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<mods:title id="17C1895441FC0B9AC55A72F65C11F5F0">Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983</mods:title>
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<heading id="D0BE6F55BC30FFF5FC8EECEBD777FB77" box="[883,1344,1200,1221]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="1305" reason="8">
GENUS
<taxonomicName id="4C49A3BABC30FFF5FC31ECEBD777FB77" authority="LEINDERS, 1983" authorityName="LEINDERS" authorityYear="1983" box="[972,1344,1200,1221]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="1305" phylum="Chordata" rank="genus">
<emphasis id="B93D042BBC30FFF5FC31ECEBD64BFB76" box="[972,1148,1200,1221]" italics="true" pageId="1" pageNumber="1305">HOPLITOMERYX</emphasis>
<bibRefCitation id="EFD8A5C8BC30FFF5FB7AECEBD777FB77" author="Leinders JJM" box="[1159,1344,1200,1221]" pageId="1" pageNumber="1305" pagination="1 - 68" refId="ref19702" refString="Leinders JJM. 1983. Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Part. 1: the cranial osteology of Hoplitomeryx gen. nov. and discussion on the classification of pecoran families. Scripta Geologica 70: 1 - 68." type="journal article" year="1983">LEINDERS, 1983</bibRefCitation>
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</paragraph>
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<subSubSection id="C3538BB2BC30FFF6FCDBEC8DD3A8FC2E" lastPageId="2" lastPageNumber="1306" pageId="1" pageNumber="1305" type="diagnosis">
<paragraph id="8BF6D839BC30FFF5FCDBEC8DD1B1FA70" blockId="1.[806,1422,1238,1903]" pageId="1" pageNumber="1305">
<emphasis id="B93D042BBC30FFF5FCDBEC8DD1C3FB59" box="[806,1012,1238,1259]" italics="true" pageId="1" pageNumber="1305">Revised diagnosis</emphasis>
: The genus
<taxonomicName id="4C49A3BABC30FFF5FB76EC8DD713FB59" authorityName="LEINDERS" authorityYear="1983" box="[1163,1316,1238,1259]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="1305" phylum="Chordata" rank="genus">
<emphasis id="B93D042BBC30FFF5FB76EC8DD713FB59" box="[1163,1316,1238,1259]" italics="true" pageId="1" pageNumber="1305">Hoplitomeryx</emphasis>
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includes species with two paired, nondeciduous, unbranched orbital horns, with a median, unpaired, nondeciduous, and unbranched nasal horn, and with sabre-like upper canines. The skull roof is pneumatized, there are two lacrimal orifices on the orbital rim, and large and smooth-surfaced auditory bullae. The jawbone is sinuous.
</paragraph>
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Dentally the genus is characterized by brachyodont to mesodont dentitions; p1 is always absent, p2 is present only in the more primitive Scontrone specimens, whereas it is lost in the specimens from Gargano: the upper and lower cheek dental formulae are therefore 3
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the Scontrone specimens,
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the Gargano ones. p4 is not molarized, generally simple, either elongated or compressed mesiodistally.
</paragraph>
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More specifically,
<taxonomicName id="4C49A3BABC30FFF5FBF1EE9BD692F967" authorityName="LEINDERS" authorityYear="1983" box="[1036,1189,1728,1749]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="1305" phylum="Chordata" rank="genus">
<emphasis id="B93D042BBC30FFF5FBF1EE9BD692F967" box="[1036,1189,1728,1749]" italics="true" pageId="1" pageNumber="1305">Hoplitomeryx</emphasis>
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is characterized by: (1) cheek teeth brachyodont or mesodont; (2) upper molars often wider than they are long; (3) variously developed parastyle and mesostyle in upper cheek teeth; (4) P3 with centrally situated and lingually directed protocone; (5) P4 and M3 with large metaconule; (6) incipient entostyle occasionally from mesial wall of metaconule of first and second upper molars; (7) cingula absent in upper cheek teeth; (8) small paraconid occasionally present; (9) nonmolarized p4; (10) metastylid generally well developed in lower molars; (11) ectostylid occasionally present, sometimes quite developed, on lower molars; (12) complete postentocristid in lower molars; (13) infrequent mesiolabial cingulum; (14) distal lobe of m3 constantly bi-lophed, normally constituted by crescent hypoconulid and large, tear-shaped entoconulid, both with distally extending cristids, not fused together along the distal margin of the tooth or otherwise fused, but only in fairly worn teeth; (14) mandible slender or pachyostotic. Sabre-like upper canines can sometimes be present. There is no
<taxonomicName id="4C49A3BABC33FFF6FDCBEACAD0E3FD14" box="[566,724,657,678]" class="Mammalia" family="Tragulidae" genus="Dorcatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="1306" phylum="Chordata" rank="genus">
<emphasis id="B93D042BBC33FFF6FDCBEACAD0E3FD14" box="[566,724,657,678]" italics="true" pageId="2" pageNumber="1306">Dorcatherium</emphasis>
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fold on lower molars, nor any
<taxonomicName id="4C49A3BABC33FFF6FDF0EAF4D0AAFD76" authorityName="Von Meyer" authorityYear="1851" box="[525,669,687,708]" class="Mammalia" family="Palaeomerycidae" genus="Palaeomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="1306" phylum="Chordata" rank="genus">
<emphasis id="B93D042BBC33FFF6FDF0EAF4D0AAFD76" box="[525,669,687,708]" italics="true" pageId="2" pageNumber="1306">Palaeomeryx</emphasis>
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fold in brachyodont species of the genus. Postcranially, the genus is characterized by patellae with elongated distal apophyses probably as a result of ossification of the ligamentum patellae, astragalus with trochleae not parallel, navicocuboid fused to the metatarsus, distally closed metatarsal gullies, cannon bones with complete distal keels.
</paragraph>
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