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<document id="936F32DE56E60F9C165E78F451C0EAB7" ID-DOI="10.5281/zenodo.175935" ID-GBIF-Dataset="0cd00333-558e-43a9-9944-fe1be57e0085" ID-ISSN="1175-5326" ID-Zenodo-Dep="175935" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459824946232" checkinUser="plazi" docAuthor="Erwin, Terry L. &amp; Moore, Wendy" docDate="2007" docId="0384CA77FFD5FFCA7BB6FF40FCD98C76" docLanguage="en" docName="zt01438p063.pdf" docOrigin="Zootaxa 1438" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Moriosomus Motschulsky 1855" docType="treatment" docVersion="9" lastPageNumber="52" masterDocId="FFBDB20FFFD6FFC97B21FFD7FFC18942" masterDocTitle="Taxonomic review of the Neotropical genus Moriosomus Motschulsky (Insecta: Coleoptera, Carabidae, Morionini) with notes on the way of life of the species" masterLastPageNumber="63" masterPageNumber="49" pageNumber="52" updateTime="1698222138880" updateUser="plazi">
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<mods:title id="E1ACED7122D64B7A33FCD7D82C959B8A">Taxonomic review of the Neotropical genus Moriosomus Motschulsky (Insecta: Coleoptera, Carabidae, Morionini) with notes on the way of life of the species</mods:title>
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<mods:namePart id="EE0BC699B1BA0E2BB5B06108030F233D">Erwin, Terry L.</mods:namePart>
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<mods:date id="7F848779E4662D70F2433F64740704C8">2007</mods:date>
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<treatment id="0384CA77FFD5FFCA7BB6FF40FCD98C76" ID-DOI="http://doi.org/10.5281/zenodo.6241324" ID-GBIF-Taxon="119356567" ID-Zenodo-Dep="6241324" LSID="urn:lsid:plazi:treatment:0384CA77FFD5FFCA7BB6FF40FCD98C76" httpUri="http://treatment.plazi.org/id/0384CA77FFD5FFCA7BB6FF40FCD98C76" lastPageNumber="52" pageId="3" pageNumber="52">
<subSubSection id="C33728EAFFD5FFCA7BB6FF40FDDC89F3" box="[151,541,151,177]" pageId="3" pageNumber="52" type="nomenclature">
<paragraph id="8B927B61FFD5FFCA7BB6FF40FDDC89F3" blockId="3.[151,541,151,177]" box="[151,541,151,177]" pageId="3" pageNumber="52">
<heading id="D0DACC0DFFD5FFCA7BB6FF40FDDC89F3" bold="true" box="[151,541,151,177]" fontSize="11" level="1" pageId="3" pageNumber="52" reason="1">
<taxonomicName id="4C2D00E2FFD5FFCA7BB6FF40FDDC89F3" ID-CoL="8KXVC" authority="Motschulsky, 1855" authorityName="Motschulsky" authorityYear="1855" box="[151,541,151,177]" class="Insecta" family="Carabidae" genus="Moriosomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B959A773FFD5FFCA7BB6FF40FDDC89F3" bold="true" box="[151,541,151,177]" pageId="3" pageNumber="52">
<emphasis id="B959A773FFD5FFCA7BB6FF40FEEE89F3" bold="true" box="[151,303,151,177]" italics="true" pageId="3" pageNumber="52">Moriosomus</emphasis>
<bibRefCitation id="EFBC0690FFD5FFCA7A14FF40FDDC89F3" author="Motschulsky" box="[309,541,151,177]" pageId="3" pageNumber="62" refString="Motschulsky, V. de (1855) Voyages. Lettre de M. de Motschulsky a M. Menetries. Etudes Enomologiques, pt. 4, 8 - 25." type="journal article" year="1855">Motschulsky, 1855</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33728EAFFD5FFCA7BB6FF35FC5C880E" pageId="3" pageNumber="52" type="materials_examined">
<paragraph id="8B927B61FFD5FFCA7BB6FF35FBFC89BE" blockId="3.[151,1437,226,1332]" box="[151,1085,226,252]" pageId="3" pageNumber="52">
<typeStatus id="5496C5C3FFD5FFCA7BB6FF35FF1089BE" box="[151,209,226,252]" pageId="3" pageNumber="52">Type</typeStatus>
species:
<taxonomicName id="4C2D00E2FFD5FFCA7A1DFF35FC9089BE" authority="Motschulsky, 1855:19" authorityName="Motschulsky" authorityPageNumber="19" authorityYear="1855" box="[316,849,226,252]" class="Insecta" family="Carabidae" genus="Moriosomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="sylvestris">
<emphasis id="B959A773FFD5FFCA7A1DFF35FD8289B9" box="[316,579,226,251]" italics="true" pageId="3" pageNumber="52">Moriosomus sylvestris</emphasis>
<bibRefCitation id="EFBC0690FFD5FFCA796AFF35FCF189BE" author="Motschulsky" box="[587,816,226,252]" pageId="3" pageNumber="62" refString="Motschulsky, V. de (1855) Voyages. Lettre de M. de Motschulsky a M. Menetries. Etudes Enomologiques, pt. 4, 8 - 25." type="journal article" year="1855">Motschulsky, 1855</bibRefCitation>
:19
</taxonomicName>
, original monotypy.
</paragraph>
<paragraph id="8B927B61FFD5FFCA7BE7FEDDFDF68866" blockId="3.[151,1437,226,1332]" box="[198,567,266,292]" pageId="3" pageNumber="52">
<typeStatus id="5496C5C3FFD5FFCA7BE7FEDDFEC18866" box="[198,256,266,292]" pageId="3" pageNumber="52">Type</typeStatus>
locality: Obispo,
<collectingCountry id="F33A3BF1FFD5FFCA7AF3FEDDFDF28866" box="[466,563,266,292]" name="Panama" pageId="3" pageNumber="52">Panamá</collectingCountry>
.
</paragraph>
<paragraph id="8B927B61FFD5FFCA7BE6FEE5FC5C880E" blockId="3.[151,1437,226,1332]" box="[199,925,306,332]" pageId="3" pageNumber="52">
<emphasis id="B959A773FFD5FFCA7BE6FEE5FD45880E" bold="true" box="[199,644,306,332]" pageId="3" pageNumber="52">Proposed English vernacular name.</emphasis>
Robust carabid beetles.
</paragraph>
</subSubSection>
<subSubSection id="C33728EAFFD5FFCA7BE6FE8DFA4788DE" pageId="3" pageNumber="52" type="diagnosis">
<paragraph id="8B927B61FFD5FFCA7BE6FE8DFA4788DE" blockId="3.[151,1437,226,1332]" pageId="3" pageNumber="52">
<emphasis id="B959A773FFD5FFCA7BE6FE8DFE868836" bold="true" box="[199,327,346,372]" pageId="3" pageNumber="52">Diagnosis.</emphasis>
Mentum tooth without medial notch. Body robust, pronotum and elytra convex, pronotum with lateral margins rounded anteriorly, posteriorly with short distinct sinuation near hind angle (
<figureCitation id="131667E4FFD5FFCA7E27FE55FAB888DE" box="[1286,1401,386,412]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,255,1369,1393]" captionStartId-1="5.[151,255,1662,1686]" captionStartId-2="6.[151,255,2011,2035]" captionTargetBox-0="[563,1035,707,1368]" captionTargetBox-1="[629,956,1115,1630]" captionTargetBox-2="[602,952,1464,2010]" captionTargetId-0="figure@4.[542,1044,673,1397]" captionTargetId-1="figure@5.[598,988,1079,1661]" captionTargetId-2="figure@6.[598,988,1433,2013]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 2. Moriosomus motschulskyi, Erwin &amp; Moore, new species, male, ADP 108774, habitus, ABL = 14.5 mm." captionText-1="FIGURE 3. Moriosomus seticollis Straneo, male, ADP 108773, habitus, ABL = 13.5 mm." captionText-2="FIGURE 4. Moriosomus sylvestris Motschulsky, male, ADP 007358, habitus, ABL = 12.8 mm." httpUri-0="https://zenodo.org/record/175937/files/figure.png" httpUri-1="https://zenodo.org/record/175938/files/figure.png" httpUri-2="https://zenodo.org/record/175939/files/figure.png" pageId="3" pageNumber="52">Figs. 24</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33728EAFFD5FFCA7BE6FE7DFCD98C76" pageId="3" pageNumber="52" type="description">
<paragraph id="8B927B61FFD5FFCA7BE6FE7DFF3A8D5E" blockId="3.[151,1437,226,1332]" pageId="3" pageNumber="52">
<emphasis id="B959A773FFD5FFCA7BE6FE7DFE9E8886" bold="true" box="[199,351,426,452]" pageId="3" pageNumber="52">Description.</emphasis>
Size medium for tribe and family, ABL = 11.5 to 17.2 mm, SBL = 11.3 to
<quantity id="4CD5D684FFD5FFCA7FEBFE7DFAE18886" box="[1226,1312,426,452]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="52" unit="mm" value="17.0">17 mm</quantity>
, TW = 4.0 to 5.8 mm.
<emphasis id="B959A773FFD5FFCA7A3EFE05FEB588A9" box="[287,372,466,491]" italics="true" pageId="3" pageNumber="52">Luster:</emphasis>
Surface shiny.
<emphasis id="B959A773FFD5FFCA790AFE05FDB588A9" box="[555,628,466,491]" italics="true" pageId="3" pageNumber="52">Head:</emphasis>
Antennae with scape glabrous, antennomeres 23 with ring of setae at apex, antennomere 4 sparsely setiferous in apical 2/3; antennomeres 511 densely setose on lateral margins, sparsely setiferous along midline; labial palpus with penultimate palpomere bisetose, other palpomeres glabrous.
<emphasis id="B959A773FFD5FFCA7BC7FD9DFEBA8B21" box="[230,379,586,611]" italics="true" pageId="3" pageNumber="52">Pterothorax:</emphasis>
Scutellum small; elytra convex; humerus moderately prominent with small dentiform projection; parascutellar interneur absent; intervals convex.
<emphasis id="B959A773FFD5FFCA787AFDA5FC5D8BC9" box="[859,924,626,651]" italics="true" pageId="3" pageNumber="52">Legs:</emphasis>
<collectingCountry id="F33A3BF1FFD5FFCA788AFDA5FC278BCE" box="[939,998,626,652]" name="India" pageId="3" pageNumber="52">Hind</collectingCountry>
legs arched, otherwise normal for Morionini. Male with biseriate white squamo-setae ventrally on tarsomeres 13 of forelegs.
<emphasis id="B959A773FFD5FFCA7FF2FD4DFA838BF1" box="[1235,1346,666,691]" italics="true" pageId="3" pageNumber="52">Abdomen</emphasis>
: Abdomen with sterna IIIVI glabrous other than a pair of ambulatory setae on each; sternum VII with a latching device on the lateral dorsal margin (as noted in
<bibRefCitation id="EFBC0690FFD5FFCA79F3FD3DFC968A46" author="Allen" box="[722,855,746,772]" pageId="3" pageNumber="61" refString="Allen, R. T. (1968) A synopsis of the tribe Morionini in the Western Hemisphere with descriptions of two new species (Coleoptera: Carabidae). Caribbean Journal of Science, 8 (3 - 4), 141 - 163." type="journal article" year="1968">Allen 1968</bibRefCitation>
for
<taxonomicName id="4C2D00E2FFD5FFCA78ABFD3DFC238A41" box="[906,994,746,771]" class="Insecta" family="Carabidae" genus="Morion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B959A773FFD5FFCA78ABFD3DFC238A41" box="[906,994,746,771]" italics="true" pageId="3" pageNumber="52">Morion</emphasis>
</taxonomicName>
).
<emphasis id="B959A773FFD5FFCA78DDFD3DFB688A41" box="[1020,1193,746,771]" italics="true" pageId="3" pageNumber="52">Male genitalia</emphasis>
(
<figureCitation id="131667E4FFD5FFCA7F9AFD3DFAF18A46" box="[1211,1328,746,772]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="7.[151,255,960,984]" captionStartId-1="7.[151,255,1937,1961]" captionStartId-2="8.[151,255,967,991]" captionTargetBox-0="[212,1415,204,927]" captionTargetBox-1="[211,1343,1119,1874]" captionTargetBox-2="[294,1360,245,916]" captionTargetId-0="figure@7.[148,1438,176,955]" captionTargetId-1="figure@7.[192,1394,1107,1914]" captionTargetId-2="figure@8.[191,1395,194,943]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 5. Moriosomus motschulskyi Erwin &amp; Moore, new species, ADP 108774. Male genitalia, including phallus, endophallus, and parameres (a) dorsal aspect, (b) ventral aspect, and (c) left lateral aspect. Legend: bb— basal bulb; bo—basal opening; c—crest of basal bulb; s—shaft; o—ostium; rp—right paramere; lp—left paramere." captionText-1="FIGURE 6. Moriosomus seticollis Straneo, ADP 108773. Male genitalia, including phallus, endophallus, and parameres (a) dorsal aspect, (b) ventral aspect, and (c) left lateral aspect." captionText-2="FIGURE 7. Moriosomus sylvestris Motschulsky, ADP 003301. Male genitalia, including phallus, endophallus, and parameres (a) dorsal aspect, (b) ventral aspect, and (c) left lateral aspect." httpUri-0="https://zenodo.org/record/175940/files/figure.png" httpUri-1="https://zenodo.org/record/175941/files/figure.png" httpUri-2="https://zenodo.org/record/175942/files/figure.png" pageId="3" pageNumber="52">Figs. 57</figureCitation>
): Phallus anopic; in dorsoventral aspect (
<figureCitation id="131667E4FFD5FFCA792AFCC5FD9C8A6E" box="[523,605,786,812]" captionStart="FIGURE 5" captionStartId="7.[151,255,960,984]" captionTargetBox="[212,1415,204,927]" captionTargetId="figure@7.[148,1438,176,955]" captionTargetPageId="7" captionText="FIGURE 5. Moriosomus motschulskyi Erwin &amp; Moore, new species, ADP 108774. Male genitalia, including phallus, endophallus, and parameres (a) dorsal aspect, (b) ventral aspect, and (c) left lateral aspect. Legend: bb— basal bulb; bo—basal opening; c—crest of basal bulb; s—shaft; o—ostium; rp—right paramere; lp—left paramere." httpUri="https://zenodo.org/record/175940/files/figure.png" pageId="3" pageNumber="52">Figs. 5</figureCitation>
ab, 7ab) broad, basal bulb crested, dorsal surface extensively membranous, apex more or less broadly subtruncate; laterally (
<figureCitation id="131667E4FFD5FFCA7863FCEDFC538A16" box="[834,914,826,852]" captionStart="FIGURE 5" captionStartId="7.[151,255,960,984]" captionTargetBox="[212,1415,204,927]" captionTargetId="figure@7.[148,1438,176,955]" captionTargetPageId="7" captionText="FIGURE 5. Moriosomus motschulskyi Erwin &amp; Moore, new species, ADP 108774. Male genitalia, including phallus, endophallus, and parameres (a) dorsal aspect, (b) ventral aspect, and (c) left lateral aspect. Legend: bb— basal bulb; bo—basal opening; c—crest of basal bulb; s—shaft; o—ostium; rp—right paramere; lp—left paramere." httpUri="https://zenodo.org/record/175940/files/figure.png" pageId="3" pageNumber="52">Figs. 5</figureCitation>
c7c), with shaft curved ventrad, apical portion narrowed and bent sharply. Endophallus with or without patches of microtrichia. Parameres (
<figureCitation id="131667E4FFD5FFCA7E26FCB5FA978A3E" box="[1287,1366,866,892]" captionStart="FIGURE 5" captionStartId="7.[151,255,960,984]" captionTargetBox="[212,1415,204,927]" captionTargetId="figure@7.[148,1438,176,955]" captionTargetPageId="7" captionText="FIGURE 5. Moriosomus motschulskyi Erwin &amp; Moore, new species, ADP 108774. Male genitalia, including phallus, endophallus, and parameres (a) dorsal aspect, (b) ventral aspect, and (c) left lateral aspect. Legend: bb— basal bulb; bo—basal opening; c—crest of basal bulb; s—shaft; o—ostium; rp—right paramere; lp—left paramere." httpUri="https://zenodo.org/record/175940/files/figure.png" pageId="3" pageNumber="52">Figs. 5</figureCitation>
b7b) glabrous, left wider than right, latter more or less digitiform.
<emphasis id="B959A773FFD5FFCA787FFC5DFB4E8AE1" box="[862,1167,906,931]" italics="true" pageId="3" pageNumber="52">Female reproductive tract</emphasis>
(
<figureCitation id="131667E4FFD5FFCA7FBEFC5DFB278AE6" box="[1183,1254,906,932]" captionStart="FIGURE 8" captionStartId="8.[151,255,1679,1703]" captionTargetBox="[512,1074,1079,1654]" captionTargetId="figure@8.[512,1075,1079,1655]" captionTargetPageId="8" captionText="FIGURE 8. Moriosomus seticollis Straneo. Ventral view of female reproductive tract. bc = bursa copulatrix; co = common oviduct; sp = spermatheca; sg = spermatheca appended gland; sgd = spermatheca gland duct diverticula; gc 1 = gonocoxite- 1; gc 2 = gonocoxite- 2." httpUri="https://zenodo.org/record/175943/files/figure.png" pageId="3" pageNumber="52">Fig. 8</figureCitation>
): Spermathecal gland and associated diverticula attach at the base of the spermatheca; gonocoxite-2 as long as gonocoxite-1, narrow and with acute apex, otherwise similar to
<taxonomicName id="4C2D00E2FFD5FFCA79FDFC0DFBFA8AB6" authority="Latreille" authorityName="Latreille" box="[732,1083,986,1012]" class="Insecta" family="Carabidae" genus="Morion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="monilicornis">
<emphasis id="B959A773FFD5FFCA79FDFC0DFC118AB1" box="[732,976,986,1011]" italics="true" pageId="3" pageNumber="52">Morion monilicornis</emphasis>
Latreille
</taxonomicName>
(see
<bibRefCitation id="EFBC0690FFD5FFCA7F58FC0DFA568AB6" author="Liebherr" box="[1145,1431,986,1012]" pageId="3" pageNumber="62" refString="Liebherr, J. K. &amp; Will, K. W. (1998) Inferring phylogenetic relationships within Carabidae (Insecta, Coleoptera) from characters of the female reproductive tract. In: G. E. Ball, Casale, A. &amp; Vigna Taglianti, A. (Eds) Phylogeny and classification of Caraboidea (Coleoptera: Adephaga). Museo Regionale di Scienze Naturali, Torino, pp. 107 - 170." type="book chapter" year="1998">Liebherr and Will, 1998</bibRefCitation>
, Fig. 41).
</paragraph>
<paragraph id="8B927B61FFD5FFCA7BE6FBFDFDAD8DFE" blockId="3.[151,1437,226,1332]" pageId="3" pageNumber="52">
<emphasis id="B959A773FFD5FFCA7BE6FBFDFE3C8D06" bold="true" box="[199,509,1066,1092]" pageId="3" pageNumber="52">Geographic distribution.</emphasis>
The geographical range of this genus extends in Lower Middle
<collectingCountry id="F33A3BF1FFD5FFCA7FD7FBFDFA9A8D06" box="[1270,1371,1066,1092]" name="United States of America" pageId="3" pageNumber="52">America</collectingCountry>
from
<collectingCountry id="F33A3BF1FFD5FFCA7BB6FB85FECE8D2E" box="[151,271,1106,1132]" name="Nicaragua" pageId="3" pageNumber="52">Nicaragua</collectingCountry>
to
<collectingCountry id="F33A3BF1FFD5FFCA7A19FB85FE598D2E" box="[312,408,1106,1132]" name="Panama" pageId="3" pageNumber="52">Panamá</collectingCountry>
, and in northwestern South
<collectingCountry id="F33A3BF1FFD5FFCA79C8FB85FC8F8D2E" box="[745,846,1106,1132]" name="United States of America" pageId="3" pageNumber="52">America</collectingCountry>
from
<collectingCountry id="F33A3BF1FFD5FFCA78B8FB85FBCC8D2E" box="[921,1037,1106,1132]" name="Colombia" pageId="3" pageNumber="52">Colombia</collectingCountry>
southward to
<collectingCountry id="F33A3BF1FFD5FFCA7F9BFB85FB318D2E" box="[1210,1264,1106,1132]" name="Peru" pageId="3" pageNumber="52">Perú</collectingCountry>
(
<figureCitation id="131667E4FFD5FFCA7E20FB85FA898D2E" box="[1281,1352,1106,1132]" captionStart="FIGURE 1" captionStartId="1.[151,255,700,724]" captionTargetBox="[566,1031,167,679]" captionTargetId="figure@1.[566,1031,167,679]" captionTargetPageId="1" captionText="FIGURE 1. Map of Lower Middle America, the West Indies and northwestern South America in the Western Hemisphere, featuring the known distribution of Moriosomus Motschulsky." httpUri="https://zenodo.org/record/175936/files/figure.png" pageId="3" pageNumber="52">Fig. 1</figureCitation>
). Both South American species are known from the cis-Andean highlands above
<quantity id="4CD5D684FFD5FFCA78D9FBADFB8D8DD6" box="[1016,1100,1146,1172]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="52" unit="m" value="1500.0">1500m</quantity>
, whereas the Central American species is found only below
<quantity id="4CD5D684FFD5FFCA7932FB75FDA68DFE" box="[531,615,1186,1212]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="52" unit="m" value="1100.0">1100m</quantity>
.
</paragraph>
<paragraph id="8B927B61FFD5FFCA7BE6FB1DFCD98C76" blockId="3.[151,1437,226,1332]" pageId="3" pageNumber="52">
<emphasis id="B959A773FFD5FFCA7BE6FB1DFED58DA6" bold="true" box="[199,276,1226,1252]" pageId="3" pageNumber="52">Notes.</emphasis>
Date of the first description of
<taxonomicName id="4C2D00E2FFD5FFCA79ADFB1CFCDE8DA6" box="[652,799,1227,1252]" class="Insecta" family="Carabidae" genus="Moriosomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B959A773FFD5FFCA79ADFB1CFCDE8DA6" box="[652,799,1227,1252]" italics="true" pageId="3" pageNumber="52">Moriosomus</emphasis>
</taxonomicName>
is cited by many previous authors as 1864. However, Motschulsky first described the
<typeStatus id="5496C5C3FFD5FFCA7932FB25FD848C4E" box="[531,581,1266,1292]" pageId="3" pageNumber="52">type</typeStatus>
species,
<emphasis id="B959A773FFD5FFCA7993FB24FC8E8C4E" box="[690,847,1267,1292]" italics="true" pageId="3" pageNumber="52">
<taxonomicName id="4C2D00E2FFD5FFCA7993FB24FC8B8C4E" box="[690,842,1267,1292]" class="Insecta" family="Carabidae" genus="Moriosomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="sylvestris">M. sylvestris</taxonomicName>
,
</emphasis>
in 1855 at which time he included that species in the new genus
<taxonomicName id="4C2D00E2FFD5FFCA7A67FACCFE188C76" box="[326,473,1307,1332]" class="Insecta" family="Carabidae" genus="Moriosomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B959A773FFD5FFCA7A67FACCFE188C76" box="[326,473,1307,1332]" italics="true" pageId="3" pageNumber="52">Moriosomus</emphasis>
</taxonomicName>
(see Appendix for details).
</paragraph>
</subSubSection>
</treatment>
</document>