581 lines
61 KiB
XML
581 lines
61 KiB
XML
<document ID-DOI="10.11646/zootaxa.5231.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7571485" ID-ZooBank="9C66A388-C221-4D17-B2AA-F9EB7F1EC940" checkinTime="1674711822273" checkinUser="plazi" docAuthor="Kraus, Fred" docDate="2023" docId="0381A239FFFCFFDA7E8DF9D4FB48FB12" docLanguage="en" docName="zootaxa.5231.1.1.pdf" docOrigin="Zootaxa 5231 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gerrhopilus polyadenus Kraus 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="FFB8DA41FFF4FFD17E1AFFE2FFBAFFD7" masterDocTitle="A revision of Gerrhopilus inornatus (Squamata: Gerrhopilidae) reveals a multi-species complex" masterLastPageNumber="23" masterPageNumber="1" pageNumber="9" updateTime="1674753466069" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of Gerrhopilus inornatus (Squamata: Gerrhopilidae) reveals a multi-species complex</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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||
<mods:namePart>Kraus, Fred</mods:namePart>
|
||
<mods:affiliation>Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, U. S. A</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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||
<mods:date>2023</mods:date>
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||
<mods:detail type="pubDate">
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<mods:number>2023-01-26</mods:number>
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||
</mods:detail>
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||
<mods:detail type="volume">
|
||
<mods:number>5231</mods:number>
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||
</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>23</mods:end>
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</mods:extent>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5231.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5231.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7571485</mods:identifier>
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<mods:identifier type="ZooBank">9C66A388-C221-4D17-B2AA-F9EB7F1EC940</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:0381A239FFFCFFDA7E8DF9D4FB48FB12" httpUri="http://treatment.plazi.org/id/0381A239FFFCFFDA7E8DF9D4FB48FB12" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
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<subSubSection box="[151,543,1590,1617]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph blockId="8.[151,543,1590,1653]" box="[151,543,1590,1617]" pageId="8" pageNumber="9">
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<heading bold="true" box="[151,543,1590,1617]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis bold="true" box="[151,543,1590,1617]" pageId="8" pageNumber="9">
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<taxonomicName authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[151,442,1590,1617]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="polyadenus" status="sp. nov.">
|
||
<emphasis bold="true" box="[151,442,1590,1617]" italics="true" pageId="8" pageNumber="9">Gerrhopilus polyadenus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[449,543,1591,1617]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,280,1627,1653]" pageId="8" pageNumber="9" type="description">
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<paragraph blockId="8.[151,543,1590,1653]" box="[151,280,1627,1653]" pageId="8" pageNumber="9">
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<figureCitation box="[151,280,1627,1653]" captionStart="FIGURE 4" captionStartId="10.[152,255,1935,1959]" captionTargetBox="[169,1419,363,1893]" captionTargetId="figure-122@10.[151,1436,345,1911]" captionTargetPageId="10" captionText="FIGURE 4. Dorsal and lateral views of heads for members of the Gerrhopilus inornatus Group having 22 midbody scale rows and uniformly dark venters. A, B holotype of Gerrhopilus papuanorum (BPBM 17236); C, D holotype of Gerrhopilus polyadenus (BMNH 1901.11.27.10)." figureDoi="http://doi.org/10.5281/zenodo.7571497" httpUri="https://zenodo.org/record/7571497/files/figure.png" pageId="8" pageNumber="9">Figs. 4C, D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="reference_group">
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<paragraph blockId="8.[151,662,1695,1750]" box="[151,541,1695,1719]" pageId="8" pageNumber="9">
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<treatmentCitationGroup box="[151,541,1695,1719]" pageId="8" pageNumber="9">
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<treatmentCitation author="de Rooij, N." box="[151,537,1695,1719]" page="14" pageId="8" pageNumber="9" year="1917">
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<taxonomicName authority="de Rooij 1917: 14" authorityName="de Rooij" authorityPageNumber="14" authorityYear="1917" box="[151,537,1695,1719]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[151,346,1695,1719]" italics="true" pageId="8" pageNumber="9">Typhlops inornatus</emphasis>
|
||
<bibRefCitation author="de Rooij, N." box="[352,537,1695,1719]" pageId="8" pageNumber="9" refId="ref18970" refString="de Rooij, N. (1917) The reptiles of the Indo-Australian Archipelago, II: Ophidia. E. J. Brill, Leiden, 334 pp." type="book" year="1917">de Rooij 1917: 14</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,662,1695,1750]" box="[151,662,1726,1750]" pageId="8" pageNumber="9">
|
||
<treatmentCitationGroup box="[151,662,1726,1750]" pageId="8" pageNumber="9">
|
||
<treatmentCitation author="Pyron, R. A. & Wallach, V." box="[151,658,1726,1750]" page="79" pageId="8" pageNumber="9" year="2014">
|
||
<taxonomicName authority="Pyron & Wallach 2014: 79" authorityName="Pyron & Wallach" authorityPageNumber="79" authorityYear="2014" box="[151,658,1726,1750]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[151,378,1726,1750]" italics="true" pageId="8" pageNumber="9">Gerrhopilus inornatus</emphasis>
|
||
<bibRefCitation author="Pyron, R. A. & Wallach, V." box="[384,658,1726,1750]" pageId="8" pageNumber="9" pagination="1 - 81" refId="ref19325" refString="Pyron, R. A. & Wallach, V. (2014) Systematics of the blind snakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence. Zootaxa, 3829 (1), 1 - 81. https: // doi. org / 10.11646 / zootaxa. 3829.1.1" type="journal article" year="2014">Pyron & Wallach 2014: 79</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="9" type="materials_examined">
|
||
<paragraph blockId="8.[151,1436,1796,2039]" box="[151,1275,1796,1823]" pageId="8" pageNumber="9">
|
||
<materialsCitation box="[151,1275,1796,1823]" collectionCode="BMNH" collectorName="H. S. Rohu." country="Papua New Guinea" location="Mt Albert Edward" pageId="8" pageNumber="9" specimenCode="BMNH 1901.11" specimenCount="1" typeStatus="holotype">
|
||
<emphasis bold="true" box="[151,267,1796,1822]" pageId="8" pageNumber="9">
|
||
<typeStatus box="[151,260,1796,1822]" pageId="8" pageNumber="9">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode box="[274,457,1796,1823]" collectionCode="BMNH" pageId="8" pageNumber="9">BMNH 1901.11</specimenCode>
|
||
.27.10, collected at
|
||
<location LSID="urn:lsid:plazi:treatment:0381A239FFFCFFDA7E8DF9D4FB48FB12:8EF745F4FFFCFFD97CB8F8E7FCCBF8C9" box="[674,881,1796,1823]" country="Papua New Guinea" name="Mt Albert Edward" pageId="8" pageNumber="9">Mt Albert Edward</location>
|
||
,
|
||
<collectingCountry box="[892,1108,1796,1823]" name="Papua New Guinea" pageId="8" pageNumber="9">Papua New Guinea</collectingCountry>
|
||
by
|
||
<collectorName box="[1150,1275,1796,1822]" pageId="8" pageNumber="9">H.S. Rohu.</collectorName>
|
||
</materialsCitation>
|
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</paragraph>
|
||
<paragraph blockId="8.[151,1436,1796,2039]" pageId="8" pageNumber="9">
|
||
<materialsCitation collectionCode="MCZ" country="Papua New Guinea" elevation="120" latitude="-8.76" location="Oro Province" longLatPrecision="781" longitude="148.18" pageId="8" pageNumber="9" specimenCode="MCZ 140728" specimenCount="1" stateProvince="Northern" typeStatus="paratype">
|
||
<emphasis bold="true" box="[199,315,1832,1858]" pageId="8" pageNumber="9">
|
||
<typeStatus box="[199,310,1832,1858]" pageId="8" pageNumber="9">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode box="[324,481,1832,1858]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="9" type="Museum">MCZ 140728</specimenCode>
|
||
, collected at Sangara [
|
||
<geoCoordinate box="[741,830,1832,1858]" degrees="8.76" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="-8.76">8.76° S</geoCoordinate>
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||
,
|
||
<geoCoordinate box="[845,964,1832,1859]" degrees="148.18" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="555" value="148.18">148.18° E</geoCoordinate>
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||
],
|
||
<elevation box="[986,1122,1832,1859]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="m" value="120.0">
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<quantity box="[986,1059,1832,1859]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="m" value="120.0">120 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<collectingRegion box="[1137,1292,1832,1858]" country="Papua New Guinea" name="Northern" pageId="8" pageNumber="9">Oro Province</collectingRegion>
|
||
,
|
||
<collectingCountry name="Papua New Guinea" pageId="8" pageNumber="9">Papua New Guinea</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
|
||
<paragraph blockId="8.[151,1436,1796,2039]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[199,322,1904,1930]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
|
||
A
|
||
<taxonomicName box="[353,576,1904,1930]" pageId="8" pageNumber="9">
|
||
<emphasis box="[353,489,1904,1930]" italics="true" pageId="8" pageNumber="9">Gerrhopilus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
characterized by the unique combination of having a rounded snout in lateral view; LSR = 22 at midbody; TSR = 320; loreal absent; supralabial imbrication pattern T-V; subocular scale one; presubocular scale absent; a sharp, protruding tail spine that is dark brown with a white or corneous-brown tip; 29 glands in the prefrontal,
|
||
<quantity box="[472,545,2012,2038]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.286" metricValueMin="2.032" pageId="8" pageNumber="9" unit="in" value="8.5" valueMax="9.0" valueMin="8.0">8–9 in</quantity>
|
||
the supraocular,
|
||
<quantity box="[743,845,2012,2038]" metricMagnitude="-1" metricUnit="m" metricValue="5.715" metricValueMax="5.842" metricValueMin="5.588" pageId="8" pageNumber="9" unit="in" value="22.5" valueMax="23.0" valueMin="22.0">22–23 in</quantity>
|
||
the ocular,
|
||
<quantity box="[982,1083,2012,2039]" metricMagnitude="0" metricUnit="m" metricValue="1.2192" metricValueMax="1.27" metricValueMin="1.1684" pageId="8" pageNumber="9" unit="in" value="48.0" valueMax="50.0" valueMin="46.0">46–50 in</quantity>
|
||
the preocular,
|
||
<quantity box="[1257,1302,2012,2038]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="9" unit="in" value="6.0">6 in</quantity>
|
||
the frontal,
|
||
<quantity box="[151,209,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="9" pageNumber="10" unit="in" value="16.0">16 in</quantity>
|
||
each subocular; L/W = 36.2–36.3; TL/SVL =
|
||
<geoCoordinate box="[737,801,151,177]" degrees="0.028" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="55" value="0.028">0.028</geoCoordinate>
|
||
<geoCoordinate box="[801,881,151,177]" degrees="0.036" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="55" value="-0.036">–0.036</geoCoordinate>
|
||
; dorsum and venter uniformly pale brown with a contrasting dark-brown patch around tail spine. Its size is moderate for this species group (
|
||
<tableCitation box="[1166,1249,187,213]" captionStart="TABLE 1" captionStartId="17.[151,244,552,578]" captionTargetPageId="17" captionText="TABLE 1. Mensural and count data for members of the Gerrhopilus inornatus Group. Forward slashes separate counts on the right and left sides, respectively." pageId="9" pageNumber="10">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">
|
||
<emphasis bold="true" box="[199,586,222,249]" pageId="9" pageNumber="10">Comparisons with other species.</emphasis>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[594,866,223,249]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="polyadenus">
|
||
<emphasis box="[594,866,223,249]" italics="true" pageId="9" pageNumber="10">Gerrhopilus polyadenus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" box="[1118,1279,223,249]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="fredparkeri">
|
||
<emphasis box="[1118,1279,223,249]" italics="true" pageId="9" pageNumber="10">G. fredparkeri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[1294,1394,223,249]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hades">
|
||
<emphasis box="[1294,1394,223,249]" italics="true" pageId="9" pageNumber="10">G. hades</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis italics="true" pageId="9" pageNumber="10">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[272,429,259,285]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[272,429,259,285]" italics="true" pageId="9" pageNumber="10">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[489,620,259,285]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis box="[489,620,259,285]" italics="true" pageId="9" pageNumber="10">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
in having 22 longitudinal scale rows at midbody (vs
|
||
<quantity box="[1207,1263,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="9" pageNumber="10" unit="in" value="16.0">16 in</quantity>
|
||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" box="[1269,1429,259,285]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="fredparkeri">
|
||
<emphasis box="[1269,1429,259,285]" italics="true" pageId="9" pageNumber="10">G. fredparkeri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity box="[151,211,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="9" pageNumber="10" unit="in" value="18.0">18 in</quantity>
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[220,322,295,321]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hades">
|
||
<emphasis box="[220,322,295,321]" italics="true" pageId="9" pageNumber="10">G. hades</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[382,517,295,321]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis box="[382,517,295,321]" italics="true" pageId="9" pageNumber="10">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity box="[584,644,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="10" unit="in" value="20.0">20 in</quantity>
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[653,797,295,321]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[653,797,295,321]" italics="true" pageId="9" pageNumber="10">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[857,1018,295,321]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[857,1018,295,321]" italics="true" pageId="9" pageNumber="10">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
). It is further distinguished from
|
||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="fredparkeri">
|
||
<emphasis italics="true" pageId="9" pageNumber="10">G. fredparkeri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[331,472,331,357]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[331,472,331,357]" italics="true" pageId="9" pageNumber="10">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
in having fewer transverse scale rows (294–320 vs
|
||
<quantity box="[1048,1118,330,357]" metricMagnitude="1" metricUnit="m" metricValue="1.36906" pageId="9" pageNumber="10" unit="in" value="539.0">539 in</quantity>
|
||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" box="[1125,1285,331,357]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="fredparkeri">
|
||
<emphasis box="[1125,1285,331,357]" italics="true" pageId="9" pageNumber="10">G. fredparkeri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity metricMagnitude="0" metricUnit="m" metricValue="9.5123" metricValueMax="9.525" metricValueMin="9.4996" pageId="9" pageNumber="10" unit="in" value="374.5" valueMax="375.0" valueMin="374.0">374–375 in</quantity>
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[183,327,367,393]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[183,327,367,393]" italics="true" pageId="9" pageNumber="10">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[418,520,367,393]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hades">
|
||
<emphasis box="[418,520,367,393]" italics="true" pageId="9" pageNumber="10">G. hades</emphasis>
|
||
</taxonomicName>
|
||
in having 22 longitudinal scale rows behind the head (vs
|
||
<quantity box="[1197,1257,367,393]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="9" pageNumber="10" unit="in" value="18.0">18 in</quantity>
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[1267,1369,367,393]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hades">
|
||
<emphasis box="[1267,1369,367,393]" italics="true" pageId="9" pageNumber="10">G. hades</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[216,351,403,429]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis box="[216,351,403,429]" italics="true" pageId="9" pageNumber="10">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
in having a single postocular on each side of the head (vs two in
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[1127,1262,403,429]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis box="[1127,1262,403,429]" italics="true" pageId="9" pageNumber="10">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
) and lacking a presubocular (vs present in
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[470,604,439,465]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis box="[470,604,439,465]" italics="true" pageId="9" pageNumber="10">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[630,903,439,465]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="polyadenus">
|
||
<emphasis box="[630,903,439,465]" italics="true" pageId="9" pageNumber="10">Gerrhopilus polyadenus</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguished from
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[1249,1392,439,465]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[1249,1392,439,465]" italics="true" pageId="9" pageNumber="10">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis italics="true" pageId="9" pageNumber="10">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[341,521,475,501]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[341,521,475,501]" italics="true" pageId="9" pageNumber="10">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
by its pale-brown color (dark brown or black in the other species), dark-brown scales around the tail spine (area black or white in the other species), and much greater number of glands in the ocular (11–23 vs
|
||
<quantity box="[342,412,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="15.24" metricValueMin="0.0" pageId="9" pageNumber="10" unit="in" value="3.0" valueMax="6.0" valueMin="0.0">0–6 in</quantity>
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[418,558,547,573]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="22" phylum="Chordata" rank="species" species="inornatus">
|
||
G.
|
||
<emphasis box="[451,558,547,572]" italics="true" pageId="9" pageNumber="10">inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity box="[571,641,547,573]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.286" metricValueMin="1.016" pageId="9" pageNumber="10" unit="in" value="6.5" valueMax="9.0" valueMin="4.0">4–9 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[647,804,547,573]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[647,804,547,573]" italics="true" pageId="9" pageNumber="10">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity box="[864,934,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="7.62" metricValueMin="0.0" pageId="9" pageNumber="10" unit="in" value="1.5" valueMax="3.0" valueMin="0.0">0–3 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[940,1116,547,573]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[940,1116,547,573]" italics="true" pageId="9" pageNumber="10">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
), preocular (42–50 vs
|
||
<quantity metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.096" metricValueMin="5.588" pageId="9" pageNumber="10" unit="in" value="23.0" valueMax="24.0" valueMin="22.0">22–24 in</quantity>
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[180,322,583,609]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[180,322,583,609]" italics="true" pageId="9" pageNumber="10">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity box="[337,436,582,609]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" metricValueMax="9.906" metricValueMin="8.89" pageId="9" pageNumber="10" unit="in" value="37.0" valueMax="39.0" valueMin="35.0">35–39 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[443,602,583,609]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[443,602,583,609]" italics="true" pageId="9" pageNumber="10">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity box="[665,764,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="7.112" metricValueMin="4.572" pageId="9" pageNumber="10" unit="in" value="23.0" valueMax="28.0" valueMin="18.0">18–28 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[771,949,583,609]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[771,949,583,609]" italics="true" pageId="9" pageNumber="10">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
), prefrontal (11–29 vs
|
||
<quantity box="[1209,1280,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="9" pageNumber="10" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[1287,1429,583,609]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[1287,1429,583,609]" italics="true" pageId="9" pageNumber="10">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity box="[151,222,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="9" pageNumber="10" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">1–2 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[229,388,619,645]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[229,388,619,645]" italics="true" pageId="9" pageNumber="10">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity box="[450,521,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="12.7" metricValueMin="2.54" pageId="9" pageNumber="10" unit="in" value="3.0" valueMax="5.0" valueMin="1.0">1–5 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[528,706,619,645]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[528,706,619,645]" italics="true" pageId="9" pageNumber="10">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
), supraocular (5–9 vs
|
||
<quantity box="[958,1029,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="9" pageNumber="10" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[1036,1178,619,645]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[1036,1178,619,645]" italics="true" pageId="9" pageNumber="10">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity box="[1192,1263,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="7.62" metricValueMin="0.0" pageId="9" pageNumber="10" unit="in" value="1.5" valueMax="3.0" valueMin="0.0">0–3 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1270,1429,619,645]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[1270,1429,619,645]" italics="true" pageId="9" pageNumber="10">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity box="[201,275,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="10.16" metricValueMin="0.0" pageId="9" pageNumber="10" unit="in" value="2.0" valueMax="4.0" valueMin="0.0">0–4 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[285,465,655,681]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[285,465,655,681]" italics="true" pageId="9" pageNumber="10">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
), and subocular (5–16 vs
|
||
<quantity box="[770,843,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" metricValueMax="5.08" metricValueMin="0.0" pageId="9" pageNumber="10" unit="in" value="1.0" valueMax="2.0" valueMin="0.0">0–2 in</quantity>
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[853,997,655,681]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[853,997,655,681]" italics="true" pageId="9" pageNumber="10">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity box="[1014,1088,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="9" pageNumber="10" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1097,1258,655,681]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[1097,1258,655,681]" italics="true" pageId="9" pageNumber="10">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity box="[1326,1399,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" metricValueMax="5.08" metricValueMin="0.0" pageId="9" pageNumber="10" unit="in" value="1.0" valueMax="2.0" valueMin="0.0">0–2 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis italics="true" pageId="9" pageNumber="10">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
). It is further distinguished from
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[665,842,691,717]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[665,842,691,717]" italics="true" pageId="9" pageNumber="10">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
by its stouter habitus (L/W = 36.2–36.3 vs
|
||
<quantity metricMagnitude="0" metricUnit="m" metricValue="1.3296899999999998" metricValueMax="1.3589000000000002" metricValueMin="1.3004799999999999" pageId="9" pageNumber="10" unit="in" value="52.35" valueMax="53.5" valueMin="51.2">51.2–53.5 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[180,357,727,753]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[180,357,727,753]" italics="true" pageId="9" pageNumber="10">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
) and longer tail (TL/SVL =
|
||
<geoCoordinate box="[675,737,727,753]" degrees="0.028" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="55" value="0.028">0.028</geoCoordinate>
|
||
<geoCoordinate box="[738,815,727,753]" degrees="0.036" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="55" value="-0.036">–0.036</geoCoordinate>
|
||
vs
|
||
<quantity box="[854,1023,727,753]" metricMagnitude="-4" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.604" metricValueMin="5.334" pageId="9" pageNumber="10" unit="in" value="0.0235" valueMax="0.026" valueMin="0.021">
|
||
<geoCoordinate box="[854,916,727,753]" degrees="0.021" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="55" value="0.021">0.021</geoCoordinate>
|
||
<geoCoordinate box="[916,994,727,753]" degrees="0.026" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="55" value="-0.026">–0.026</geoCoordinate>
|
||
in
|
||
</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1030,1207,727,753]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[1030,1207,727,753]" italics="true" pageId="9" pageNumber="10">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
|
||
<paragraph blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">
|
||
<emphasis bold="true" box="[199,533,762,789]" pageId="9" pageNumber="10">
|
||
Description of the
|
||
<typeStatus box="[424,527,763,789]" pageId="9" pageNumber="10">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female. L =
|
||
<quantity box="[688,785,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="1.9899999999999998" pageId="9" pageNumber="10" unit="mm" value="199.0">199 mm</quantity>
|
||
, SVL =
|
||
<quantity box="[882,1000,762,789]" metricMagnitude="-1" metricUnit="m" metricValue="1.935" pageId="9" pageNumber="10" unit="mm" value="193.5">193.5 mm</quantity>
|
||
, TL =
|
||
<quantity box="[1078,1168,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="10" unit="mm" value="5.5">5.5 mm</quantity>
|
||
, HW =
|
||
<quantity box="[1260,1363,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="9" pageNumber="10" unit="mm" value="10.1">10.1 mm</quantity>
|
||
, SN =
|
||
<quantity box="[151,241,798,825]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="10" unit="mm" value="5.5">5.5 mm</quantity>
|
||
, SW =
|
||
<quantity box="[327,417,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="9" pageNumber="10" unit="mm" value="8.6">8.6 mm</quantity>
|
||
, PSN =
|
||
<quantity box="[514,603,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="10" unit="mm" value="2.4">2.4 mm</quantity>
|
||
, RW =
|
||
<quantity box="[691,781,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="10" unit="mm" value="3.7">3.7 mm</quantity>
|
||
, EW =
|
||
<quantity box="[869,959,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="10" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, W =
|
||
<quantity box="[1029,1133,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="9" pageNumber="10" unit="mm" value="10.6">10.6 mm</quantity>
|
||
, VW =
|
||
<quantity box="[1224,1313,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="9" pageNumber="10" unit="mm" value="8.7">8.7 mm</quantity>
|
||
, TW =
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="9" pageNumber="10" unit="mm" value="7.2">7.2 mm</quantity>
|
||
, L/W = 39.0, TL/SVL = 0.028. Head slightly wider than neck. Snout rounded in dorsal and lateral views. Snout anterior to lower jaw horizontal, parallel to body axis. Rostral rather narrow (RW/HW = 0.36), oval dorsally, lateral margins convex, posterior border extending approximately half way between naris and eye, posterior margin straight; ventrally surface papillose, with sides slightly diverging anteriorly and posterior margin straight. Nasals separated dorsally by prefrontal (
|
||
<figureCitation box="[534,626,979,1005]" captionStart="FIGURE 4" captionStartId="10.[152,255,1935,1959]" captionTargetBox="[169,1419,363,1893]" captionTargetId="figure-122@10.[151,1436,345,1911]" captionTargetPageId="10" captionText="FIGURE 4. Dorsal and lateral views of heads for members of the Gerrhopilus inornatus Group having 22 midbody scale rows and uniformly dark venters. A, B holotype of Gerrhopilus papuanorum (BPBM 17236); C, D holotype of Gerrhopilus polyadenus (BMNH 1901.11.27.10)." figureDoi="http://doi.org/10.5281/zenodo.7571497" httpUri="https://zenodo.org/record/7571497/files/figure.png" pageId="9" pageNumber="10">Fig. 4C</figureCitation>
|
||
); superior nasal large, with slightly sinuous posterior margin, concave dorsally, convex ventrally (
|
||
<figureCitation box="[461,554,1015,1041]" captionStart="FIGURE 4" captionStartId="10.[152,255,1935,1959]" captionTargetBox="[169,1419,363,1893]" captionTargetId="figure-122@10.[151,1436,345,1911]" captionTargetPageId="10" captionText="FIGURE 4. Dorsal and lateral views of heads for members of the Gerrhopilus inornatus Group having 22 midbody scale rows and uniformly dark venters. A, B holotype of Gerrhopilus papuanorum (BPBM 17236); C, D holotype of Gerrhopilus polyadenus (BMNH 1901.11.27.10)." figureDoi="http://doi.org/10.5281/zenodo.7571497" httpUri="https://zenodo.org/record/7571497/files/figure.png" pageId="9" pageNumber="10">Fig. 4D</figureCitation>
|
||
). External naris semicircular, oriented obliquely, close to rostral, anterior half covered by inferior nasal; superior nasal suture extending anterodorsally from naris to rostral; inferior nasal suture complete, contacting second supralabial just posterior to latter’s contact with first supralabial. Prefrontal larger than supraoculars, which are larger than parietals and interparietal, which are subequal in size. Frontal fused with both parietals. Preocular large, triangular; larger than ocular but smaller than superior nasal. Ocular large, smaller than preocular, extending dorsally well above preocular, extending ventrally to ~2/3 depth of preocular, bordered posteroventrally by subocular of approximately two-thirds its size. Eye distinct, with pale pupil, situated at widest point of ocular and approximately midway along its height, anterior 1/4 (R) or 1/3 (L) covered by preocular plate in lateral view. Four postoculars bordering ocular and subocular between parietal and fourth supralabial. Four supralabials, third the largest, all except first with long axis oblique to long axis of body, first approximately square. Supralabial imbrication pattern T-V, posterior border of second supralabial overlaps anteroventral margin of preocular, that of third supralabial overlaps anteroventral margin of subocular and extends posterior to rear margin of preocular. Mental hexagonal, wider than long, projecting slightly beyond curve of lower jaw and fitting into notch on upper lip when mouth is closed. Infralabials two on each side, second much longer.
|
||
</paragraph>
|
||
<paragraph blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">
|
||
Longitudinal scale rows uniformly 22 behind head, at midbody, and anterior to vent; transverse scale rows between rostral and tail tip 320,
|
||
<specimenCount box="[518,678,1555,1581]" pageId="9" pageNumber="10" type="intercalary">six intercalary</specimenCount>
|
||
scales along vertebral row; subcaudals 16; dorsocaudals 16; apical region with blunt spine that barely extends past last scales.
|
||
</paragraph>
|
||
<paragraph blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">Rostral, nasals, and preoculars densely covered in large pale glands; oculars with 22 (R) and 23 (L) glands, preoculars with 50 (R) and 46 (L), supraoculars with 8 (R) and 9 (L), prefrontal with 29; frontal with 6, and suboculars with 16.</paragraph>
|
||
<paragraph blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">In preservative, ~121 years after preservation, dorsum faded medium brown, gradually fading laterally to paler brown on venter; no sharp distinction between dorsal and ventral coloration; each dorsal scale uniformly dark; each ventral scale slightly darker anteriorly. Anterior half of rostral, area around nares, first supralabial, mental, and center of throat to six scales behind mental pale straw yellow; second, third, and fourth supralabials pale straw yellow with brown along dorsal margins. Head glands pale straw yellow; tail spine and adjacent scales dark brown (darker than rest of body), spine with white tip. Iris dark gray; pupil pale gray.</paragraph>
|
||
<paragraph blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1436,150,321]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
<emphasis bold="true" box="[199,319,1950,1976]" pageId="9" pageNumber="10">Variation.</emphasis>
|
||
The
|
||
<typeStatus box="[379,475,1951,1977]" pageId="9" pageNumber="10">paratype</typeStatus>
|
||
is an immature female and differs from the
|
||
<typeStatus box="[983,1081,1951,1977]" pageId="9" pageNumber="10">holotype</typeStatus>
|
||
in being shorter (L =
|
||
<quantity box="[1335,1432,1951,1977]" metricMagnitude="-1" metricUnit="m" metricValue="1.38" pageId="9" pageNumber="10" unit="mm" value="138.0">138 mm</quantity>
|
||
, SVL =
|
||
<quantity box="[235,331,1987,2013]" metricMagnitude="-1" metricUnit="m" metricValue="1.33" pageId="9" pageNumber="10" unit="mm" value="133.0">133 mm</quantity>
|
||
, TL =
|
||
<quantity box="[408,476,1986,2013]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="mm" value="5.0">5 mm</quantity>
|
||
,
|
||
<tableCitation box="[489,571,1987,2013]" captionStart="TABLE 1" captionStartId="17.[151,244,552,578]" captionTargetPageId="17" captionText="TABLE 1. Mensural and count data for members of the Gerrhopilus inornatus Group. Forward slashes separate counts on the right and left sides, respectively." pageId="9" pageNumber="10">Table 1</tableCitation>
|
||
), the same relative width (W =
|
||
<quantity box="[936,1025,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="10" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, L/W = 36.3), and with a longer tail (TL/SVL = 0.038). It has 294 transverse scale rows, 15 subcaudals and dorsocaudals, and fewer head glands than the
|
||
<typeStatus box="[192,293,187,213]" pageId="10" pageNumber="11">holotype</typeStatus>
|
||
, with 11 (R) and 16 (L) glands in the oculars, 42 (R) and 43 (L) in the preoculars, 5 (R) and 6 (L) in the supraoculars,
|
||
<quantity box="[306,361,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="10" pageNumber="11" unit="in" value="11.0">11 in</quantity>
|
||
the prefrontal, 0 in the frontal, and 5 (R) and 8 (L) in the subocular. It has a sharp, protruding tail spine that is longer than that in the
|
||
<typeStatus box="[549,651,259,285]" pageId="10" pageNumber="11">holotype</typeStatus>
|
||
; base of spine and adjacent scales dark brown, but the tip of the spine is corneous brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571497" ID-Zenodo-Dep="7571497" httpUri="https://zenodo.org/record/7571497/files/figure.png" pageId="10" pageNumber="11" startId="10.[152,255,1935,1959]" targetBox="[169,1419,363,1893]" targetPageId="10">
|
||
<paragraph blockId="10.[151,1437,1935,2031]" pageId="10" pageNumber="11">
|
||
<emphasis bold="true" box="[152,283,1935,1959]" pageId="10" pageNumber="11">FIGURE 4.</emphasis>
|
||
Dorsal and lateral views of heads for members of the
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[862,1091,1935,1959]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[862,1091,1935,1959]" italics="true" pageId="10" pageNumber="11">Gerrhopilus inornatus</emphasis>
|
||
</taxonomicName>
|
||
Group having 22 midbody scale rows and uniformly dark venters.
|
||
<emphasis bold="true" box="[503,552,1971,1995]" pageId="10" pageNumber="11">A, B</emphasis>
|
||
holotype of
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[686,946,1971,1995]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[686,946,1971,1995]" italics="true" pageId="10" pageNumber="11">Gerrhopilus papuanorum</emphasis>
|
||
</taxonomicName>
|
||
(BPBM 17236);
|
||
<emphasis bold="true" box="[1128,1179,1971,1995]" pageId="10" pageNumber="11">C, D</emphasis>
|
||
holotype of
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="polyadenus">
|
||
<emphasis italics="true" pageId="10" pageNumber="11">Gerrhopilus polyadenus</emphasis>
|
||
</taxonomicName>
|
||
(BMNH 1901.11.27.10).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="11" pageNumber="12" type="etymology">
|
||
<paragraph blockId="11.[151,1437,151,1221]" pageId="11" pageNumber="12">
|
||
<emphasis bold="true" box="[199,334,151,177]" pageId="11" pageNumber="12">Etymology.</emphasis>
|
||
The specific epithet is a masculine Latinized compound adjective, from the Greek
|
||
<emphasis box="[1268,1326,151,177]" italics="true" pageId="11" pageNumber="12">poly-</emphasis>
|
||
, meaning
|
||
<emphasis box="[151,212,187,212]" italics="true" pageId="11" pageNumber="12">many</emphasis>
|
||
, and
|
||
<emphasis box="[273,341,187,213]" italics="true" pageId="11" pageNumber="12">adeno</emphasis>
|
||
, meaning
|
||
<emphasis box="[459,523,187,213]" italics="true" pageId="11" pageNumber="12">gland</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="12" type="distribution">
|
||
<paragraph blockId="11.[151,1437,151,1221]" pageId="11" pageNumber="12">
|
||
<emphasis bold="true" box="[199,353,222,248]" pageId="11" pageNumber="12">Distribution.</emphasis>
|
||
Known from the
|
||
<typeStatus box="[565,613,223,249]" pageId="11" pageNumber="12">type</typeStatus>
|
||
locality at an unspecified location on Mt Albert Edward and from the nearby lowlands of
|
||
<collectingRegion box="[373,525,259,285]" country="Papua New Guinea" name="Northern" pageId="11" pageNumber="12">Oro Province</collectingRegion>
|
||
,
|
||
<collectingCountry box="[537,753,259,285]" name="Papua New Guinea" pageId="11" pageNumber="12">Papua New Guinea</collectingCountry>
|
||
(
|
||
<figureCitation box="[768,838,259,285]" captionStart="FIGURE 2" captionStartId="5.[152,255,1069,1093]" captionTargetBox="[181,1408,456,1045]" captionTargetId="figure-148@5.[179,1408,453,1045]" captionTargetPageId="5" captionText="FIGURE 2. Map of southern Papua New Guinea showing collecting localities for Gerrhopilus inornatus (star), G. flavinotatus (diamond), G. lorealis (hexagon), G. papuanorum (square), G. polyadenus (open circles), G. slapcinskyi (filled circle), and G. wallachi (triangle)." figureDoi="http://doi.org/10.5281/zenodo.7571491" httpUri="https://zenodo.org/record/7571491/files/figure.png" pageId="11" pageNumber="12">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="12" type="discussion">
|
||
<paragraph blockId="11.[151,1437,151,1221]" pageId="11" pageNumber="12">
|
||
<emphasis bold="true" box="[199,318,295,321]" pageId="11" pageNumber="12">Remarks.</emphasis>
|
||
The BMNH catalogue for the
|
||
<typeStatus box="[699,800,295,321]" pageId="11" pageNumber="12">holotype</typeStatus>
|
||
states that it was purchased from Mr. H.S. Rohu of Stepney, a district on the east end of London. This catalogue states that the specimen came from “not less than
|
||
<quantity box="[151,242,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="11" pageNumber="12" unit="ft" value="6000.0">6000 ft.</quantity>
|
||
” on the Albert Edward Range, but this information is doubtful for two reasons. First,
|
||
<bibRefCitation author="Wichmann, A." pageId="11" pageNumber="12" pagination="371 - 1026" refId="ref19812" refString="Wichmann, A. (1912) Resultats de l'expedition scientifique Neerlandaise la Nouvelle-Guinee en 1903 sous les auspices de Arthur Wichmann, vol. II, 2 eme partie, Entdeckungsgeschichte von Neu-Guinea (1885 bis 1902). Nova Guinea, 2, 371 - 1026." type="journal article" year="1912">Wichmann (1912: 754)</bibRefCitation>
|
||
states that Rohu’s expedition up Mt Albert Edward reached no farther than a height of
|
||
<quantity box="[1208,1298,403,429]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="11" pageNumber="12" unit="ft" value="6000.0">6000 ft.</quantity>
|
||
(“...dort bis in die Höhe von
|
||
<quantity box="[347,455,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="11" pageNumber="12" unit="ft" value="6000.0">6000 feet</quantity>
|
||
gelangte.). Wichmann does not state the source of his information, but he may have been in direct correspondence with Rohu on the matter. Rohu was an overseer on the Tamata Station, along the Mambare River (
|
||
<bibRefCitation author="Administrator & British New Guinea" box="[351,833,511,537]" pageId="11" pageNumber="12" refId="ref18844" refString="Administrator, British New Guinea. (1902) Annual Report on British New Guinea from 1 st July, 1900, to 30 th June, 1901; with appendices. Government of the Commonwealth of Australia, Brisbane, xxx + 134 pp." type="book" year="1902">
|
||
Administrator, British New
|
||
<collectingCountry box="[680,763,511,537]" name="Guinea-Bissau" pageId="11" pageNumber="12">Guinea</collectingCountry>
|
||
1902
|
||
</bibRefCitation>
|
||
), which lies on the northern versant in what is now
|
||
<collectingRegion box="[151,309,547,573]" country="Papua New Guinea" name="Northern" pageId="11" pageNumber="12">Oro Province</collectingRegion>
|
||
,
|
||
<collectingCountry box="[320,381,547,573]" name="Papua New Guinea" pageId="11" pageNumber="12">PNG</collectingCountry>
|
||
. He quit service at Tamata Station on 18 October, 1900, and his expedition would thus have occurred in late 1900 or the first part of
|
||
<quantity box="[623,711,583,609]" metricMagnitude="1" metricUnit="m" metricValue="4.828539999999999" pageId="11" pageNumber="12" unit="in" value="1901.0">1901 in</quantity>
|
||
order for the specimens to have reached BMNH in time to be catalogued in late November, 1901. Mt Albert Edward is one of the higher peaks of the Owen Stanley Range; it now lies largely within Central Province,
|
||
<collectingCountry box="[661,722,655,681]" name="Papua New Guinea" pageId="11" pageNumber="12">PNG</collectingCountry>
|
||
, but straddles the division between the northern and southern versants of the Owen Stanley Mts. Given Rohu’s residence at Tamata Station, I presume he made his ascent from the east (
|
||
<collectingRegion box="[253,414,727,753]" country="Papua New Guinea" name="Northern" pageId="11" pageNumber="12">Oro Province</collectingRegion>
|
||
), which is consistent with the
|
||
<typeStatus box="[759,858,727,753]" pageId="11" pageNumber="12">paratype</typeStatus>
|
||
being from Sangara,
|
||
<quantity box="[1107,1223,726,753]" metricMagnitude="4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="11" pageNumber="12" unit="km" value="55.0" valueMax="60.0" valueMin="50.0">50–60 km</quantity>
|
||
S of the Mambare River in the same province. Inasmuch as the
|
||
<typeStatus box="[678,777,763,789]" pageId="11" pageNumber="12">paratype</typeStatus>
|
||
came from an elevation of approximately
|
||
<quantity box="[1275,1349,763,789]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="12" unit="m" value="120.0">120 m</quantity>
|
||
a.s.l., I presume that the
|
||
<typeStatus box="[352,453,799,825]" pageId="11" pageNumber="12">holotype</typeStatus>
|
||
did not, in fact, come from as high as “not less than”
|
||
<quantity box="[1085,1176,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="11" pageNumber="12" unit="ft" value="6000.0">6000 ft.</quantity>
|
||
(
|
||
<quantity box="[1192,1285,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="11" pageNumber="12" unit="m" value="1830.0">1830 m</quantity>
|
||
), as stated in the BMNH catalogue, and that
|
||
<bibRefCitation author="Wichmann, A." box="[513,742,835,861]" pageId="11" pageNumber="12" pagination="371 - 1026" refId="ref19812" refString="Wichmann, A. (1912) Resultats de l'expedition scientifique Neerlandaise la Nouvelle-Guinee en 1903 sous les auspices de Arthur Wichmann, vol. II, 2 eme partie, Entdeckungsgeschichte von Neu-Guinea (1885 bis 1902). Nova Guinea, 2, 371 - 1026." type="journal article" year="1912">Wichmann’s (1912)</bibRefCitation>
|
||
account is correct. Second, the BMNH accession of Rohu’s herpetological material includes
|
||
<specimenCount box="[533,691,871,897]" pageId="11" pageNumber="12" type="generic">24 specimens</specimenCount>
|
||
of 15 species. Of these, at least four (e.g.,
|
||
<taxonomicName baseAuthorityName="Meyer" baseAuthorityYear="1875" box="[1199,1429,871,897]" class="Amphibia" family="Ceratobatrachidae" genus="Cornufer" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="papuensis">
|
||
<emphasis box="[1199,1429,871,897]" italics="true" pageId="11" pageNumber="12">Cornufer papuensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Kopstein" baseAuthorityYear="1926" box="[151,302,907,933]" class="Reptilia" family="Scincidae" genus="Emoia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="jakati">
|
||
<emphasis box="[151,302,907,933]" italics="true" pageId="11" pageNumber="12">Emoia jakati</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[318,495,907,933]" class="Squamata" family="Scincidae" genus="Palaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptilia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pulchra">
|
||
<emphasis box="[318,495,907,933]" italics="true" pageId="11" pageNumber="12">Palaia pulchra</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Oudemans" baseAuthorityYear="1894" box="[511,771,907,933]" class="Reptilia" family="Scincidae" genus="Prasinohaema" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="semoni">
|
||
<emphasis box="[511,771,907,933]" italics="true" pageId="11" pageNumber="12">Prasinohaema semoni</emphasis>
|
||
</taxonomicName>
|
||
) are lowland species that are not found as high as
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="12" unit="m" value="1800.0">1800 m</quantity>
|
||
a.s.l., much less occurring “not less than that elevation”. Several of the other species are also predominantly lowland inhabitants though they may attain elevations as high as
|
||
<quantity box="[936,1026,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="12" unit="m" value="1800.0">1800 m</quantity>
|
||
a.s.l. Hence, the exact elevational provenance of the
|
||
<typeStatus box="[362,463,1015,1041]" pageId="11" pageNumber="12">holotype</typeStatus>
|
||
is uncertain, but it seems to have been collected at a relatively low elevation.
|
||
<bibRefCitation author="Zweifel, R. G." pageId="11" pageNumber="12" pagination="1 - 21" refId="ref19914" refString="Zweifel, R. G. (1979) Variation in the scincid lizard Lipinia noctua and notes on other Lipinia from the New Guinea region. American Museum Novitates, 2676, 1 - 21. [http: // hdl. handle. net / 2246 / 5434]" type="journal article" year="1979">Zweifel (1979)</bibRefCitation>
|
||
makes this same inference that the reported elevation for the
|
||
<typeStatus box="[959,1060,1051,1077]" pageId="11" pageNumber="12">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[1102,1278,1051,1077]" class="Squamata" family="Scincidae" genus="Palaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptilia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pulchra">
|
||
<emphasis box="[1102,1278,1051,1077]" italics="true" pageId="11" pageNumber="12">Palaia pulchra</emphasis>
|
||
</taxonomicName>
|
||
, collected by Rohu on the same expedition, is erroneous.
|
||
</paragraph>
|
||
<paragraph blockId="11.[151,1437,151,1221]" pageId="11" pageNumber="12">
|
||
The
|
||
<typeStatus box="[254,352,1123,1149]" pageId="11" pageNumber="12">holotype</typeStatus>
|
||
of this species was examined and referred to
|
||
<taxonomicName authorityName="de Rooij" authorityYear="1917" box="[905,1124,1123,1149]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[905,1124,1123,1149]" italics="true" pageId="11" pageNumber="12">Typhlops inornatus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="de Rooij, N." box="[1176,1364,1123,1149]" pageId="11" pageNumber="12" refId="ref18970" refString="de Rooij, N. (1917) The reptiles of the Indo-Australian Archipelago, II: Ophidia. E. J. Brill, Leiden, 334 pp." type="book" year="1917">de Rooij (1917)</bibRefCitation>
|
||
, who, however, did not remark on the differences between this specimen and the
|
||
<typeStatus box="[996,1094,1159,1185]" pageId="11" pageNumber="12">holotype</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="de Rooij" authorityYear="1917" box="[1132,1274,1159,1185]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[1132,1274,1159,1185]" italics="true" pageId="11" pageNumber="12">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus box="[1340,1436,1159,1185]" pageId="11" pageNumber="12">paratype</typeStatus>
|
||
was examined by
|
||
<bibRefCitation author="Wallach, V." box="[352,525,1195,1221]" pageId="11" pageNumber="12" pagination="107 - 118" refId="ref19774" refString="Wallach, V. (1996) Two new blind snakes of the Typhlops ater species group from Papua New Guinea (Serpentes: Typhlopidae). Russian Journal of Herpetology, 3, 107 - 118." type="journal article" year="1996">Wallach (1996)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Pyron, R. A. & Wallach, V." box="[579,857,1195,1221]" pageId="11" pageNumber="12" pagination="1 - 81" refId="ref19325" refString="Pyron, R. A. & Wallach, V. (2014) Systematics of the blind snakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence. Zootaxa, 3829 (1), 1 - 81. https: // doi. org / 10.11646 / zootaxa. 3829.1.1" type="journal article" year="2014">Pyron & Wallach (2014)</bibRefCitation>
|
||
, who listed it as
|
||
<taxonomicName authorityName="de Rooij" authorityYear="1917" box="[1044,1258,1195,1221]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[1044,1258,1195,1221]" italics="true" pageId="11" pageNumber="12">Typhlops inornatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |