFBE6CBBB09B" box="[1375,1447,1963,1987]" country="United Kingdom" name="Surrey" pageId="1" pageNumber="256">Surrey</collectingRegion>
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,
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</paragraph>
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<paragraph id="8BB55F6FEC01FFFCFCDBFFDB6CB0BF51" blockId="1.[810,1452,1751,2092]" pageId="1" pageNumber="256">
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<collectingCountry id="F31D1FFFEC01FFFCFCDBFFDB6A48B0BD" box="[810,852,1998,2021]" name="United Kingdom" pageId="1" pageNumber="256">UK</collectingCountry>
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;
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<bibRefCitation id="EF9B229EEC01FFFCFC95FFDB6D67B0BE" author="Austen, P. A. & Batten, D. J." box="[868,1147,1998,2022]" pageId="1" pageNumber="256" pagination="171 - 201" refId="ref2154" refString="Austen, P. A. & Batten, D. J. (2018) English Wealden fossils: an update. Proceedings of the Geologists' Association, 129 (2), 171 - 201. https: // doi. org / 10.1016 / j. pgeola. 2018.02.007" type="journal article" year="2018">Austen & Batten (2018)</bibRefCitation>
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bed 5 (lower), Bevhalstia and siltstone band, upper Weald Clay Formation, lower
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</paragraph>
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<paragraph id="8BB55F6FEC01FFFCFCDBF0016ABEBF74" blockId="1.[810,1452,1751,2092]" box="[810,930,2068,2092]" pageId="1" pageNumber="256">Barremian.</paragraph>
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</subSubSection>
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<caption id="DF750FE7EC02FFFFFF79FF8A6AA3B082" ID-DOI="http://doi.org/10.5281/zenodo.8073099" ID-Zenodo-Dep="8073099" httpUri="https://zenodo.org/record/8073099/files/figure.png" pageId="2" pageNumber="257" startId="2.[136,235,1951,1975]" targetBox="[174,1415,189,1924]" targetPageId="2" targetType="figure">
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<paragraph id="8BB55F6FEC02FFFFFF79FF8A6AA3B082" blockId="2.[136,1451,1951,2017]" pageId="2" pageNumber="257">
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<emphasis id="B97E837DEC02FFFFFF79FF8A681AB0EF" bold="true" box="[136,262,1951,1975]" pageId="2" pageNumber="257">FIGURE 2.</emphasis>
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?
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<taxonomicName id="4C0A24ECEC02FFFFFEEAFF8A68F8B0EF" authority="BASHKUEV & JARZEMBOWSKI, 2023" authorityName="BASHKUEV & JARZEMBOWSKI" authorityYear="2023" box="[283,484,1951,1975]" class="Insecta" family="Eomeropidae" genus="Typhothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="agari" status="sp. nov.">
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<emphasis id="B97E837DEC02FFFFFEEAFF8A68F8B0EF" box="[283,484,1951,1975]" italics="true" pageId="2" pageNumber="257">Typhothauma agari</emphasis>
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</taxonomicName>
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<emphasis id="B97E837DEC02FFFFFE1CFF8A6B47B0EF" bold="true" box="[493,603,1951,1975]" pageId="2" pageNumber="257">
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<taxonomicNameLabel id="A24D3E06EC02FFFFFE1CFF8A6B5CB0EF" box="[493,576,1951,1975]" pageId="2" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
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A
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</emphasis>
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,
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<emphasis id="B97E837DEC02FFFFFD9AFF8A6B60B0EF" bold="true" box="[619,636,1951,1975]" pageId="2" pageNumber="257">B</emphasis>
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, Photographs of the holotype in different optical and illumination settings.
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<emphasis id="B97E837DEC02FFFFFA62FF8A6CB9B0EF" bold="true" box="[1427,1445,1951,1975]" pageId="2" pageNumber="257">C</emphasis>
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, Venation interpretation, with the hypothetical position of Sc
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<subScript id="178E5D2AEC02FFFFFD16FFC66BF2B0B9" attach="right" box="[743,750,2003,2017]" fontSize="6" pageId="2" pageNumber="257">3</subScript>
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. Scale bars = 2 mm.
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</paragraph>
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</caption>
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<subSubSection id="C3100CE4EC03FFFEFF4CF88A6835B584" pageId="3" pageNumber="258" type="description">
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<paragraph id="8BB55F6FEC03FFFEFF4CF88A6A16B784" blockId="3.[136,778,159,732]" pageId="3" pageNumber="258">
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<emphasis id="B97E837DEC03FFFEFF4CF88A6854B7EF" bold="true" box="[189,328,159,183]" pageId="3" pageNumber="258">Description.</emphasis>
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Almost complete forewing with distal part of anterior margin missing,
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<quantity id="4CF2F28AEC03FFFEFE41F8D16B12B783" box="[432,526,196,220]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="258" unit="mm" value="10.2">10.2 mm</quantity>
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long and
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<quantity id="4CF2F28AEC03FFFEFD86F8D16BD4B783" box="[631,712,196,220]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="258" unit="mm" value="3.9">3.9 mm</quantity>
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wide,
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</paragraph>
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<paragraph id="8BB55F6FEC03FFFEFF79F8FD6A16B67D" blockId="3.[136,778,159,732]" pageId="3" pageNumber="258">length to width ratio 2.6. Venation relatively simplified as expected from rather small size of wing. Sc with two long,</paragraph>
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<paragraph id="8BB55F6FEC03FFFEFF79F92469B8B558" blockId="3.[136,778,159,732]" pageId="3" pageNumber="258">
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zigzagging longitudinal branches, cells between them and costal margin indiscernible. Sc
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<subScript id="178E5D2AEC03FFFEFE25F97368C0B62C" attach="left" box="[468,476,358,372]" fontSize="6" pageId="3" pageNumber="258">1</subScript>
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reaching somewhat beyond wing mid-length. R
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<subScript id="178E5D2AEC03FFFEFE97F99E6872B6C1" attach="left" box="[358,366,395,409]" fontSize="6" pageId="3" pageNumber="258">1</subScript>
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forking at pterostigma; pterostigma itself not preserved. Rs branching with at least six veins reaching wing margin, with apical forks on Rs
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<subScript id="178E5D2AEC03FFFEFD83F9C16B66B6BA" attach="left" box="[626,634,468,482]" fontSize="6" pageId="3" pageNumber="258">1</subScript>
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and either on Rs
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</paragraph>
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<paragraph id="8BB55F6FEC03FFFEFF55F9EC69B0B55F" blockId="3.[136,778,159,732]" box="[164,172,505,519]" pageId="3" pageNumber="258">3</paragraph>
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<paragraph id="8BB55F6FEC03FFFEFF42F9FC6810B558" blockId="3.[136,778,159,732]" box="[179,268,489,512]" pageId="3" pageNumber="258">or on Rs</paragraph>
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<paragraph id="8BB55F6FEC03FFFEFEFCF9EC6809B55F" blockId="3.[136,778,159,732]" box="[269,277,505,519]" pageId="3" pageNumber="258">4</paragraph>
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<paragraph id="8BB55F6FEC03FFFEFEE5F9FC6835B584" blockId="3.[136,778,159,732]" pageId="3" pageNumber="258">
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. Venation of M mostly obscure, but apparently with more branches than Rs. Cu
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<subScript id="178E5D2AEC03FFFEFE0DFA086B18B573" attach="left" box="[508,516,541,555]" fontSize="6" pageId="3" pageNumber="258">1</subScript>
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reaching beyond wing mid-length at the same level as Sc
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<subScript id="178E5D2AEC03FFFEFE09FA576B1CB508" attach="left" box="[504,512,578,592]" fontSize="6" pageId="3" pageNumber="258">1</subScript>
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. Space between Cu
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<subScript id="178E5D2AEC03FFFEFD25FA576BC0B508" attach="left" box="[724,732,578,592]" fontSize="6" pageId="3" pageNumber="258">1</subScript>
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and Cu
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<subScript id="178E5D2AEC03FFFEFF56FA7369B3B52C" attach="left" box="[167,175,614,628]" fontSize="6" pageId="3" pageNumber="258">2</subScript>
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with one row of five cells. Cu
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<subScript id="178E5D2AEC03FFFEFDF1FA736B14B52C" attach="left" box="[512,520,614,628]" fontSize="6" pageId="3" pageNumber="258">2</subScript>
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and anal veins in usual position, A
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<subScript id="178E5D2AEC03FFFEFF0DFA9E6818B5C1" attach="left" box="[252,260,651,665]" fontSize="6" pageId="3" pageNumber="258">3</subScript>
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forked. Cross-veins numerous, faintly preserved in distal part of wing, longitudinal veins mostly zigzagged by cross-veins.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |