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<document id="CF3B4E182878A1B6C7E8E37FDC7080F8" ID-DOI="10.11646/zootaxa.3990.4.3" ID-GBIF-Dataset="6c8c507c-e532-4603-957e-d28e2b97253a" ID-ISSN="1175-5326" ID-Zenodo-Dep="240688" ID-ZooBank="04B6F8A3-671D-4EE2-8271-2CA9E3BA333F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461270714269" checkinUser="plazi" docAuthor="Ferragut, Francisco &amp; Navia, Denise" docDate="2015" docId="039BEE1EFFEB8E2AFF6DFE66FBE1BBEA" docLanguage="en" docName="zt03990p550.pdf" docOrigin="Zootaxa 3990 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Typhlodromus (Anthoseius) anomalos Ferragut, n. sp." docType="treatment" docVersion="9" lastPageNumber="545" masterDocId="FFA29666FFF98E3EFFFAFF9FFFD2BC76" masterDocTitle="Phytoseiid mites (Acari: Phytoseiidae) from Patagonia and Tierra del Fuego" masterLastPageNumber="550" masterPageNumber="525" pageNumber="543" updateTime="1698611595168" updateUser="plazi">
<mods:mods id="5FFCFDAA8CE7B7345AD3D0755FC05C6A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C6E0E8DC9172C11F103F3E879B7D9703">
<mods:title id="6EB96F200FD1134A7578649FADD3B8DB">Phytoseiid mites (Acari: Phytoseiidae) from Patagonia and Tierra del Fuego</mods:title>
</mods:titleInfo>
<mods:name id="5C66CA1E85FB1008EF0062202624DA34" type="personal">
<mods:role id="8D5DF7BE803C5E9579950FF4D82F59CD">
<mods:roleTerm id="98E823E5A9C6411CA40E5E315D4ECB3F">Author</mods:roleTerm>
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<mods:namePart id="785C0F2A573DB15D43452B8894DC0191">Ferragut, Francisco</mods:namePart>
</mods:name>
<mods:name id="BE12B1CC93F5171BB8322A12BE27CA44" type="personal">
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<mods:roleTerm id="5E00AB85914CFB366C072237075707BB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5F9198841B3644F2B2010D4D553BBB55">Navia, Denise</mods:namePart>
</mods:name>
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<mods:title id="CCFAECC48959C55262DA26906CA4A115">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="56BEFFA47C3AB9673BCA13B0CCCFD53B">
<mods:date id="CE624AF97D8D4285C9552E71BF132CD2">2015</mods:date>
<mods:detail id="0D934D3DFAFB03329A1608BE7D11D6E0" type="volume">
<mods:number id="CB967BEA4B87198105797F25A6607C8F">3990</mods:number>
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<mods:detail id="418CE4C4C4D3374F0A8CBA1BC342760E" type="issue">
<mods:number id="40D488218A4AC09758988AE2993456E1">4</mods:number>
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<mods:classification id="4129CBFBC02276005E4A889FC8BAF272">journal article</mods:classification>
<mods:identifier id="03D11E595D863BCDCBDC93519B7EDB05" type="DOI">10.11646/zootaxa.3990.4.3</mods:identifier>
<mods:identifier id="D1323230BC279605B7D92DBCE6E9CA98" type="GBIF-Dataset">6c8c507c-e532-4603-957e-d28e2b97253a</mods:identifier>
<mods:identifier id="95EE0B8125FAD4C15AD2B401EBA20370" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3B605A2359D807D49B1D902C4337FEFC" type="Zenodo-Dep">240688</mods:identifier>
<mods:identifier id="773408E385DB4F3607E0C58BE89D32A6" type="ZooBank">04B6F8A3-671D-4EE2-8271-2CA9E3BA333F</mods:identifier>
</mods:mods>
<treatment id="039BEE1EFFEB8E2AFF6DFE66FBE1BBEA" ID-DOI="http://doi.org/10.5281/zenodo.5684476" ID-GBIF-Taxon="119638762" ID-Zenodo-Dep="5684476" LSID="urn:lsid:plazi:treatment:039BEE1EFFEB8E2AFF6DFE66FBE1BBEA" httpUri="http://treatment.plazi.org/id/039BEE1EFFEB8E2AFF6DFE66FBE1BBEA" lastPageId="20" lastPageNumber="545" pageId="18" pageNumber="543">
<subSubSection id="C3280C83FFEB8E2CFF6DFE66FE98BE43" pageId="18" pageNumber="543" type="nomenclature">
<paragraph id="8B8D5F08FFEB8E2CFF6DFE66FCF4BE65" blockId="18.[151,806,505,565]" box="[151,806,505,531]" pageId="18" pageNumber="543">
<heading id="D0C5E864FFEB8E2CFF6DFE66FCF4BE65" bold="true" box="[151,806,505,531]" fontSize="11" level="1" pageId="18" pageNumber="543" reason="1">
<emphasis id="B946831AFFEB8E2CFF6DFE66FCF4BE65" bold="true" box="[151,806,505,531]" pageId="18" pageNumber="543">
<taxonomicName id="4C32248BFFEB8E2CFF6DFE66FD0EBE65" authority="Ferragut" authorityName="Ferragut" box="[151,732,505,531]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="543" phylum="Arthropoda" rank="species" species="anomalos" status="sp. nov." subGenus="Anthoseius">
<emphasis id="B946831AFFEB8E2CFF6DFE66FE94BE65" bold="true" box="[151,326,505,531]" italics="true" pageId="18" pageNumber="543">Typhlodromus</emphasis>
(
<emphasis id="B946831AFFEB8E2CFEACFE66FE0FBE65" bold="true" box="[342,477,505,531]" italics="true" pageId="18" pageNumber="543">Anthoseius</emphasis>
)
<emphasis id="B946831AFFEB8E2CFE15FE66FDB6BE65" bold="true" box="[495,612,505,531]" italics="true" pageId="18" pageNumber="543">anomalos</emphasis>
Ferragut
</taxonomicName>
<taxonomicNameLabel id="A2753E61FFEB8E2CFD19FE66FCF4BE65" box="[739,806,505,531]" pageId="18" pageNumber="543" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF6DFD83FE98BE43" blockId="18.[151,806,505,565]" box="[151,330,540,565]" pageId="18" pageNumber="543">
(
<figureCitation id="1309438DFFEB8E2CFF64FD83FE90BE43" box="[158,322,540,565]" captionStart-0="FIGURES 46 – 49" captionStart-1="FIGURES 50 – 51" captionStartId-0="19.[151,264,1957,1980]" captionStartId-1="20.[151,264,1064,1087]" captionTargetBox-0="[213,1369,353,1924]" captionTargetBox-1="[323,1259,210,1005]" captionTargetId-0="figure@19.[212,1374,337,1936]" captionTargetId-1="figure@20.[306,1281,193,1043]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 46 – 49. Typhlodromus (Anthoseius) anomalos n. sp. female. 46. Dorsal shield, 47. Ventral surface, 48. Spermatheca, 49. Genu, tibia and tarsus of leg IV." captionText-1="FIGURES 50 – 51. Typhlodromus (Anthoseius) anomalos n. sp. male. 50. Dorsal shield, 51. Ventrianal shield." httpUri-0="https://zenodo.org/record/240700/files/figure.png" httpUri-1="https://zenodo.org/record/240701/files/figure.png" pageId="18" pageNumber="543">Figures 46–51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3280C83FFEB8E2CFF6DFDFBFE96BFB6" pageId="18" pageNumber="543" type="diagnosis">
<paragraph id="8B8D5F08FFEB8E2CFF6DFDFBFE96BFB6" blockId="18.[151,1437,612,2005]" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF6DFDFBFEC3BE0B" bold="true" box="[151,273,612,637]" pageId="18" pageNumber="543">Diagnosis.</emphasis>
Female dorsal shield elongate, striate and weakly reticulate. Dorsal setal pattern 12A:8A. Dorsal setae short, smooth and acute except for
<emphasis id="B946831AFFEB8E2CFDD3FD17FD95BEE9" box="[553,583,648,671]" italics="true" pageId="18" pageNumber="543">Z5</emphasis>
, thick and serrate. Position of seta
<emphasis id="B946831AFFEB8E2CFC21FD17FC25BED6" box="[987,1015,648,672]" italics="true" pageId="18" pageNumber="543">S4</emphasis>
variable in the
<typeStatus id="5489E1AAFFEB8E2CFB56FD18FAC3BED6" box="[1196,1297,647,672]" pageId="18" pageNumber="543" type="holotype">holotype</typeStatus>
; inserted on soft integument in the left side and on the dorsal shield in the right side. Six pairs of dorsal solenostomes (
<emphasis id="B946831AFFEB8E2CFAC9FD33FA8FBEB3" box="[1331,1373,684,709]" italics="true" pageId="18" pageNumber="543">gd1</emphasis>
,
<emphasis id="B946831AFFEB8E2CFA91FD33FA47BEB3" box="[1387,1429,684,709]" italics="true" pageId="18" pageNumber="543">gd2</emphasis>
,
<emphasis id="B946831AFFEB8E2CFF6DFD50FF13BE9E" box="[151,193,719,744]" italics="true" pageId="18" pageNumber="543">gd5</emphasis>
,
<emphasis id="B946831AFFEB8E2CFF2AFD50FF28BE9E" box="[208,250,719,744]" italics="true" pageId="18" pageNumber="543">gd6</emphasis>
,
<emphasis id="B946831AFFEB8E2CFEF2FD50FEE0BE9E" box="[264,306,719,744]" italics="true" pageId="18" pageNumber="543">gd8</emphasis>
,
<emphasis id="B946831AFFEB8E2CFEBBFD50FEB9BE9E" box="[321,363,719,744]" italics="true" pageId="18" pageNumber="543">gd9</emphasis>
). Peritremes almost reaching setae
<emphasis id="B946831AFFEB8E2CFD04FD4FFCC1BE9E" box="[766,787,720,744]" italics="true" pageId="18" pageNumber="543">j1</emphasis>
. Sternal shield with three pairs of setae, posterior margin not well discernible, apparently concave. Ventrianal shield smooth, with four pairs of pre-anal setae and without pores. Three pairs of setae around the ventrianal. Seta
<emphasis id="B946831AFFEB8E2CFD06FC87FCFABF46" box="[764,808,792,816]" italics="true" pageId="18" pageNumber="543">JV4</emphasis>
absent. Spermathecal calyx elongate, with a cylindrical base and a distal cup. Chelicerae apparently with two teeth on fixed digit, movable digit unidentate. Genu II with seven setae. One short and curved macroseta on tarsus IV. Ventrianal shield in the male oblong, reduced and with five pairs of pre-anal setae. Two large and rounded metapodal plates on the soft integument, near the ventrianal anterior angles.
</paragraph>
</subSubSection>
<subSubSection id="C3280C83FFEB8E2CFF3DFC53FC7ABBC6" pageId="18" pageNumber="543" type="description">
<paragraph id="8B8D5F08FFEB8E2CFF3DFC53FD2BBF93" blockId="18.[151,1437,612,2005]" box="[199,761,972,997]" pageId="18" pageNumber="543">
FEMALE (
<figureCitation id="1309438DFFEB8E2CFEBFFC53FE38BF93" box="[325,490,972,997]" captionStart="FIGURES 46 – 49" captionStartId="19.[151,264,1957,1980]" captionTargetBox="[213,1369,353,1924]" captionTargetId="figure@19.[212,1374,337,1936]" captionTargetPageId="19" captionText="FIGURES 46 – 49. Typhlodromus (Anthoseius) anomalos n. sp. female. 46. Dorsal shield, 47. Ventral surface, 48. Spermatheca, 49. Genu, tibia and tarsus of leg IV." httpUri="https://zenodo.org/record/240700/files/figure.png" pageId="18" pageNumber="543">Figures 46–49</figureCitation>
) (one female measured)
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF3DFC70FD86B8CA" blockId="18.[151,1437,612,2005]" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF3DFC70FEA1B87E" box="[199,371,1007,1032]" italics="true" pageId="18" pageNumber="543">Dorsal surface</emphasis>
. Dorsal shield elongate, 390 long, 190 wide, striated to reticulate. Eighteen pairs of thick, smooth and acute setae, except for
<emphasis id="B946831AFFEB8E2CFDE7FB8AFDE9B85A" box="[541,571,1045,1068]" italics="true" pageId="18" pageNumber="543">Z5</emphasis>
serrated. Setae
<emphasis id="B946831AFFEB8E2CFD11FB8BFCD6B85A" box="[747,772,1044,1068]" italics="true" pageId="18" pageNumber="543">r3</emphasis>
and
<emphasis id="B946831AFFEB8E2CFCC3FB8BFC8AB85A" box="[825,856,1044,1068]" italics="true" pageId="18" pageNumber="543">R1</emphasis>
on lateral integument; seta
<emphasis id="B946831AFFEB8E2CFB76FB8BFB7AB85A" box="[1164,1192,1044,1068]" italics="true" pageId="18" pageNumber="543">S4</emphasis>
on lateral integument in the left side of the mite. Seta
<emphasis id="B946831AFFEB8E2CFDFBFBA7FDC4B826" box="[513,534,1080,1104]" italics="true" pageId="18" pageNumber="543">j1</emphasis>
23,
<emphasis id="B946831AFFEB8E2CFDB3FBA7FD8DB826" box="[585,607,1080,1104]" italics="true" pageId="18" pageNumber="543">j3</emphasis>
22,
<emphasis id="B946831AFFEB8E2CFD6BFBA7FD75B826" box="[657,679,1080,1104]" italics="true" pageId="18" pageNumber="543">j4</emphasis>
14,
<emphasis id="B946831AFFEB8E2CFD23FBA7FD3DB839" box="[729,751,1080,1103]" italics="true" pageId="18" pageNumber="543">j5</emphasis>
13,
<emphasis id="B946831AFFEB8E2CFCDBFBA8FCE5B826" box="[801,823,1079,1104]" italics="true" pageId="18" pageNumber="543">j6</emphasis>
17,
<emphasis id="B946831AFFEB8E2CFC93FBA7FC51B826" box="[873,899,1080,1104]" italics="true" pageId="18" pageNumber="543">J2</emphasis>
19,
<emphasis id="B946831AFFEB8E2CFC4FFBA7FC02B839" box="[949,976,1080,1103]" italics="true" pageId="18" pageNumber="543">J5</emphasis>
15,
<emphasis id="B946831AFFEB8E2CFBF8FBA7FBC9B826" box="[1026,1051,1080,1104]" italics="true" pageId="18" pageNumber="543">z2</emphasis>
18,
<emphasis id="B946831AFFEB8E2CFBB7FBA7FBB5B826" box="[1101,1127,1080,1104]" italics="true" pageId="18" pageNumber="543">z3</emphasis>
18,
<emphasis id="B946831AFFEB8E2CFB62FBA7FB63B826" box="[1176,1201,1080,1104]" italics="true" pageId="18" pageNumber="543">z4</emphasis>
20,
<emphasis id="B946831AFFEB8E2CFB19FBA7FB2EB839" box="[1251,1276,1080,1103]" italics="true" pageId="18" pageNumber="543">z5</emphasis>
16,
<emphasis id="B946831AFFEB8E2CFAD4FBA7FA9EB826" box="[1326,1356,1080,1104]" italics="true" pageId="18" pageNumber="543">Z4</emphasis>
26,
<emphasis id="B946831AFFEB8E2CFA84FBA7FA4EB839" box="[1406,1436,1080,1103]" italics="true" pageId="18" pageNumber="543">Z5</emphasis>
46,
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<emphasis id="B946831AFFEB8E2CFF3AFBC3FF0BB802" box="[192,217,1116,1140]" italics="true" pageId="18" pageNumber="543">s4</emphasis>
23
</geoCoordinate>
,
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<emphasis id="B946831AFFEB8E2CFEF3FBC3FEF0B803" box="[265,290,1116,1141]" italics="true" pageId="18" pageNumber="543">s6</emphasis>
26
</geoCoordinate>
,
<geoCoordinate id="EE0639CFFFEB8E2CFEA8FBC3FE40B802" box="[338,402,1116,1140]" direction="south" orientation="latitude" pageId="18" pageNumber="543" precision="925" value="-2.4">
<emphasis id="B946831AFFEB8E2CFEA8FBC3FEBCB802" box="[338,366,1116,1140]" italics="true" pageId="18" pageNumber="543">S2</emphasis>
24
</geoCoordinate>
,
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<emphasis id="B946831AFFEB8E2CFE64FBC3FE68B802" box="[414,442,1116,1140]" italics="true" pageId="18" pageNumber="543">S4</emphasis>
22
</geoCoordinate>
,
<geoCoordinate id="EE0639CFFFEB8E2CFE10FBC2FDF8B802" box="[490,554,1116,1140]" direction="south" orientation="latitude" pageId="18" pageNumber="543" precision="925" value="-5.5">
<emphasis id="B946831AFFEB8E2CFE10FBC2FDD4B802" box="[490,518,1117,1140]" italics="true" pageId="18" pageNumber="543">S5</emphasis>
30
</geoCoordinate>
,
<emphasis id="B946831AFFEB8E2CFDCCFBC3FD9DB802" box="[566,591,1116,1140]" italics="true" pageId="18" pageNumber="543">r3</emphasis>
23,
<emphasis id="B946831AFFEB8E2CFD84FBC3FD4CB802" box="[638,670,1116,1140]" italics="true" pageId="18" pageNumber="543">R1</emphasis>
20. Six pairs of solenostomes, being
<emphasis id="B946831AFFEB8E2CFBC7FBC3FBB5B803" box="[1085,1127,1116,1141]" italics="true" pageId="18" pageNumber="543">gd4</emphasis>
absent;
<emphasis id="B946831AFFEB8E2CFB39FBC3FB3FB803" box="[1219,1261,1116,1141]" italics="true" pageId="18" pageNumber="543">gd1</emphasis>
anterolateral to
<emphasis id="B946831AFFEB8E2CFF6DFB1FFF7FB8EE" box="[151,173,1152,1176]" italics="true" pageId="18" pageNumber="543">j3</emphasis>
, distance
<emphasis id="B946831AFFEB8E2CFEDAFBE0FEBCB8EE" box="[288,366,1151,1176]" italics="true" pageId="18" pageNumber="543">gd1–j3</emphasis>
7;
<emphasis id="B946831AFFEB8E2CFE6FFBE0FE6DB8EE" box="[405,447,1151,1176]" italics="true" pageId="18" pageNumber="543">gd2</emphasis>
close and medial to
<emphasis id="B946831AFFEB8E2CFD54FB1FFD15B8EE" box="[686,711,1152,1176]" italics="true" pageId="18" pageNumber="543">z4</emphasis>
, distance
<emphasis id="B946831AFFEB8E2CFCC0FBE0FC59B8EE" box="[826,907,1151,1176]" italics="true" pageId="18" pageNumber="543">gd2–z4</emphasis>
14;
<emphasis id="B946831AFFEB8E2CFC3AFBE0FC3BB8EE" box="[960,1001,1151,1176]" italics="true" pageId="18" pageNumber="543">gd9</emphasis>
posteromedial to
<emphasis id="B946831AFFEB8E2CFB42FB1FFB06B8E1" box="[1208,1236,1152,1175]" italics="true" pageId="18" pageNumber="543">S5</emphasis>
, distance
<emphasis id="B946831AFFEB8E2CFAB2FBE0FA4EB8EE" box="[1352,1436,1151,1176]" italics="true" pageId="18" pageNumber="543">
gd9–
<geoCoordinate id="EE0639CFFFEB8E2CFA7AFBE0FA4EB8EE" box="[1408,1436,1151,1176]" direction="south" orientation="latitude" pageId="18" pageNumber="543" precision="925" value="-5.2">S5</geoCoordinate>
</emphasis>
12. Peritremes almost reaching setae
<emphasis id="B946831AFFEB8E2CFDC2FB3BFD9FB8CA" box="[568,589,1188,1212]" italics="true" pageId="18" pageNumber="543">j1</emphasis>
.
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF3DFB58FD57B9E3" blockId="18.[151,1437,612,2005]" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF3DFB58FEA2B896" box="[199,368,1223,1248]" italics="true" pageId="18" pageNumber="543">Ventral surface</emphasis>
. Sternal shield smooth and weakly sclerotised; posterior margin difficult to discern. Apparently with three pairs of setae;
<emphasis id="B946831AFFEB8E2CFE4DFB73FE0AB972" box="[439,472,1260,1284]" italics="true" pageId="18" pageNumber="543">st4</emphasis>
inserted on the tegument. Distance
<emphasis id="B946831AFFEB8E2CFC8BFB73FC13B972" box="[881,961,1260,1284]" italics="true" pageId="18" pageNumber="543">st1–st3</emphasis>
74, distance
<emphasis id="B946831AFFEB8E2CFBADFB73FB75B972" box="[1111,1191,1260,1284]" italics="true" pageId="18" pageNumber="543">st2–st2</emphasis>
64. Genital shield 66 wide. Smooth ventrianal shield 127 long, 99 wide at level of
<emphasis id="B946831AFFEB8E2CFC8AFA8FFC4DB95E" box="[880,927,1296,1320]" italics="true" pageId="18" pageNumber="543">ZV2</emphasis>
, 92 wide at level of anus. Anterior margin convex, laterals concave, two lateral areas of muscle attachments on the posterior margin, at the level of anus. Four pairs of pre-anal setae, very short, 7–9 long. Pre-anal pores absent. Three pairs of setae around the shield,
<emphasis id="B946831AFFEB8E2CFACBFAC7FAB2B906" box="[1329,1376,1368,1392]" italics="true" pageId="18" pageNumber="543">ZV1</emphasis>
,
<emphasis id="B946831AFFEB8E2CFA97FAC7FA4EB906" box="[1389,1436,1368,1392]" italics="true" pageId="18" pageNumber="543">ZV3</emphasis>
and
<emphasis id="B946831AFFEB8E2CFF3CFAE2FF20B9E2" box="[198,242,1405,1428]" italics="true" pageId="18" pageNumber="543">JV5</emphasis>
. Seta
<emphasis id="B946831AFFEB8E2CFECDFAE3FEB1B9E2" box="[311,355,1404,1428]" italics="true" pageId="18" pageNumber="543">JV4</emphasis>
absent. Seta
<emphasis id="B946831AFFEB8E2CFE0CFAE2FDF0B9E2" box="[502,546,1405,1428]" italics="true" pageId="18" pageNumber="543">JV5</emphasis>
34 long.
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF3DFA00FEC1B9AB" blockId="18.[151,1437,612,2005]" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF3DFA00FE90B9CE" box="[199,322,1439,1464]" italics="true" pageId="18" pageNumber="543">Chelicerae</emphasis>
. Not well discernible due to their position. Apparently, fixed digit with two teeth, movable digit unidentate.
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF3DFA76FD1FBA53" blockId="18.[151,1437,612,2005]" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF3DFA76FE00BA76" box="[199,466,1513,1536]" italics="true" pageId="18" pageNumber="543">Insemination apparatus</emphasis>
. Major duct broad, atrium not prominent. Calyx 22 long, with distal cup 12 long and a proximal narrow and cylindrical structure 10 long.
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF3DF9AEFA8BBA3E" blockId="18.[151,1437,612,2005]" box="[199,1369,1583,1608]" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF3DF9AEFF2EBA3E" box="[199,252,1585,1608]" italics="true" pageId="18" pageNumber="543">Legs</emphasis>
. Genu II with seven setae; 2–2/0, 2/0–1. One short and curved macroseta on
<emphasis id="B946831AFFEB8E2CFBA6F9B0FB1DBA3E" box="[1116,1231,1583,1608]" italics="true" pageId="18" pageNumber="543">basitarsus</emphasis>
IV, 26 long.
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF3DF9CBFD19BA1B" blockId="18.[151,1437,612,2005]" box="[199,715,1620,1645]" pageId="18" pageNumber="543">
MALE (
<figureCitation id="1309438DFFEB8E2CFEDFF9CBFE1BBA1B" box="[293,457,1620,1645]" captionStart="FIGURES 50 – 51" captionStartId="20.[151,264,1064,1087]" captionTargetBox="[323,1259,210,1005]" captionTargetId="figure@20.[306,1281,193,1043]" captionTargetPageId="20" captionText="FIGURES 50 – 51. Typhlodromus (Anthoseius) anomalos n. sp. male. 50. Dorsal shield, 51. Ventrianal shield." httpUri="https://zenodo.org/record/240701/files/figure.png" pageId="18" pageNumber="543">Figures 50–51</figureCitation>
) (one male measured):
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF3DF9E8FD90BA8A" blockId="18.[151,1437,612,2005]" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF3DF9E8FEBCBAE6" box="[199,366,1655,1680]" italics="true" pageId="18" pageNumber="543">Dorsal surface</emphasis>
. Dorsal shield 330 long, 176 wide, anteriorly striated and reticulate posteriorly. Twenty pairs of smooth and acute setae, being
<emphasis id="B946831AFFEB8E2CFE16F902FDDBBAC2" box="[492,521,1693,1716]" italics="true" pageId="18" pageNumber="543">Z5</emphasis>
lightly serrated. Seta
<emphasis id="B946831AFFEB8E2CFD07F903FCC1BAC2" box="[765,787,1692,1716]" italics="true" pageId="18" pageNumber="543">j1</emphasis>
22,
<emphasis id="B946831AFFEB8E2CFCB9F903FC8BBAC2" box="[835,857,1692,1716]" italics="true" pageId="18" pageNumber="543">j3</emphasis>
18,
<emphasis id="B946831AFFEB8E2CFC70F903FC72BAC2" box="[906,928,1692,1716]" italics="true" pageId="18" pageNumber="543">j4</emphasis>
13,
<emphasis id="B946831AFFEB8E2CFC2AF902FC34BAC2" box="[976,998,1693,1716]" italics="true" pageId="18" pageNumber="543">j5</emphasis>
11,
<emphasis id="B946831AFFEB8E2CFBEFF903FBF9BAC3" box="[1045,1067,1692,1717]" italics="true" pageId="18" pageNumber="543">j6</emphasis>
15,
<emphasis id="B946831AFFEB8E2CFBA6F903FBA4BAC2" box="[1116,1142,1692,1716]" italics="true" pageId="18" pageNumber="543">J2</emphasis>
14,
<emphasis id="B946831AFFEB8E2CFB5CF902FB13BAC2" box="[1190,1217,1693,1716]" italics="true" pageId="18" pageNumber="543">J5</emphasis>
11,
<emphasis id="B946831AFFEB8E2CFB0AF903FADBBAC2" box="[1264,1289,1692,1716]" italics="true" pageId="18" pageNumber="543">z2</emphasis>
15,
<emphasis id="B946831AFFEB8E2CFAC0F903FA81BAC2" box="[1338,1363,1692,1716]" italics="true" pageId="18" pageNumber="543">z3</emphasis>
12,
<emphasis id="B946831AFFEB8E2CFA79F903FA4EBAC2" box="[1411,1436,1692,1716]" italics="true" pageId="18" pageNumber="543">z4</emphasis>
14,
<emphasis id="B946831AFFEB8E2CFF3EF95FFF0FBAA1" box="[196,221,1728,1751]" italics="true" pageId="18" pageNumber="543">z5</emphasis>
12,
<emphasis id="B946831AFFEB8E2CFEE9F95FFEE2BAAE" box="[275,304,1728,1752]" italics="true" pageId="18" pageNumber="543">Z4</emphasis>
23,
<emphasis id="B946831AFFEB8E2CFE9DF95FFE57BAA1" box="[359,389,1728,1751]" italics="true" pageId="18" pageNumber="543">Z5</emphasis>
33,
<geoCoordinate id="EE0639CFFFEB8E2CFE41F95FFE29BAAE" box="[443,507,1728,1752]" direction="south" orientation="latitude" pageId="18" pageNumber="543" precision="925" value="-4.3333335">
<emphasis id="B946831AFFEB8E2CFE41F95FFE01BAAE" box="[443,467,1728,1752]" italics="true" pageId="18" pageNumber="543">s4</emphasis>
20
</geoCoordinate>
,
<geoCoordinate id="EE0639CFFFEB8E2CFDF0F95FFD9CBAAE" box="[522,590,1728,1752]" direction="south" orientation="latitude" pageId="18" pageNumber="543" precision="925" value="-2.3333333">
<emphasis id="B946831AFFEB8E2CFDF0F95FFDF4BAAE" box="[522,550,1728,1752]" italics="true" pageId="18" pageNumber="543">S2</emphasis>
20
</geoCoordinate>
,
<geoCoordinate id="EE0639CFFFEB8E2CFDA6F95FFD73BAAE" box="[604,673,1728,1752]" direction="south" orientation="latitude" pageId="18" pageNumber="543" precision="925" value="-4.3333335">
<emphasis id="B946831AFFEB8E2CFDA6F95FFDABBAAE" box="[604,633,1728,1752]" italics="true" pageId="18" pageNumber="543">S4</emphasis>
20
</geoCoordinate>
,
<geoCoordinate id="EE0639CFFFEB8E2CFD55F95FFD21BAAE" box="[687,755,1728,1752]" direction="south" orientation="latitude" pageId="18" pageNumber="543" precision="925" value="-5.366667">
<emphasis id="B946831AFFEB8E2CFD55F95FFD19BAA1" box="[687,715,1728,1751]" italics="true" pageId="18" pageNumber="543">S5</emphasis>
22
</geoCoordinate>
,
<emphasis id="B946831AFFEB8E2CFCFBF95FFCC9BAAE" box="[769,795,1728,1752]" italics="true" pageId="18" pageNumber="543">r3</emphasis>
19,
<emphasis id="B946831AFFEB8E2CFCABF95FFCA2BAAE" box="[849,880,1728,1752]" italics="true" pageId="18" pageNumber="543">R1</emphasis>
17. Six pairs of solenostomes, with
<emphasis id="B946831AFFEB8E2CFAE0F920FA96BAAE" box="[1306,1348,1727,1752]" italics="true" pageId="18" pageNumber="543">gd4</emphasis>
absent. Peritremes extend to bases of setae
<emphasis id="B946831AFFEB8E2CFDDFF97BFDE9BA8A" box="[549,571,1764,1788]" italics="true" pageId="18" pageNumber="543">j3</emphasis>
.
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF3DF898FA49BB1E" blockId="18.[151,1437,612,2005]" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF3DF898FEA3BB56" box="[199,369,1799,1824]" italics="true" pageId="18" pageNumber="543">Ventral surface</emphasis>
. Ventrianal shield oblong in shape and reduced, showing two well-developed metapodal plates at the level of anterior corners, 16–18 long, 8–10 wide. Anterior margin straight, laterals strongly convex; length 120, width at level of
<emphasis id="B946831AFFEB8E2CFE76F8CFFE69BB1E" box="[396,443,1872,1896]" italics="true" pageId="18" pageNumber="543">ZV2</emphasis>
115, at level of anus 88. Five pairs of pre-anal setae; pre-anal pores absent. Seta
<emphasis id="B946831AFFEB8E2CFABFF8CFFAA3BB11" box="[1349,1393,1872,1895]" italics="true" pageId="18" pageNumber="543">JV5</emphasis>
24.
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF3DF8EBFAFDBBFB" blockId="18.[151,1437,612,2005]" box="[199,1327,1908,1933]" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF3DF8EBFE90BBFB" box="[199,322,1908,1933]" italics="true" pageId="18" pageNumber="543">Chelicerae</emphasis>
. Inner margin not discernible. Spermatodactyl L-shaped, with a small anterior projection.
</paragraph>
<paragraph id="8B8D5F08FFEB8E2CFF3DF806FC7ABBC6" blockId="18.[151,1437,612,2005]" box="[199,936,1943,1968]" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF3DF806FF2EBBC6" box="[199,252,1945,1968]" italics="true" pageId="18" pageNumber="543">Legs</emphasis>
. Macroseta on leg IV (
<emphasis id="B946831AFFEB8E2CFE05F808FDA0BBC6" box="[511,626,1943,1968]" italics="true" pageId="18" pageNumber="543">basitarsus</emphasis>
) short and curved, 24 long.
</paragraph>
</subSubSection>
<subSubSection id="C3280C83FFEB8E2DFF3DF823FCE5BD6B" lastPageId="19" lastPageNumber="544" pageId="18" pageNumber="543" type="materials_examined">
<paragraph id="8B8D5F08FFEB8E2DFF3DF823FCE5BD6B" blockId="18.[151,1437,612,2005]" lastBlockId="19.[151,1437,151,285]" lastPageId="19" lastPageNumber="544" pageId="18" pageNumber="543">
<emphasis id="B946831AFFEB8E2CFF3DF823FEA8BBA3" bold="true" box="[199,378,1980,2005]" pageId="18" pageNumber="543">
<typeStatus id="5489E1AAFFEB8E2CFF3DF823FED0BBA3" box="[199,258,1980,2005]" pageId="18" pageNumber="543">Type</typeStatus>
material.
</emphasis>
One female
<typeStatus id="5489E1AAFFEB8E2CFDE6F823FDACBBA3" box="[540,638,1980,2005]" pageId="18" pageNumber="543" type="holotype">holotype</typeStatus>
and one male
<typeStatus id="5489E1AAFFEB8E2CFCC3F822FC4BBBA3" box="[825,921,1981,2005]" pageId="18" pageNumber="543" type="paratype">paratype</typeStatus>
on
<taxonomicName id="4C32248BFFEB8E2DFC35F823FE9CBCC6" authority="(Molina) K. Koch" authorityName="(Molina) K. Koch" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" lastPageId="19" lastPageNumber="544" order="Pinales" pageId="18" pageNumber="543" phylum="Tracheophyta" rank="species" species="araucana">
<emphasis id="B946831AFFEB8E2CFC35F823FB68BBA2" box="[975,1210,1980,2004]" italics="true" pageId="18" pageNumber="543">Araucaria araucana</emphasis>
(Molina) K. Koch (Araucariaceae)
</taxonomicName>
. Mamuil-Malal Pass, 65 Km from Junín de los Andes (
<collectingCountry id="F3251F98FFEA8E2DFC15FF08FBB6BCC6" box="[1007,1124,151,176]" name="Argentina" pageId="19" pageNumber="544">Argentina</collectingCountry>
), near the border between
<collectingCountry id="F3251F98FFEA8E2DFF6DFF23FED4BCA3" box="[151,262,188,213]" name="Argentina" pageId="19" pageNumber="544">Argentina</collectingCountry>
and
<collectingCountry id="F3251F98FFEA8E2DFEC7FF23FEAFBCA3" box="[317,381,188,213]" name="Chile" pageId="19" pageNumber="544">Chile</collectingCountry>
; 39º 34'56&quot; S, 71º 27'44&quot; W;
<quantity id="4CCAF2EDFFEA8E2DFD3DFF23FCCEBCA2" box="[711,796,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="19" pageNumber="544" unit="m" value="1210.0">1210 m</quantity>
asl.
<date id="FF8C79C8FFEA8E2DFCABFF22FBD3BCA2" box="[849,1025,188,213]" pageId="19" pageNumber="544" value="2013-01-07">7 January 2013</date>
.
<typeStatus id="5489E1AAFFEA8E2DFBF1FF23FBA1BCA3" box="[1035,1139,188,213]" pageId="19" pageNumber="544" type="holotype">Holotype</typeStatus>
female and male
<typeStatus id="5489E1AAFFEA8E2DFAC6FF22FA4EBCA3" box="[1340,1436,189,213]" pageId="19" pageNumber="544" type="paratype">paratype</typeStatus>
were deposited at Museo Nacional de Ciencias Naturales, Madrid,
<collectingCountry id="F3251F98FFEA8E2DFC7EFF40FC16BC8E" box="[900,964,223,248]" name="Spain" pageId="19" pageNumber="544">Spain</collectingCountry>
(
<collectionCode id="ED23C7CDFFEA8E2DFC28FF7FFBFEBC8E" LSID="urn:lsid:biocol.org:col:35275" box="[978,1068,224,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="19" pageNumber="544">MNCN</collectionCode>
). Reference number of
<typeStatus id="5489E1AAFFEA8E2DFAC9FF40FA4BBC8E" box="[1331,1433,223,248]" pageId="19" pageNumber="544" type="holotype">holotype</typeStatus>
:
<collectionCode id="ED23C7CDFFEA8E2DFF6DFE9BFF39BD6A" LSID="urn:lsid:biocol.org:col:35275" box="[151,235,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="19" pageNumber="544">MNCN</collectionCode>
20.02/17395;
<specimenCount id="9D349481FFEA8E2DFE74FE9BFE0ABD6B" box="[398,472,260,285]" pageId="19" pageNumber="544" type="male">1 male</specimenCount>
<typeStatus id="5489E1AAFFEA8E2DFE1AFE9AFD92BD6B" box="[480,576,261,285]" pageId="19" pageNumber="544" type="paratype">paratype</typeStatus>
<collectionCode id="ED23C7CDFFEA8E2DFDBDFE9BFD49BD6A" LSID="urn:lsid:biocol.org:col:35275" box="[583,667,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="19" pageNumber="544">MNCN</collectionCode>
20.02/17396.
</paragraph>
</subSubSection>
<caption id="DF4D0F80FFEA8E2DFF6DF83AFE2DBBAC" httpUri="https://zenodo.org/record/240700/files/figure.png" pageId="19" pageNumber="544" targetBox="[213,1369,353,1924]" targetPageId="19">
<paragraph id="8B8D5F08FFEA8E2DFF6DF83AFE2DBBAC" blockId="19.[151,1436,1957,2010]" pageId="19" pageNumber="544">
<emphasis id="B946831AFFEA8E2DFF6DF83AFE81BBCD" bold="true" box="[151,339,1957,1980]" pageId="19" pageNumber="544">FIGURES 46–49.</emphasis>
<taxonomicName id="4C32248BFFEA8E2DFEA3F839FD05BBCA" box="[345,727,1958,1980]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="544" phylum="Arthropoda" rank="species" species="anomalos" status="sp. nov." subGenus="Anthoseius">
<emphasis id="B946831AFFEA8E2DFEA3F839FE3ABBCA" box="[345,488,1958,1980]" italics="true" pageId="19" pageNumber="544">Typhlodromus</emphasis>
(
<emphasis id="B946831AFFEA8E2DFE0CF839FDB7BBCA" box="[502,613,1958,1980]" italics="true" pageId="19" pageNumber="544">Anthoseius</emphasis>
)
<emphasis id="B946831AFFEA8E2DFD8EF839FD05BBCA" box="[628,727,1958,1980]" italics="true" pageId="19" pageNumber="544">anomalos</emphasis>
</taxonomicName>
<emphasis id="B946831AFFEA8E2DFD26F839FCC7BBCD" bold="true" box="[732,789,1958,1979]" pageId="19" pageNumber="544">
<taxonomicNameLabel id="A2753E61FFEA8E2DFD26F839FCC7BBCD" box="[732,789,1958,1979]" pageId="19" pageNumber="544" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
female. 46. Dorsal shield, 47. Ventral surface, 48. Spermatheca, 49. Genu, tibia and tarsus of leg IV.
</paragraph>
</caption>
<caption id="DF4D0F80FFED8E2AFF6DFBB7FB3EB848" box="[151,1260,1064,1087]" httpUri="https://zenodo.org/record/240701/files/figure.png" pageId="20" pageNumber="545" targetBox="[323,1259,210,1005]" targetPageId="20">
<paragraph id="8B8D5F08FFED8E2AFF6DFBB7FB3EB848" blockId="20.[151,1260,1064,1087]" box="[151,1260,1064,1087]" pageId="20" pageNumber="545">
<emphasis id="B946831AFFED8E2AFF6DFBB7FE86B84B" bold="true" box="[151,340,1064,1087]" pageId="20" pageNumber="545">FIGURES 50–51.</emphasis>
<taxonomicName id="4C32248BFFED8E2AFEA0FBB7FD0AB848" box="[346,728,1064,1086]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="anomalos" status="sp. nov." subGenus="Anthoseius">
<emphasis id="B946831AFFED8E2AFEA0FBB7FE3BB848" box="[346,489,1064,1086]" italics="true" pageId="20" pageNumber="545">Typhlodromus</emphasis>
(
<emphasis id="B946831AFFED8E2AFE02FBB7FDB5B848" box="[504,615,1064,1086]" italics="true" pageId="20" pageNumber="545">Anthoseius</emphasis>
)
<emphasis id="B946831AFFED8E2AFD8FFBB7FD0AB848" box="[629,728,1064,1086]" italics="true" pageId="20" pageNumber="545">anomalos</emphasis>
</taxonomicName>
<emphasis id="B946831AFFED8E2AFD24FBB6FCC5B848" bold="true" box="[734,791,1065,1086]" pageId="20" pageNumber="545">
<taxonomicNameLabel id="A2753E61FFED8E2AFD24FBB6FCC5B848" box="[734,791,1065,1086]" pageId="20" pageNumber="545" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
male. 50. Dorsal shield, 51. Ventrianal shield.
</paragraph>
</caption>
<subSubSection id="C3280C83FFED8E2AFF3DFBF4FC9FB8DE" pageId="20" pageNumber="545" type="etymology">
<paragraph id="8B8D5F08FFED8E2AFF3DFBF4FC9FB8DE" blockId="20.[151,1437,1131,1948]" pageId="20" pageNumber="545">
<emphasis id="B946831AFFED8E2AFF3DFBF4FE9CB8F2" bold="true" box="[199,334,1131,1156]" pageId="20" pageNumber="545">Etymology.</emphasis>
The Greek word
<taxonomicName id="4C32248BFFED8E2AFDC4FBF4FD79B8F2" box="[574,683,1131,1156]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="anomalos">
<emphasis id="B946831AFFED8E2AFDC4FBF4FD79B8F2" box="[574,683,1131,1156]" italics="true" pageId="20" pageNumber="545">anomalos</emphasis>
</taxonomicName>
means &quot;unusual&quot; and &quot;different&quot; and refers to the freakish morphological features in the female and male of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3280C83FFED8E2AFF3DFB2CFBE1BBEA" pageId="20" pageNumber="545" type="discussion">
<paragraph id="8B8D5F08FFED8E2AFF3DFB2CFD0EB9D2" blockId="20.[151,1437,1131,1948]" pageId="20" pageNumber="545">
<emphasis id="B946831AFFED8E2AFF3DFB2CFE9DB8BA" bold="true" box="[199,335,1203,1228]" pageId="20" pageNumber="545">Comments.</emphasis>
<taxonomicName id="4C32248BFFED8E2AFE99FB2CFCC2B8BA" box="[355,784,1203,1228]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="anomalos" status="sp. nov." subGenus="Anthoseius">
<emphasis id="B946831AFFED8E2AFE99FB2CFDD0B8BA" box="[355,514,1203,1228]" italics="true" pageId="20" pageNumber="545">Typhlodromus</emphasis>
(
<emphasis id="B946831AFFED8E2AFDEFFB2CFD42B8BA" box="[533,656,1203,1228]" italics="true" pageId="20" pageNumber="545">Anthoseius</emphasis>
)
<emphasis id="B946831AFFED8E2AFD59FB2CFCC2B8BA" box="[675,784,1203,1228]" italics="true" pageId="20" pageNumber="545">anomalos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2753E61FFED8E2AFCE0FB2BFCA6B8BD" box="[794,884,1204,1227]" pageId="20" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
is the unique species in the genus with seta
<emphasis id="B946831AFFED8E2AFA7AFB2BFA4EB8BA" box="[1408,1436,1204,1228]" italics="true" pageId="20" pageNumber="545">S4</emphasis>
present and
<emphasis id="B946831AFFED8E2AFEE7FB48FE98B899" box="[285,330,1239,1263]" italics="true" pageId="20" pageNumber="545">JV4</emphasis>
absent, and it does not fit with the species groups created by
<bibRefCitation id="EFA322F9FFED8E2AFC0FFB48FAF6B899" author="Chant" box="[1013,1316,1239,1264]" pageId="20" pageNumber="545" refString="Chant, D. A. &amp; McMurtry, J. A. (1994) A review of the subfamilies Phytoseiinae and Typhlodrominae (Acari: Phytoseiidae). International Journal of Acarology, 20, 223 - 310. http: // dx. doi. org / 10.1080 / 01647959408684022" type="journal article" year="1994">Chant &amp; McMurtry (1994)</bibRefCitation>
. Seta
<emphasis id="B946831AFFED8E2AFA9CFB48FA51B899" box="[1382,1411,1239,1263]" italics="true" pageId="20" pageNumber="545">S4</emphasis>
is inserted well ahead of seta
<emphasis id="B946831AFFED8E2AFE35FB63FE3FB962" box="[463,493,1276,1300]" italics="true" pageId="20" pageNumber="545">Z4</emphasis>
and much closer to
<emphasis id="B946831AFFED8E2AFD2CFB63FD20B962" box="[726,754,1276,1300]" italics="true" pageId="20" pageNumber="545">S2</emphasis>
than to
<emphasis id="B946831AFFED8E2AFCA8FB63FCBCB965" box="[850,878,1276,1299]" italics="true" pageId="20" pageNumber="545">S5</emphasis>
. Because this species is represented only by two adults we do not know if this seta is found in the same position in immature stages. Despite of its position we consider this seta
<emphasis id="B946831AFFED8E2AFE82FADBFE46B92A" box="[376,404,1348,1372]" italics="true" pageId="20" pageNumber="545">S4</emphasis>
because
<emphasis id="B946831AFFED8E2AFDF1FADBFDF5B92A" box="[523,551,1348,1372]" italics="true" pageId="20" pageNumber="545">S3</emphasis>
, present in the families
<taxonomicName id="4C32248BFFED8E2AFCA4FADCFC6AB92A" box="[862,952,1347,1372]" class="Arachnida" family="Ascidae" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="family">Ascidae</taxonomicName>
and Blattisocidae, is suppressed in the
<taxonomicName id="4C32248BFFED8E2AFF6DFAF8FDB3B909" authority="Chant &amp; McMurtry, 2007" authorityName="Chant &amp; McMurtry" authorityYear="2007" box="[151,609,1383,1408]" class="Arachnida" family="Phytoseiidae" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="family">
Phytoseiidae (
<bibRefCitation id="EFA322F9FFED8E2AFECEFAF8FD8BB909" author="Chant" box="[308,601,1383,1408]" pageId="20" pageNumber="545" refString="Chant, D. A. &amp; McMurtry, J. A. (2007) Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata). Indira Publishing House, Michigan, 220 pp." type="book" year="2007">Chant &amp; McMurtry, 2007</bibRefCitation>
)
</taxonomicName>
. On the right side of
<typeStatus id="5489E1AAFFED8E2AFCB0FAF8FC7EB9F6" box="[842,940,1383,1408]" pageId="20" pageNumber="545" type="holotype">holotype</typeStatus>
<emphasis id="B946831AFFED8E2AFC49FAF8FC1DB909" box="[947,975,1383,1407]" italics="true" pageId="20" pageNumber="545">S4</emphasis>
is inserted on the dorsal shield; however, on the left side is inserted on the lateral integument.
</paragraph>
<paragraph id="8B8D5F08FFED8E2AFF3DFA30FEF0BB7D" blockId="20.[151,1437,1131,1948]" pageId="20" pageNumber="545">
The new species shares the absence of seta
<emphasis id="B946831AFFED8E2AFD4AFA30FD0EB9B1" box="[688,732,1455,1479]" italics="true" pageId="20" pageNumber="545">JV4</emphasis>
with
<taxonomicName id="4C32248BFFED8E2AFCE6FA30FA4BB9B1" authority="Tuttle &amp; Muma" authorityName="Tuttle &amp; Muma" box="[796,1433,1455,1480]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="arizonicus" subGenus="Anthoseius">
<emphasis id="B946831AFFED8E2AFCE6FA30FC68B9BE" box="[796,954,1455,1480]" italics="true" pageId="20" pageNumber="545">Typhlodromus</emphasis>
(
<emphasis id="B946831AFFED8E2AFC31FA30FB94B9BE" box="[971,1094,1455,1480]" italics="true" pageId="20" pageNumber="545">Anthoseius</emphasis>
)
<emphasis id="B946831AFFED8E2AFBADFA2FFB1EB9B1" box="[1111,1228,1456,1479]" italics="true" pageId="20" pageNumber="545">arizonicus</emphasis>
(Tuttle &amp; Muma)
</taxonomicName>
. This apomorphy is shared with species in the tribes Typhloseiopsini and Metaseiulini and it was considered as a secondary adaptation by
<bibRefCitation id="EFA322F9FFED8E2AFE55FA68FD32BA79" author="Chant" box="[431,736,1527,1552]" pageId="20" pageNumber="545" refString="Chant, D. A. &amp; McMurtry, J. A. (1994) A review of the subfamilies Phytoseiinae and Typhlodrominae (Acari: Phytoseiidae). International Journal of Acarology, 20, 223 - 310. http: // dx. doi. org / 10.1080 / 01647959408684022" type="journal article" year="1994">Chant &amp; McMurtry (1994)</bibRefCitation>
. Based on this atypical setal pattern
<bibRefCitation id="EFA322F9FFED8E2AFB80FA68FA4EBA79" author="Denmark" box="[1146,1436,1527,1552]" pageId="20" pageNumber="545" refString="Denmark, H. A. &amp; Evans, G. A. (2011) Phytoseiidae of North America and Hawaii (Acari: Mesostigmata). Indira Publishing House, West Bloomfield, Michigan, 451 pp." type="book" year="2011">
<collectingCountry id="F3251F98FFED8E2AFB80FA68FB30BA66" box="[1146,1250,1527,1552]" name="Denmark" pageId="20" pageNumber="545">Denmark</collectingCountry>
&amp; Evans (2011)
</bibRefCitation>
placed
<taxonomicName id="4C32248BFFED8E2AFF1DF983FE79BA45" box="[231,427,1564,1588]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="arizonicus" subGenus="Anthoseius">
<emphasis id="B946831AFFED8E2AFF1DF983FF26BA45" box="[231,244,1564,1587]" italics="true" pageId="20" pageNumber="545">T</emphasis>
. (
<emphasis id="B946831AFFED8E2AFEF7F983FECCBA45" box="[269,286,1564,1587]" italics="true" pageId="20" pageNumber="545">A</emphasis>
.)
<emphasis id="B946831AFFED8E2AFECCF983FE79BA45" box="[310,427,1564,1587]" italics="true" pageId="20" pageNumber="545">arizonicus</emphasis>
</taxonomicName>
in the genus
<taxonomicName id="4C32248BFFED8E2AFDB8F983FD78BA45" box="[578,682,1564,1587]" class="Arachnida" family="Phytoseiidae" genus="Meyerius" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="genus">
<emphasis id="B946831AFFED8E2AFDB8F983FD78BA45" box="[578,682,1564,1587]" italics="true" pageId="20" pageNumber="545">Meyerius</emphasis>
</taxonomicName>
and described
<taxonomicName id="4C32248BFFED8E2AFCAEF983FBEEBA42" box="[852,1084,1563,1588]" class="Arachnida" family="Phytoseiidae" genus="Meyerius" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="waltheriae">
<emphasis id="B946831AFFED8E2AFCAEF983FBEEBA42" box="[852,1084,1563,1588]" italics="true" pageId="20" pageNumber="545">Meyerius waltheriae</emphasis>
</taxonomicName>
<collectingCountry id="F3251F98FFED8E2AFBBEF984FB7EBA42" box="[1092,1196,1563,1588]" name="Denmark" pageId="20" pageNumber="545">Denmark</collectingCountry>
&amp; Evans as a related species. It is interesting to note that seta
<emphasis id="B946831AFFED8E2AFDA1F9A0FDA5BA21" box="[603,631,1599,1623]" italics="true" pageId="20" pageNumber="545">S2</emphasis>
is located in its normal position (more or less aligned with
<emphasis id="B946831AFFED8E2AFAF4F9A0FAFABA21" box="[1294,1320,1599,1623]" italics="true" pageId="20" pageNumber="545">J2</emphasis>
) in
<taxonomicName id="4C32248BFFED8E2AFAACF9DFFEDEBA0D" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="arizonicus" subGenus="Anthoseius">
<emphasis id="B946831AFFED8E2AFAACF9DFFAB1BA21" box="[1366,1379,1600,1623]" italics="true" pageId="20" pageNumber="545">T</emphasis>
. (
<emphasis id="B946831AFFED8E2AFA80F9DFFA59BA21" box="[1402,1419,1600,1623]" italics="true" pageId="20" pageNumber="545">A</emphasis>
.)
<emphasis id="B946831AFFED8E2AFF6DF9FBFEDEBA0D" box="[151,268,1636,1659]" italics="true" pageId="20" pageNumber="545">arizonicus</emphasis>
</taxonomicName>
but distinctly displaced in a posterior position (anterolaterad to solenostome
<emphasis id="B946831AFFED8E2AFB97F9FCFB45BA0A" box="[1133,1175,1635,1660]" italics="true" pageId="20" pageNumber="545">gd8</emphasis>
) in
<taxonomicName id="4C32248BFFED8E2AFB3FF9FBFAB0BA0A" box="[1221,1378,1635,1660]" class="Arachnida" family="Phytoseiidae" genus="Meyerius" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="waltheriae">
<emphasis id="B946831AFFED8E2AFB3FF9FBFB0EBA0D" box="[1221,1244,1636,1659]" italics="true" pageId="20" pageNumber="545">M</emphasis>
.
<emphasis id="B946831AFFED8E2AFB10F9FCFAB0BA0A" box="[1258,1378,1635,1660]" italics="true" pageId="20" pageNumber="545">waltheriae</emphasis>
</taxonomicName>
. The insertion of
<emphasis id="B946831AFFED8E2AFEE7F918FEEBBAE9" box="[285,313,1671,1695]" italics="true" pageId="20" pageNumber="545">
S
<quantity id="4CCAF2EDFFED8E2AFED6F918FEEBBAE9" box="[300,313,1671,1695]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="20" pageNumber="545" unit="in" value="2.0">2</quantity>
</emphasis>
in this latter species is similar to the position of that we have considered seta
<emphasis id="B946831AFFED8E2AFB66F918FB6ABAE9" box="[1180,1208,1671,1695]" italics="true" pageId="20" pageNumber="545">
S
<quantity id="4CCAF2EDFFED8E2AFB50F918FB6ABAE9" box="[1194,1208,1671,1695]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="20" pageNumber="545" unit="in" value="4.0">4</quantity>
</emphasis>
in
<taxonomicName id="4C32248BFFED8E2AFB21F917FA47BAD6" box="[1243,1429,1671,1696]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="anomalos" subGenus="Anthoseius">
<emphasis id="B946831AFFED8E2AFB21F917FB3ABAE9" box="[1243,1256,1672,1695]" italics="true" pageId="20" pageNumber="545">T</emphasis>
. (
<emphasis id="B946831AFFED8E2AFAFAF917FAC3BAE9" box="[1280,1297,1672,1695]" italics="true" pageId="20" pageNumber="545">A</emphasis>
.)
<emphasis id="B946831AFFED8E2AFAD2F918FA47BAD6" box="[1320,1429,1671,1696]" italics="true" pageId="20" pageNumber="545">anomalos</emphasis>
</taxonomicName>
. As
<taxonomicName id="4C32248BFFED8E2AFF47F933FE52BAB5" box="[189,384,1708,1732]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="arizonicus" subGenus="Anthoseius">
<emphasis id="B946831AFFED8E2AFF47F933FF19BAB5" box="[189,203,1708,1731]" italics="true" pageId="20" pageNumber="545">T</emphasis>
. (
<emphasis id="B946831AFFED8E2AFF19F933FF26BAB5" box="[227,244,1708,1731]" italics="true" pageId="20" pageNumber="545">A</emphasis>
.)
<emphasis id="B946831AFFED8E2AFEF6F933FE52BAB5" box="[268,384,1708,1731]" italics="true" pageId="20" pageNumber="545">arizonicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C32248BFFED8E2AFE4DF933FD87BAB2" box="[439,597,1707,1732]" class="Arachnida" family="Phytoseiidae" genus="Meyerius" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="waltheriae">
<emphasis id="B946831AFFED8E2AFE4DF933FE1CBAB5" box="[439,462,1708,1731]" italics="true" pageId="20" pageNumber="545">M</emphasis>
.
<emphasis id="B946831AFFED8E2AFE27F934FD87BAB2" box="[477,597,1707,1732]" italics="true" pageId="20" pageNumber="545">waltheriae</emphasis>
</taxonomicName>
have been described from only one female and
<taxonomicName id="4C32248BFFED8E2AFB88F933FAFFBAB2" box="[1138,1325,1707,1732]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="545" phylum="Arthropoda" rank="species" species="anomalos" subGenus="Anthoseius">
<emphasis id="B946831AFFED8E2AFB88F933FBADBAB5" box="[1138,1151,1708,1731]" italics="true" pageId="20" pageNumber="545">T</emphasis>
. (
<emphasis id="B946831AFFED8E2AFB6DF933FB7ABAB5" box="[1175,1192,1708,1731]" italics="true" pageId="20" pageNumber="545">A</emphasis>
.)
<emphasis id="B946831AFFED8E2AFB3AF934FAFFBAB2" box="[1216,1325,1707,1732]" italics="true" pageId="20" pageNumber="545">anomalos</emphasis>
</taxonomicName>
from two adults will be necessary to examine more individuals, including immature stages, to end the uncertainty about these dorsal setae.
</paragraph>
<paragraph id="8B8D5F08FFED8E2AFF3DF888FD1DBB22" blockId="20.[151,1437,1131,1948]" pageId="20" pageNumber="545">The shape of the male ventrianal shield in this species, broadly pentagonal, and the presence of two large metapodal plates are also a very unusual character.</paragraph>
<paragraph id="8B8D5F08FFED8E2AFF3DF8C0FBE1BBEA" blockId="20.[151,1437,1131,1948]" pageId="20" pageNumber="545">
The mite was collected on the Chilean pine or &quot;Pehuén&quot;,
<taxonomicName id="4C32248BFFED8E2AFCBEF8FFFBFBBB01" box="[836,1065,1888,1911]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="20" pageNumber="545" phylum="Tracheophyta" rank="species" species="araucana">
<emphasis id="B946831AFFED8E2AFCBEF8FFFBFBBB01" box="[836,1065,1888,1911]" italics="true" pageId="20" pageNumber="545">Araucaria araucana</emphasis>
</taxonomicName>
, considered as a living fossil and now restricted to Chilean and Argentinian south–central Andes above
<quantity id="4CCAF2EDFFED8E2AFC50F81BFBD4BBED" box="[938,1030,1924,1948]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="545" unit="m" value="1000.0">1,000 m</quantity>
asl.
</paragraph>
</subSubSection>
</treatment>
</document>
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