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<document id="6D85D13C0993E1DA513C3094D3B24471" ID-DOI="10.11646/zootaxa.5005.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5141485" ID-ZooBank="86F985AB-BB8D-474A-8A70-19103C316174" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627465624485" checkinUser="plazi" docAuthor="Mawadda, N. V. Ayisha, Kumar, P. Girish &amp; Sureshan, P. M." docDate="2021" docId="03D49323FFF7FF9019C2FB37FD32FEB0" docLanguage="en" docName="zootaxa.5005.2.6.pdf" docOrigin="Zootaxa 5005 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyroda binghami Tsuneki 1983" docType="treatment" docVersion="3" lastPageNumber="215" masterDocId="FFEDEB5BFFF0FF9E1955FF92FFA7FFAD" masterDocTitle="Additions to the knowledge of the genus Lyroda Say (Hymenoptera: Crabronidae) from India with the descriptions of two new species" masterLastPageNumber="217" masterPageNumber="201" pageNumber="208" updateTime="1699025541757" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="BED1000FDA690B39C5E0660245BA1967">
<mods:title id="3B1602E5D45E7584F1AC4E4ABB6779F5">Additions to the knowledge of the genus Lyroda Say (Hymenoptera: Crabronidae) from India with the descriptions of two new species</mods:title>
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<mods:namePart id="9633D3FDB249EA17EF1E1EBAB85088F0">Mawadda, N. V. Ayisha</mods:namePart>
<mods:nameIdentifier id="5CDA4638A263F8442904BEB0DACFF356" type="ORCID">0000-0003-4649-9428</mods:nameIdentifier>
<mods:affiliation id="5F728799D6F30733EA0080B16C84953D">ayishakmr @ gmail. com; https: // orcid. org / 0000 - 0003 - 4649 - 9428</mods:affiliation>
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<mods:namePart id="A97628ED76063B79399894B18F3A0707">Kumar, P. Girish</mods:namePart>
<mods:nameIdentifier id="E4BA876D4A082528425F98E66B1D7B84" type="ORCID">0000-0003-2121-0165</mods:nameIdentifier>
<mods:affiliation id="F356E2E48213E6CDA8A48AD03BE238C2">kpgiris @ gmail. com; https: // orcid. org / 0000 - 0003 - 2121 - 0165</mods:affiliation>
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<mods:namePart id="2F71364E060852CB4783A7E9E314BBFF">Sureshan, P. M.</mods:namePart>
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<mods:affiliation id="C2DB0EA3CC8F4501F6A03ACA5526BE15">pmsuresh 43 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4722 - 6368</mods:affiliation>
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<mods:title id="828BC474CB5D2FCF2ED5D0F7FCD93B04">Zootaxa</mods:title>
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<mods:date id="159555F930C2532E728B28EE64A14847">2021</mods:date>
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<treatment id="03D49323FFF7FF9019C2FB37FD32FEB0" ID-DOI="http://doi.org/10.5281/zenodo.5151753" ID-Zenodo-Dep="5151753" LSID="urn:lsid:plazi:treatment:03D49323FFF7FF9019C2FB37FD32FEB0" httpUri="http://treatment.plazi.org/id/03D49323FFF7FF9019C2FB37FD32FEB0" lastPageId="14" lastPageNumber="215" pageId="7" pageNumber="208">
<subSubSection id="C36771BEFFF7FF9919C2FB37FDBDFB12" box="[151,538,1189,1216]" pageId="7" pageNumber="208" type="nomenclature">
<paragraph id="8BC22235FFF7FF9919C2FB37FDBDFB12" blockId="7.[151,538,1189,1251]" box="[151,538,1189,1216]" pageId="7" pageNumber="208">
<heading id="D08A9559FFF7FF9919C2FB37FDBDFB12" bold="true" box="[151,538,1189,1216]" fontSize="11" level="1" pageId="7" pageNumber="208" reason="1">
<taxonomicName id="4C7D59B6FFF7FF9919C2FB37FDBDFB12" ID-CoL="3WTNP" authority="Tsuneki, 1983" authorityName="Tsuneki" authorityYear="1983" box="[151,538,1189,1216]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="binghami">
<emphasis id="B909FE27FFF7FF9919C2FB37FDBDFB12" bold="true" box="[151,538,1189,1216]" pageId="7" pageNumber="208">
<emphasis id="B909FE27FFF7FF9919C2FB37FECFFB6D" bold="true" box="[151,360,1189,1216]" italics="true" pageId="7" pageNumber="208">Lyroda binghami</emphasis>
Tsuneki, 1983
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF9919C2FB5BFE80FB4E" box="[151,295,1225,1251]" pageId="7" pageNumber="208" type="description">
<paragraph id="8BC22235FFF7FF9919C2FB5BFE80FB4E" blockId="7.[151,538,1189,1251]" box="[151,295,1225,1251]" pageId="7" pageNumber="208">
(
<figureCitation id="13463EB0FFF7FF9919CAFB5BFEB8FB4E" box="[159,287,1225,1251]" captionStart-0="FIGURES 28–36" captionStart-1="FIGURES 37–43" captionStartId-0="9.[151,264,1943,1967]" captionStartId-1="10.[151,264,1943,1967]" captionTargetBox-0="[195,1395,192,1910]" captionTargetBox-1="[190,1397,191,1911]" captionTargetId-0="figure-18@9.[182,1406,181,1919]" captionTargetId-1="figure-17@10.[181,1405,181,1920]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 28–36. Lyroda binghami Tsuneki. ♀. 28. Habitus, lateral view; 29. Head, frontal view; 30. Lower half of head, frontal view; 31. Head &amp; thorax; 32. Propodeum, dorsal view; 33. Propodeum, lateral view; 34. Forewing; 35. Gaster, dorsal view; 36. Pygidium." captionText-1="FIGURES 37–43. Lyroda binghami Tsuneki. ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Lower half of head, fron- tal view; 40. Head &amp; thorax; 41. Propodeum, dorsal view; 42. Pygidium; 43. Last sternum." figureDoi-0="http://doi.org/10.5281/zenodo.5141497" figureDoi-1="http://doi.org/10.5281/zenodo.5141499" httpUri-0="https://zenodo.org/record/5141497/files/figure.png" httpUri-1="https://zenodo.org/record/5141499/files/figure.png" pageId="7" pageNumber="208">Figs 28–41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF9919C2FA9CFB2CFA8B" box="[151,1163,1294,1318]" pageId="7" pageNumber="208" type="reference_group">
<paragraph id="8BC22235FFF7FF9919C2FA9CFB2CFA8B" blockId="7.[151,1163,1294,1318]" box="[151,1163,1294,1318]" pageId="7" pageNumber="208">
<treatmentCitationGroup id="AB6D051BFFF7FF9919C2FA9CFB2CFA8B" box="[151,1163,1294,1318]" pageId="7" pageNumber="208">
<treatmentCitation id="0ADC0424FFF7FF9919C2FA9CFDAFFA8B" author="Tsuneki, K." box="[151,520,1294,1318]" page="73" pageId="7" pageNumber="208" year="1983">
<taxonomicName id="4C7D59B6FFF7FF9919C2FA9CFDAFFA8B" ID-CoL="3WTNP" authority="Tsuneki, 1983 a: 73" authorityName="Tsuneki" authorityPageNumber="73" authorityYear="1983" box="[151,520,1294,1318]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="binghami">
<emphasis id="B909FE27FFF7FF9919C2FA9CFEE2FA8B" box="[151,325,1294,1318]" italics="true" pageId="7" pageNumber="208">Lyroda binghami</emphasis>
<materialsCitation id="3B152868FFF7FF99181EFA9CFDAFFA8B" box="[331,520,1294,1318]" collectionCode="NHM" country="Myanmar" location="Tenasserim" municipality="Shwegyin" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<bibRefCitation id="EFEC5FC4FFF7FF99181EFA9CFDAFFA8B" author="Tsuneki, K." box="[331,520,1294,1318]" pageId="7" pageNumber="208" pagination="69 - 81" refId="ref11594" refString="Tsuneki, K. (1983 a) Further studies on the Larrinae of the Philippine Islands, with remarks on the Indian species of the genus Lyroda, (Hymenoptera, Sphecidae). Special Publications of the Japan Hymenopterists Association, 24, 69 - 81 + 112 - 117." type="journal article" year="1983">Tsuneki, 1983a:73</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="9D7BE9BCFFF7FF991B47FA9CFD82FA88" box="[530,549,1294,1317]" pageId="7" pageNumber="208" type="female">♀</specimenCount>
,
<specimenCount id="9D7BE9BCFFF7FF991B67FA9CFDEFFA88" box="[562,584,1294,1317]" pageId="7" pageNumber="208" type="male">♂</specimenCount>
.
<typeStatus id="54C69C97FFF7FF991B04FA9CFD14FA8B" box="[593,691,1294,1318]" pageId="7" pageNumber="208" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D7BE9BCFFF7FF991BE8FA9CFD77FA88" box="[701,720,1294,1317]" pageId="7" pageNumber="208" type="female">♀</specimenCount>
,
<collectingCountry id="F36A62A5FFF7FF991B88FA9CFCE3FA8B" box="[733,836,1294,1318]" name="Myanmar" pageId="7" pageNumber="208">Myanmar</collectingCountry>
:
<location id="8EA274EEFFF7FF991A19FA9CFC64FA8B" LSID="urn:lsid:plazi:treatment:03D49323FFF7FF9019C2FB37FD32FEB0:8EA274EEFFF7FF991A19FA9CFC64FA8B" box="[844,963,1294,1318]" country="Myanmar" municipality="Shwegyin" name="Tenasserim" pageId="7" pageNumber="208">Tenasserim</location>
:
<collectingMunicipality id="6BA6B84FFFF7FF991A99FA9CFB96FA8B" box="[972,1073,1294,1318]" pageId="7" pageNumber="208">Shwegyin</collectingMunicipality>
(NHM).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF9919C2FAC6FC9AF8EF" pageId="7" pageNumber="208" type="diagnosis">
<paragraph id="8BC22235FFF7FF9919C2FAC6FC9AF8EF" blockId="7.[151,1437,1364,2038]" pageId="7" pageNumber="208">
<emphasis id="B909FE27FFF7FF9919C2FAC6FEB5FAC3" bold="true" box="[151,274,1364,1390]" pageId="7" pageNumber="208">Diagnosis.</emphasis>
FEMALE. BL.
<quantity id="4C858FD0FFF7FF991893FAC6FDCBFAC2" box="[454,620,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="9.875" metricValueMax="9.9" metricValueMin="9.85" pageId="7" pageNumber="208" unit="mm" value="9.875" valueMax="9.9" valueMin="9.85">9.85–9.90 mm</quantity>
. Black, tibio-tarsal articulation and all tarsi slightly ferruginous (
<figureCitation id="13463EB0FFF7FF991C69FAC6FA37FAC3" box="[1340,1424,1364,1390]" captionStart="FIGURES 28–36" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[195,1395,192,1910]" captionTargetId="figure-18@9.[182,1406,181,1919]" captionTargetPageId="9" captionText="FIGURES 28–36. Lyroda binghami Tsuneki. ♀. 28. Habitus, lateral view; 29. Head, frontal view; 30. Lower half of head, frontal view; 31. Head &amp; thorax; 32. Propodeum, dorsal view; 33. Propodeum, lateral view; 34. Forewing; 35. Gaster, dorsal view; 36. Pygidium." figureDoi="http://doi.org/10.5281/zenodo.5141497" httpUri="https://zenodo.org/record/5141497/files/figure.png" pageId="7" pageNumber="208">Fig. 28</figureCitation>
). Setae on lower frons and clypeus, behind eyes on vertex, pronotal collar, posterior and posterolateral sides of scutum, sides of scutellum, metanotum, all sides of propodeum, and sternal areas of thorax silvery, silvery pile bands present at apical portion of Gt1–Gt3. Gt4, Gs4, Gt5, and Gs5 on posterior portion and Gs6 extensively covered with dense golden velvety pile (
<figureCitation id="13463EB0FFF7FF9918AEFA76FDE9FA52" box="[507,590,1508,1535]" captionStart="FIGURES 28–36" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[195,1395,192,1910]" captionTargetId="figure-18@9.[182,1406,181,1919]" captionTargetPageId="9" captionText="FIGURES 28–36. Lyroda binghami Tsuneki. ♀. 28. Habitus, lateral view; 29. Head, frontal view; 30. Lower half of head, frontal view; 31. Head &amp; thorax; 32. Propodeum, dorsal view; 33. Propodeum, lateral view; 34. Forewing; 35. Gaster, dorsal view; 36. Pygidium." figureDoi="http://doi.org/10.5281/zenodo.5141497" httpUri="https://zenodo.org/record/5141497/files/figure.png" pageId="7" pageNumber="208">Fig. 35</figureCitation>
). Anterior margin of clypeus usually with one medial and two lateral sets of three teeth in female, accompanied by two or three teeth between median and lateral sets (
<figureCitation id="13463EB0FFF7FF991DD4F99AFB72F98F" box="[1153,1237,1544,1570]" captionStart="FIGURES 28–36" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[195,1395,192,1910]" captionTargetId="figure-18@9.[182,1406,181,1919]" captionTargetPageId="9" captionText="FIGURES 28–36. Lyroda binghami Tsuneki. ♀. 28. Habitus, lateral view; 29. Head, frontal view; 30. Lower half of head, frontal view; 31. Head &amp; thorax; 32. Propodeum, dorsal view; 33. Propodeum, lateral view; 34. Forewing; 35. Gaster, dorsal view; 36. Pygidium." figureDoi="http://doi.org/10.5281/zenodo.5141497" httpUri="https://zenodo.org/record/5141497/files/figure.png" pageId="7" pageNumber="208">Figs 30</figureCitation>
&amp;
<figureCitation id="13463EB0FFF7FF991DADF99AFAB0F98E" box="[1272,1303,1544,1571]" captionStart="FIGURES 58–64" captionStartId="13.[151,264,1942,1966]" captionTargetBox="[181,1400,189,1912]" captionTargetId="figure-18@13.[178,1409,181,1918]" captionTargetPageId="13" captionText="FIGURES 58–64. Anterior clypeal margins: 58. Lyroda aurea sp. nov. holotype ♀; 59. Lyroda nuda sp. nov. holotype ♀; 60. Lyroda salai Giner Marí. ♀; 61. Lyroda binghami Tsuneki. ♀; 62. Lyroda binghami Tsuneki. ♂; 63. Lyroda formosa (Smith). ♀; 64. Lyroda formosa (Smith). ♂." figureDoi="http://doi.org/10.5281/zenodo.5141505" httpUri="https://zenodo.org/record/5141505/files/figure.png" pageId="7" pageNumber="208">61</figureCitation>
). Propodeal dorsum coarsely rugoso-reticulate (
<figureCitation id="13463EB0FFF7FF991B70F9BEFDD0F9EA" box="[549,631,1580,1607]" captionStart="FIGURES 28–36" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[195,1395,192,1910]" captionTargetId="figure-18@9.[182,1406,181,1919]" captionTargetPageId="9" captionText="FIGURES 28–36. Lyroda binghami Tsuneki. ♀. 28. Habitus, lateral view; 29. Head, frontal view; 30. Lower half of head, frontal view; 31. Head &amp; thorax; 32. Propodeum, dorsal view; 33. Propodeum, lateral view; 34. Forewing; 35. Gaster, dorsal view; 36. Pygidium." figureDoi="http://doi.org/10.5281/zenodo.5141497" httpUri="https://zenodo.org/record/5141497/files/figure.png" pageId="7" pageNumber="208">Fig. 32</figureCitation>
).Wings hyaline, apically slightly darkened, veins dark brown (
<figureCitation id="13463EB0FFF7FF991C6EF9BEFA37F9EB" box="[1339,1424,1580,1606]" captionStart="FIGURES 28–36" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[195,1395,192,1910]" captionTargetId="figure-18@9.[182,1406,181,1919]" captionTargetPageId="9" captionText="FIGURES 28–36. Lyroda binghami Tsuneki. ♀. 28. Habitus, lateral view; 29. Head, frontal view; 30. Lower half of head, frontal view; 31. Head &amp; thorax; 32. Propodeum, dorsal view; 33. Propodeum, lateral view; 34. Forewing; 35. Gaster, dorsal view; 36. Pygidium." figureDoi="http://doi.org/10.5281/zenodo.5141497" httpUri="https://zenodo.org/record/5141497/files/figure.png" pageId="7" pageNumber="208">Fig. 34</figureCitation>
). Pygidial plate broadly ferruginous and triangular, densely covered with golden setae, its lateral carina somewhat rounded (
<figureCitation id="13463EB0FFF7FF991856F9E6FEFEF922" box="[259,345,1652,1679]" captionStart="FIGURES 28–36" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[195,1395,192,1910]" captionTargetId="figure-18@9.[182,1406,181,1919]" captionTargetPageId="9" captionText="FIGURES 28–36. Lyroda binghami Tsuneki. ♀. 28. Habitus, lateral view; 29. Head, frontal view; 30. Lower half of head, frontal view; 31. Head &amp; thorax; 32. Propodeum, dorsal view; 33. Propodeum, lateral view; 34. Forewing; 35. Gaster, dorsal view; 36. Pygidium." figureDoi="http://doi.org/10.5281/zenodo.5141497" httpUri="https://zenodo.org/record/5141497/files/figure.png" pageId="7" pageNumber="208">Fig. 36</figureCitation>
). Length of F1 3.30–3.35 × its apical width. MALE. BL.
<quantity id="4C858FD0FFF7FF991ABDF9E6FB28F922" box="[1000,1167,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="7.55" metricValueMin="7.45" pageId="7" pageNumber="208" unit="mm" value="7.5" valueMax="7.55" valueMin="7.45">7.45–7.55 mm</quantity>
. Setae as in female, but denser, silvery pile bands present at apical portion of Gt1–Gt3. Anterior margin of clypeus not excavated in middle (
<figureCitation id="13463EB0FFF7FF9919F5F92EFF50F97B" box="[160,247,1724,1750]" captionStart="FIGURES 37–43" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[190,1397,191,1911]" captionTargetId="figure-17@10.[181,1405,181,1920]" captionTargetPageId="10" captionText="FIGURES 37–43. Lyroda binghami Tsuneki. ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Lower half of head, fron- tal view; 40. Head &amp; thorax; 41. Propodeum, dorsal view; 42. Pygidium; 43. Last sternum." figureDoi="http://doi.org/10.5281/zenodo.5141499" httpUri="https://zenodo.org/record/5141499/files/figure.png" pageId="7" pageNumber="208">Fig. 39</figureCitation>
). Tibio-tarsal articulation paler, tibial spurs and apical tarsomeres of all legs ferruginous. Length of F1 2–2.05 × its apical width. Propodeal dorsum coarsely irregularly rugoso-reticulate (
<figureCitation id="13463EB0FFF7FF991D1CF972FB39F957" box="[1097,1182,1760,1786]" captionStart="FIGURES 37–43" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[190,1397,191,1911]" captionTargetId="figure-17@10.[181,1405,181,1920]" captionTargetPageId="10" captionText="FIGURES 37–43. Lyroda binghami Tsuneki. ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Lower half of head, fron- tal view; 40. Head &amp; thorax; 41. Propodeum, dorsal view; 42. Pygidium; 43. Last sternum." figureDoi="http://doi.org/10.5281/zenodo.5141499" httpUri="https://zenodo.org/record/5141499/files/figure.png" pageId="7" pageNumber="208">Fig. 41</figureCitation>
). Gaster black (except pygidium brownish black). Pygidial plate broad and subtruncate, its lateral carina not rounded (
<figureCitation id="13463EB0FFF7FF991D86F896FA8EF8B3" box="[1235,1321,1796,1822]" captionStart="FIGURES 37–43" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[190,1397,191,1911]" captionTargetId="figure-17@10.[181,1405,181,1920]" captionTargetPageId="10" captionText="FIGURES 37–43. Lyroda binghami Tsuneki. ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Lower half of head, fron- tal view; 40. Head &amp; thorax; 41. Propodeum, dorsal view; 42. Pygidium; 43. Last sternum." figureDoi="http://doi.org/10.5281/zenodo.5141499" httpUri="https://zenodo.org/record/5141499/files/figure.png" pageId="7" pageNumber="208">Fig. 42</figureCitation>
). Sternum VIII without median tooth or protuberance at apex (
<figureCitation id="13463EB0FFF7FF991B89F8BAFC97F8EF" box="[732,816,1832,1858]" captionStart="FIGURES 37–43" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[190,1397,191,1911]" captionTargetId="figure-17@10.[181,1405,181,1920]" captionTargetPageId="10" captionText="FIGURES 37–43. Lyroda binghami Tsuneki. ♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Lower half of head, fron- tal view; 40. Head &amp; thorax; 41. Propodeum, dorsal view; 42. Pygidium; 43. Last sternum." figureDoi="http://doi.org/10.5281/zenodo.5141499" httpUri="https://zenodo.org/record/5141499/files/figure.png" pageId="7" pageNumber="208">Fig. 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF991992F8DEFB04F8CB" box="[199,1187,1868,1895]" pageId="7" pageNumber="208" type="distribution">
<paragraph id="8BC22235FFF7FF991992F8DEFB04F8CB" blockId="7.[151,1437,1364,2038]" box="[199,1187,1868,1895]" pageId="7" pageNumber="208">
<emphasis id="B909FE27FFF7FF991992F8DEFEC6F8CB" bold="true" box="[199,353,1868,1894]" pageId="7" pageNumber="208">Distribution.</emphasis>
<collectingCountry id="F36A62A5FFF7FF991832F8DEFE06F8CB" box="[359,417,1868,1894]" name="India" pageId="7" pageNumber="208">India</collectingCountry>
<emphasis id="B909FE27FFF7FF9918FDF8DEFD9FF8CB" bold="true" box="[424,568,1868,1894]" pageId="7" pageNumber="208">(new record</emphasis>
):
<collectingRegion id="49B9ECD7FFF7FF991B1AF8DEFD7AF8CB" box="[591,733,1868,1894]" country="India" name="Uttarakhand" pageId="7" pageNumber="208">Uttarakhand</collectingRegion>
.
<emphasis id="B909FE27FFF7FF991BBDF8DEFCFBF8CB" box="[744,860,1868,1894]" italics="true" pageId="7" pageNumber="208">Elsewhere</emphasis>
:
<collectingCountry id="F36A62A5FFF7FF991A3FF8DFFC70F8CA" box="[874,983,1869,1895]" name="Myanmar" pageId="7" pageNumber="208">Myanmar</collectingCountry>
(
<bibRefCitation id="EFEC5FC4FFF7FF991AB3F8DEFB31F8CB" author="Tsuneki, K." box="[998,1174,1868,1894]" pageId="7" pageNumber="208" pagination="69 - 81" refId="ref11594" refString="Tsuneki, K. (1983 a) Further studies on the Larrinae of the Philippine Islands, with remarks on the Indian species of the genus Lyroda, (Hymenoptera, Sphecidae). Special Publications of the Japan Hymenopterists Association, 24, 69 - 81 + 112 - 117." type="journal article" year="1983">Tsuneki, 1983a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF991992F8E2FE90F85B" pageId="7" pageNumber="208" type="discussion">
<paragraph id="8BC22235FFF7FF991992F8E2FE90F85B" blockId="7.[151,1437,1364,2038]" pageId="7" pageNumber="208">
<emphasis id="B909FE27FFF7FF991992F8E2FE93F827" bold="true" box="[199,308,1904,1930]" pageId="7" pageNumber="208">Remarks</emphasis>
. In the female, teeth on anterior clypeal margin vary among specimens. Only some specimens agree well with the original description of the clypeus, whereas others differ considerably. In the male, foretarsal coloration varies, mostly only apical tarsomeres being ferruginous,
<specimenCount id="9D7BE9BCFFF7FF991A18F82BFC41F87F" box="[845,998,1976,2003]" pageId="7" pageNumber="208" type="generic">one specimen</specimenCount>
having all fore tarsi ferruginous except the basitarsus.
</paragraph>
</subSubSection>
<caption id="DF0272BDFFF8FF9619C2F80AFEA7F855" ID-DOI="http://doi.org/10.5281/zenodo.5141493" ID-Zenodo-Dep="5141493" httpUri="https://zenodo.org/record/5141493/files/figure.png" pageId="8" pageNumber="209" startId="8.[151,264,1944,1968]" targetBox="[185,1398,193,1912]" targetPageId="8">
<paragraph id="8BC22235FFF8FF9619C2F80AFEA7F855" blockId="8.[151,1437,1944,2040]" pageId="8" pageNumber="209">
<emphasis id="B909FE27FFF8FF9619C2F80AFEE9F81D" bold="true" box="[151,334,1944,1968]" pageId="8" pageNumber="209">FIGURES 19–27</emphasis>
.
<taxonomicName id="4C7D59B6FFF8FF96180FF80AFDFEF81D" authority="Giner Mari." authorityName="Giner Mari." box="[346,601,1944,1968]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="salai">
<emphasis id="B909FE27FFF8FF96180FF80AFE7EF81D" box="[346,473,1944,1968]" italics="true" pageId="8" pageNumber="209">Lyroda salai</emphasis>
Giner Marí.
</taxonomicName>
♀. 19. Habitus, lateral view; 20. Head, frontal view; 21. Lower half of head, fron- tal view; 22. Head &amp; thorax; 23. Propodeum, dorsal view; 24. Thorax, lateral view; 25. Forewing; 26. Gaster, dorsal view; 27. Antennae.
</paragraph>
</caption>
<caption id="DF0272BDFFF9FF9719C2F805FEC1F85A" ID-DOI="http://doi.org/10.5281/zenodo.5141497" ID-Zenodo-Dep="5141497" httpUri="https://zenodo.org/record/5141497/files/figure.png" pageId="9" pageNumber="210" startId="9.[151,264,1943,1967]" targetBox="[195,1395,192,1910]" targetPageId="9">
<paragraph id="8BC22235FFF9FF9719C2F805FEC1F85A" blockId="9.[151,1436,1943,2039]" pageId="9" pageNumber="210">
<emphasis id="B909FE27FFF9FF9719C2F805FEF7F802" bold="true" box="[151,336,1943,1967]" pageId="9" pageNumber="210">FIGURES 28–36</emphasis>
.
<taxonomicName id="4C7D59B6FFF9FF97180AF805FDC8F802" authority="Tsuneki." authorityName="Tsuneki." box="[351,623,1943,1967]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="binghami">
<emphasis id="B909FE27FFF9FF97180AF805FDB7F802" box="[351,528,1943,1967]" italics="true" pageId="9" pageNumber="210">Lyroda binghami</emphasis>
Tsuneki.
</taxonomicName>
♀. 28. Habitus, lateral view; 29. Head, frontal view; 30. Lower half of head, frontal view; 31. Head &amp; thorax; 32. Propodeum, dorsal view; 33. Propodeum, lateral view; 34. Forewing; 35. Gaster, dorsal view; 36. Pygidium.
</paragraph>
</caption>
<caption id="DF0272BDFFFAFF9419C2F805FB96F87E" ID-DOI="http://doi.org/10.5281/zenodo.5141499" ID-Zenodo-Dep="5141499" httpUri="https://zenodo.org/record/5141499/files/figure.png" pageId="10" pageNumber="211" startId="10.[151,264,1943,1967]" targetBox="[190,1397,191,1911]" targetPageId="10">
<paragraph id="8BC22235FFFAFF9419C2F805FB96F87E" blockId="10.[151,1436,1943,2004]" pageId="10" pageNumber="211">
<emphasis id="B909FE27FFFAFF9419C2F805FEF3F802" bold="true" box="[151,340,1943,1967]" pageId="10" pageNumber="211">FIGURES 37–43.</emphasis>
<taxonomicName id="4C7D59B6FFFAFF94180CF80AFDC4F81D" authority="Tsuneki." authorityName="Tsuneki." box="[345,611,1944,1968]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="binghami">
<emphasis id="B909FE27FFFAFF94180CF80AFDA1F81D" box="[345,518,1944,1968]" italics="true" pageId="10" pageNumber="211">Lyroda binghami</emphasis>
Tsuneki.
</taxonomicName>
♂. 37. Habitus, lateral view; 38. Head, frontal view; 39. Lower half of head, fron- tal view; 40. Head &amp; thorax; 41. Propodeum, dorsal view; 42. Pygidium; 43. Last sternum.
</paragraph>
</caption>
<caption id="DF0272BDFFFBFF9519C2F805FEA6F855" ID-DOI="http://doi.org/10.5281/zenodo.5141501" ID-Zenodo-Dep="5141501" httpUri="https://zenodo.org/record/5141501/files/figure.png" pageId="11" pageNumber="212" startId="11.[151,264,1943,1967]" targetBox="[193,1392,193,1910]" targetPageId="11">
<paragraph id="8BC22235FFFBFF9519C2F805FEA6F855" blockId="11.[151,1437,1943,2040]" pageId="11" pageNumber="212">
<emphasis id="B909FE27FFFBFF9519C2F805FEEAF802" bold="true" box="[151,333,1943,1967]" pageId="11" pageNumber="212">FIGURES 44–52</emphasis>
.
<taxonomicName id="4C7D59B6FFFBFF95180CF80AFDE8F81D" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[345,591,1944,1968]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="formosa">
<emphasis id="B909FE27FFFBFF95180CF80AFE5EF81D" box="[345,505,1944,1968]" italics="true" pageId="11" pageNumber="212">Lyroda formosa</emphasis>
(Smith)
</taxonomicName>
. ♀. 44. Habitus, lateral view; 45. Head, frontal view; 46. Lower half of head, frontal view; 47. Head &amp; thorax; 48. Gaster, dorsal view; 49. Forewing; 50. Propodeum, dorsal view; 51. Thorax, lateral view; 52. Pygidium.
</paragraph>
</caption>
<caption id="DF0272BDFFFCFF9219C2F96CFD2CF897" ID-DOI="http://doi.org/10.5281/zenodo.5141503" ID-Zenodo-Dep="5141503" httpUri="https://zenodo.org/record/5141503/files/figure.png" pageId="12" pageNumber="213" startId="12.[151,264,1790,1814]" targetBox="[157,1425,192,1756]" targetPageId="12">
<paragraph id="8BC22235FFFCFF9219C2F96CFD2CF897" blockId="12.[151,1437,1790,1851]" pageId="12" pageNumber="213">
<emphasis id="B909FE27FFFCFF9219C2F96CFEEAF8BB" bold="true" box="[151,333,1790,1814]" pageId="12" pageNumber="213">FIGURES 53–57</emphasis>
.
<taxonomicName id="4C7D59B6FFFCFF92180CF96CFDE8F8BB" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[345,591,1790,1814]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="formosa">
<emphasis id="B909FE27FFFCFF92180CF96CFE5EF8BB" box="[345,505,1790,1814]" italics="true" pageId="12" pageNumber="213">Lyroda formosa</emphasis>
(Smith)
</taxonomicName>
. ♂. 53. Habitus, lateral view; 54. Head, frontal view; 55. Lower half of head, frontal view; 56. Antennae; 57. Propodeum, dorsal view.
</paragraph>
</caption>
<caption id="DF0272BDFFFDFF9319C2F804FE5FF85B" ID-DOI="http://doi.org/10.5281/zenodo.5141505" ID-Zenodo-Dep="5141505" httpUri="https://zenodo.org/record/5141505/files/figure.png" pageId="13" pageNumber="214" startId="13.[151,264,1942,1966]" targetBox="[181,1400,189,1912]" targetPageId="13">
<paragraph id="8BC22235FFFDFF9319C2F804FE5FF85B" blockId="13.[151,1437,1942,2039]" pageId="13" pageNumber="214">
<emphasis id="B909FE27FFFDFF9319C2F804FEE9F803" bold="true" box="[151,334,1942,1966]" pageId="13" pageNumber="214">FIGURES 58–64</emphasis>
. Anterior clypeal margins: 58.
<taxonomicName id="4C7D59B6FFFDFF931BD9F805FCB2F803" authority="Mawadda &amp; Kumar &amp; Sureshan, 2021" authorityName="Mawadda &amp; Kumar &amp; Sureshan" authorityYear="2021" box="[652,789,1943,1967]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="aurea" status="sp. nov.">
<emphasis id="B909FE27FFFDFF931BD9F805FCB2F803" box="[652,789,1943,1967]" italics="true" pageId="13" pageNumber="214">Lyroda aurea</emphasis>
</taxonomicName>
<emphasis id="B909FE27FFFDFF931A49F805FCCAF802" bold="true" box="[796,877,1943,1967]" pageId="13" pageNumber="214">
<taxonomicNameLabel id="A23A435CFFFDFF931A49F805FCCAF802" box="[796,877,1943,1967]" pageId="13" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype ♀; 59.
<taxonomicName id="4C7D59B6FFFDFF931D49F805FB3AF802" authority="Mawadda &amp; Kumar &amp; Sureshan, 2021" authorityName="Mawadda &amp; Kumar &amp; Sureshan" authorityYear="2021" box="[1052,1181,1943,1967]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="nuda" status="sp. nov.">
<emphasis id="B909FE27FFFDFF931D49F805FB3AF802" box="[1052,1181,1943,1967]" italics="true" pageId="13" pageNumber="214">Lyroda nuda</emphasis>
</taxonomicName>
<emphasis id="B909FE27FFFDFF931DF1F805FB52F802" bold="true" box="[1188,1269,1943,1967]" pageId="13" pageNumber="214">
<taxonomicNameLabel id="A23A435CFFFDFF931DF1F805FB52F802" box="[1188,1269,1943,1967]" pageId="13" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype ♀; 60.
<taxonomicName id="4C7D59B6FFFDFF9319C2F829FE3FF87E" authority="Giner Mari." authorityName="Giner Mari." box="[151,408,1979,2003]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="salai">
<emphasis id="B909FE27FFFDFF9319C2F829FEB0F87E" box="[151,279,1979,2003]" italics="true" pageId="13" pageNumber="214">Lyroda salai</emphasis>
Giner Marí.
</taxonomicName>
♀; 61.
<taxonomicName id="4C7D59B6FFFDFF9318B2F829FD53F87E" authority="Tsuneki." authorityName="Tsuneki." box="[487,756,1979,2003]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="binghami">
<emphasis id="B909FE27FFFDFF9318B2F829FD31F87E" box="[487,662,1979,2003]" italics="true" pageId="13" pageNumber="214">Lyroda binghami</emphasis>
Tsuneki.
</taxonomicName>
♀; 62.
<taxonomicName id="4C7D59B6FFFDFF931A16F829FBF7F87E" authority="Tsuneki." authorityName="Tsuneki." box="[835,1104,1979,2003]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="binghami">
<emphasis id="B909FE27FFFDFF931A16F829FC55F87E" box="[835,1010,1979,2003]" italics="true" pageId="13" pageNumber="214">Lyroda binghami</emphasis>
Tsuneki.
</taxonomicName>
♂; 63.
<taxonomicName id="4C7D59B6FFFDFF931DCAF829FA3EF87E" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[1183,1433,1979,2003]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="formosa">
<emphasis id="B909FE27FFFDFF931DCAF829FAE6F87E" box="[1183,1345,1979,2003]" italics="true" pageId="13" pageNumber="214">Lyroda formosa</emphasis>
(Smith)
</taxonomicName>
. ♀; 64.
<taxonomicName id="4C7D59B6FFFDFF93198BF84DFE72F85A" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[222,469,2015,2039]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="formosa">
<emphasis id="B909FE27FFFDFF93198BF84DFED9F85A" box="[222,382,2015,2039]" italics="true" pageId="13" pageNumber="214">Lyroda formosa</emphasis>
(Smith)
</taxonomicName>
. ♂.
</paragraph>
</caption>
<subSubSection id="C36771BEFFFEFF901992FF04FD32FEB0" pageId="14" pageNumber="215" type="materials_examined">
<paragraph id="8BC22235FFFEFF901992FF04FD32FEB0" blockId="14.[151,1437,150,285]" pageId="14" pageNumber="215">
<emphasis id="B909FE27FFFEFF901992FF04FE0AFF1D" bold="true" box="[199,429,150,176]" pageId="14" pageNumber="215">Material examined.</emphasis>
<materialsCitation id="3B152868FFFEFF9018E4FF05FA9EFF78" collectingDate="2019-07-15" collectorName="Coll. P. Girish Kumar &amp; Regd. Nos." country="India" latitude="30.345001" location="Dehradun district" longLatPrecision="20" longitude="78.01028" pageId="14" pageNumber="215" specimenCount="13" specimenCount-female="8" specimenCount-male="5" stateProvince="Uttarakhand">
<collectingCountry id="F36A62A5FFFEFF9018E4FF05FDA2FF1C" box="[433,517,151,177]" name="India" pageId="14" pageNumber="215">INDIA</collectingCountry>
:
<collectingRegion id="49B9ECD7FFFEFF901B5EFF05FD3FFF1C" box="[523,664,151,177]" country="India" name="Uttarakhand" pageId="14" pageNumber="215">Uttarakhand</collectingRegion>
,
<location id="8EA274EEFFFEFF901BF5FF05FCC4FF1C" LSID="urn:lsid:plazi:treatment:03D49323FFF7FF9019C2FB37FD32FEB0:8EA274EEFFFEFF901BF5FF05FCC4FF1C" box="[672,867,151,177]" country="India" latitude="30.345001" longLatPrecision="20" longitude="78.01028" name="Dehradun district" pageId="14" pageNumber="215" stateProvince="Uttarakhand">Dehradun district</location>
,ZSI campus(
<specimenCount id="9D7BE9BCFFFEFF901AAFFF04FB84FF1C" box="[1018,1059,150,177]" pageId="14" pageNumber="215" type="male">5 ♂</specimenCount>
,
<specimenCount id="9D7BE9BCFFFEFF901D79FF05FBF7FF1C" box="[1068,1104,151,177]" pageId="14" pageNumber="215" type="female">8♀</specimenCount>
) (
<geoCoordinate id="EE4944F2FFFEFF901D32FF05FB5EFF1C" box="[1127,1273,151,177]" degrees="30" direction="north" minutes="20" orientation="latitude" pageId="14" pageNumber="215" precision="15" seconds="42" value="30.345001">30°20’42’’N</geoCoordinate>
,
<geoCoordinate id="EE4944F2FFFEFF901C57FF05FA28FF1C" box="[1282,1423,151,177]" degrees="78" direction="east" minutes="00" orientation="longitude" pageId="14" pageNumber="215" precision="15" seconds="37" value="78.01028">78°00’37’’E</geoCoordinate>
),
<date id="FFC304F5FFFEFF9019C2FF28FEBEFF78" box="[151,281,186,213]" pageId="14" pageNumber="215" value="2019-07-15">
<collectingDate id="EF87FD1DFFFEFF9019C2FF28FEBEFF78" box="[151,281,186,213]" pageId="14" pageNumber="215" value="2019-07-15">15.vii.2019</collectingDate>
</date>
,
<collectorName id="268847E3FFFEFF901872FF29FD85FF78" box="[295,546,187,213]" pageId="14" pageNumber="215">Coll. P. Girish Kumar</collectorName>
, ZSIK
<collectorName id="268847E3FFFEFF901B23FF29FD51FF78" box="[630,758,187,213]" pageId="14" pageNumber="215">Regd. Nos.</collectorName>
ZSI/ WGRC/IR/INV/15035–15042, 15045–15048
</materialsCitation>
.
<materialsCitation id="3B152868FFFEFF901C10FF29FD36FEB0" collectingDate="2017-10-08" collectorName="Coll. A. Ramesh Kumar &amp; Regd. No." country="India" latitude="30.285276" location="Dehradun district" longLatPrecision="20" longitude="77.97528" pageId="14" pageNumber="215" specimenCount="1" specimenCount-female="1" stateProvince="Uttarakhand">
<collectingCountry id="F36A62A5FFFEFF901C10FF29FA3EFF78" box="[1349,1433,187,213]" name="India" pageId="14" pageNumber="215">INDIA</collectingCountry>
:
<collectingRegion id="49B9ECD7FFFEFF9019C2FF4DFE83FF54" box="[151,292,223,249]" country="India" name="Uttarakhand" pageId="14" pageNumber="215">Uttarakhand</collectingRegion>
,
<location id="8EA274EEFFFEFF90187AFF4DFE52FF54" LSID="urn:lsid:plazi:treatment:03D49323FFF7FF9019C2FB37FD32FEB0:8EA274EEFFFEFF90187AFF4DFE52FF54" box="[303,501,223,249]" country="India" latitude="30.285276" longLatPrecision="20" longitude="77.97528" name="Dehradun district" pageId="14" pageNumber="215" stateProvince="Uttarakhand">Dehradun district</location>
, WII campus (
<specimenCount id="9D7BE9BCFFFEFF901BCFFF4DFD18FF54" box="[666,703,223,249]" pageId="14" pageNumber="215" type="female">1 ♀</specimenCount>
) (
<geoCoordinate id="EE4944F2FFFEFF901B8FFF4DFCCFFF54" box="[730,872,223,249]" degrees="30" direction="north" minutes="17" orientation="latitude" pageId="14" pageNumber="215" precision="15" seconds="07" value="30.285276">30°17’07’’N</geoCoordinate>
<geoCoordinate id="EE4944F2FFFEFF901A20FF4CFBA1FF54" box="[885,1030,222,249]" degrees="77" direction="east" minutes="58" orientation="longitude" pageId="14" pageNumber="215" precision="15" seconds="31" value="77.97528">77°58’31’’ E</geoCoordinate>
),
<date id="FFC304F5FFFEFF901D4FFF4DFBD9FF54" box="[1050,1150,223,249]" pageId="14" pageNumber="215" value="2017-10-08">
<collectingDate id="EF87FD1DFFFEFF901D4FFF4DFBD9FF54" box="[1050,1150,223,249]" pageId="14" pageNumber="215" value="2017-10-08">8.x.2017</collectingDate>
</date>
,
<collectorName id="268847E3FFFEFF901DDFFF4DFA3EFF54" box="[1162,1433,223,249]" pageId="14" pageNumber="215">Coll. A. Ramesh Kumar</collectorName>
, ZSIK
<collectorName id="268847E3FFFEFF901989FE91FEE8FEB0" box="[220,335,259,285]" pageId="14" pageNumber="215">Regd. No.</collectorName>
ZSI/ WGRC/IR/INV/15043
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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