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<document id="86044ABFD5C6745D662F86C8942B0410" ID-DOI="10.5281/zenodo.193258" ID-GBIF-Dataset="85176744-66ec-48ba-860c-671c02718c50" ID-ISSN="1175-5326" ID-Zenodo-Dep="193258" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460129092828" checkinUser="plazi" docAuthor="Johanson, Kjell Arne &amp; Oláh, János" docDate="2010" docId="03CCF00CFF881E64FF30F280EFA656D8" docLanguage="en" docName="zt02338p062.pdf" docOrigin="Zootaxa 2338" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Paduniella madagassa Johanson &amp; Oláh, 2010, new species" docType="treatment" docVersion="9" lastPageNumber="60" masterDocId="FFF58874FF8D1E62FFA7F31DEC1A551A" masterDocTitle="Descriptions of five new species of Paduniella from Madagascar (Trichoptera: Psychomyiidae)" masterLastPageNumber="62" masterPageNumber="54" pageNumber="59" updateTime="1698235096248" updateUser="plazi">
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<mods:title id="65FBA5309F3D927B84C5B70D8A24A63A">Descriptions of five new species of Paduniella from Madagascar (Trichoptera: Psychomyiidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="5B555F73DE6E4CA46E115E2B9AD3E7D3">Johanson, Kjell Arne</mods:namePart>
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<mods:namePart id="9DDB415DE1C962A33223BAFF51314E5E">Oláh, János</mods:namePart>
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<mods:typeOfResource id="1B5009776A193065643A3AA6A341D259">text</mods:typeOfResource>
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<mods:date id="F5054F5A3A1BCBD3012EDC1344E49438">2010</mods:date>
<mods:detail id="FD52F089C0F4997FA20BD25EEC542546" type="volume">
<mods:number id="42C56A7039B39141053BF5C1B6DE7F78">2338</mods:number>
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<mods:classification id="6044CD104140DC40051D79A192767F07">journal article</mods:classification>
<mods:identifier id="3057CEF46D68A11465AA9151B390F9A1" type="DOI">10.5281/zenodo.193258</mods:identifier>
<mods:identifier id="527CF5F1945573D0919ED3DA9AB087D3" type="GBIF-Dataset">85176744-66ec-48ba-860c-671c02718c50</mods:identifier>
<mods:identifier id="BF4760D675BDE707D147E6EA287C10DC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="34D809E56FC70616BC7010FDE2604C08" type="Zenodo-Dep">193258</mods:identifier>
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<treatment id="03CCF00CFF881E64FF30F280EFA656D8" ID-DOI="http://doi.org/10.5281/zenodo.6205189" ID-GBIF-Taxon="124955623" ID-Zenodo-Dep="6205189" LSID="urn:lsid:plazi:treatment:03CCF00CFF881E64FF30F280EFA656D8" httpUri="http://treatment.plazi.org/id/03CCF00CFF881E64FF30F280EFA656D8" lastPageId="6" lastPageNumber="60" pageId="5" pageNumber="59">
<subSubSection id="C37F1291FF881E67FF30F280ED0354C0" pageId="5" pageNumber="59" type="nomenclature">
<paragraph id="8BDA411AFF881E67FF30F280EE5C54AD" blockId="5.[151,582,413,474]" box="[151,582,413,439]" pageId="5" pageNumber="59">
<heading id="D092F676FF881E67FF30F280EE5C54AD" bold="true" box="[151,582,413,439]" fontSize="11" level="1" pageId="5" pageNumber="59" reason="1">
<emphasis id="B9119D08FF881E67FF30F280EE5C54AD" bold="true" box="[151,582,413,439]" pageId="5" pageNumber="59">
<taxonomicName id="4C653A99FF881E67FF30F280EDBD54AD" box="[151,423,413,439]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="59" phylum="Arthropoda" rank="species" species="madagassa" status="sp. nov.">
<emphasis id="B9119D08FF881E67FF30F280EDBD54AD" bold="true" box="[151,423,413,439]" italics="true" pageId="5" pageNumber="59">Paduniella madagassa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2222073FF881E67FE11F280EE5C54AD" box="[438,582,413,439]" pageId="5" pageNumber="59" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA411AFF881E67FF30F2DDED0354C0" blockId="5.[151,582,413,474]" box="[151,281,448,474]" pageId="5" pageNumber="59">
<figureCitation id="135E5D9FFF881E67FF30F2DDED0354C0" box="[151,281,448,474]" captionStart="FIGURES 10 12" captionStartId="5.[151,273,1729,1753]" captionTargetBox="[154,1433,752,1669]" captionTargetId="figure@5.[151,1436,714,1708]" captionTargetPageId="5" captionText="FIGURES 10 12. Paduniella madagassa, new species, holotype, male genitalia: 10, lateral; 11, phallic apparatus, lateral; 12, ventral." httpUri="https://zenodo.org/record/193262/files/figure.png" pageId="5" pageNumber="59">Figs 1012</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37F1291FF881E64FF30F117E97A561D" lastPageId="6" lastPageNumber="60" pageId="5" pageNumber="59" type="description">
<paragraph id="8BDA411AFF881E67FF30F117EEA3578E" blockId="5.[151,1436,522,660]" pageId="5" pageNumber="59">
The long median branch of the inferior appendages is bilobed in
<taxonomicName id="4C653A99FF881E67FC68F117E8435739" box="[975,1113,522,547]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="59" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B9119D08FF881E67FC68F117E8435739" box="[975,1113,522,547]" italics="true" pageId="5" pageNumber="59">P. africana</emphasis>
</taxonomicName>
,
<emphasis id="B9119D08FF881E67FBCCF117E8C25739" box="[1131,1240,522,547]" italics="true" pageId="5" pageNumber="59">P. a m b r a</emphasis>
and
<taxonomicName id="4C653A99FF881E67FABDF117E98E5739" box="[1306,1428,522,547]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="59" phylum="Arthropoda" rank="species" species="nandra">
<emphasis id="B9119D08FF881E67FABDF117E98E5739" box="[1306,1428,522,547]" italics="true" pageId="5" pageNumber="59">P. nandra</emphasis>
</taxonomicName>
.
<taxonomicName id="4C653A99FF881E67FF30F12DEDB25753" box="[151,424,560,585]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="59" phylum="Arthropoda" rank="species" species="madagassa">
<emphasis id="B9119D08FF881E67FF30F12DEDB25753" box="[151,424,560,585]" italics="true" pageId="5" pageNumber="59">Paduniella madagassa</emphasis>
</taxonomicName>
differs from these species by having tergite IX and superior appendages narrow and apparently fused, inferior appendages narrow; sternal strips broad and originating from the central part of a broad phallobase, and sternite IX semicircular.
</paragraph>
<caption id="DF1A1192FF881E67FF30F5DCED7B53E2" httpUri="https://zenodo.org/record/193262/files/figure.png" pageId="5" pageNumber="59" targetBox="[154,1433,752,1669]" targetPageId="5">
<paragraph id="8BDA411AFF881E67FF30F5DCED7B53E2" blockId="5.[151,1436,1729,1784]" pageId="5" pageNumber="59">
<emphasis id="B9119D08FF881E67FF30F5DCED7F53C3" bold="true" box="[151,357,1729,1753]" pageId="5" pageNumber="59">FIGURES 1012.</emphasis>
<taxonomicName id="4C653A99FF881E67FECBF5DFEE7053C3" box="[364,618,1730,1753]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="59" phylum="Arthropoda" rank="species" species="madagassa" status="sp. nov.">
<emphasis id="B9119D08FF881E67FECBF5DFEE7053C3" box="[364,618,1730,1753]" italics="true" pageId="5" pageNumber="59">Paduniella madagassa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2222073FF881E67FDDDF5DCEF1853C2" box="[634,770,1729,1752]" pageId="5" pageNumber="59" rank="species">new species</taxonomicNameLabel>
, holotype, male genitalia: 10, lateral; 11, phallic apparatus, lateral; 12, ventral.
</paragraph>
</caption>
<paragraph id="8BDA411AFF881E64FF61F43AE97A561D" blockId="5.[151,1436,1831,2006]" lastBlockId="6.[151,1437,152,962]" lastPageId="6" lastPageNumber="60" pageId="5" pageNumber="59">
<emphasis id="B9119D08FF881E67FF61F43AEE39525A" box="[198,547,1831,1856]" italics="true" pageId="5" pageNumber="59">
Male genitalia (
<figureCitation id="135E5D9FFF881E67FE2EF43AEE0C525A" box="[393,534,1831,1856]" captionStart="FIGURES 10 12" captionStartId="5.[151,273,1729,1753]" captionTargetBox="[154,1433,752,1669]" captionTargetId="figure@5.[151,1436,714,1708]" captionTargetPageId="5" captionText="FIGURES 10 12. Paduniella madagassa, new species, holotype, male genitalia: 10, lateral; 11, phallic apparatus, lateral; 12, ventral." httpUri="https://zenodo.org/record/193262/files/figure.png" pageId="5" pageNumber="59">Figs 1012</figureCitation>
).
</emphasis>
Sternite IX narrowly club-shaped in lateral view; both ventral arms with almost straight posterior and anterior margin, nearly 3 times longer than wide, about
<quantity id="4C9DECFFFF881E67FB10F451E90F527C" box="[1207,1301,1868,1894]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="59" unit="mm" value="0.1">0.1 mm</quantity>
long; with swelling at posteroventral corner; both dorsal arms narrowing anterad, nearly needle-shaped at apex, curving anterad along entire length, joined with tergite IX at dorsal part of genitalia, about
<quantity id="4C9DECFFFF881E67FBC5F48AE8D152AB" box="[1122,1227,1943,1969]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="59" unit="mm" value="0.15">0.15 mm</quantity>
long. Ventral and dorsal arms forming sickle-shaped plates. Tergite IX long, smooth, in lateral view nearly parallel-sided, straight; apex completely fused with superior appendages. Superior appendages completely fused with apex of tergite IX; tapering apically, with concave ventral margin and convex dorsal margin, about as wide as widest part of sternite IX; setae present at apices; apex pointing ventrad. Median branch of inferior appendages curving ventrad along entire length; in ventral view originating proximally on main branch; parallel-sided, slightly curving mesad; apex bilobed; lobes equally large; covered by setae; nearly
<quantity id="4C9DECFFFF8B1E64FA95F230E986545D" box="[1330,1436,301,327]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="60" unit="mm" value="0.14">0.14 mm</quantity>
long. Ventral lobe of inferior appendages minute, triangular in lateral and ventral view. Main branch of inferior appendages about
<quantity id="4C9DECFFFF8B1E64FE75F265EE215488" box="[466,571,376,402]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="60" unit="mm" value="0.35">0.35 mm</quantity>
long, narrowing distad to mid-length; slightly curved ventrad; distal half narrow, straight; proximal part 2 times wider than widest part of superior appendages in lateral view. Phallic apparatus long, connected to sternite IX and tergite IX by long, horizontal, posteriorly narrowing sternal strips; dorsal margin S-shaped. Phallobase about
<quantity id="4C9DECFFFF8B1E64FD41F2F5EF5B5718" box="[742,833,488,514]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="60" unit="mm" value="0.5">0.5 mm</quantity>
long, longer than phallicata, strongly pointed into long, anterior apex; posterior part with convex dorsal and ventral margins; about 2 times wider than sternal strips. Median processes about
<quantity id="4C9DECFFFF8B1E64FDA2F12FEE445756" box="[517,606,562,588]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="60" unit="mm" value="0.6">0.6 mm</quantity>
long, needle-shaped, straight along entire length; originating from mid part of phallobase; each apex reaching as far posteriorly as paramere; longer than phallicata; series of ventral teeth present on left branch, absent on right branch. Phallicata strongly club-shaped; basal two-thirds slender, straight, parallel-sided; apex broad, asymmetrical, ventral margin nearly straight, dorsal margin slightly convex. Paramere separated from phallicata at posterior apex of phallobase; needle-shaped along its length, except slightly drop-shaped at apex; running parallel with and exceeding phallicata; nearly
<quantity id="4C9DECFFFF8B1E64FB64F1F0E907561D" box="[1219,1309,749,775]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="60" unit="mm" value="0.5">0.5 mm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="C37F1291FF8B1E64FF63F00FEEA9566D" pageId="6" pageNumber="60" type="materials_examined">
<paragraph id="8BDA411AFF8B1E64FF63F00FEE4C5648" blockId="6.[151,1437,152,962]" pageId="6" pageNumber="60">
<emphasis id="B9119D08FF8B1E64FF63F00FED235636" bold="true" box="[196,313,786,812]" pageId="6" pageNumber="60">
<typeStatus id="54DEFFB8FF8B1E64FF63F00FED235636" box="[196,313,786,812]" pageId="6" pageNumber="60" type="holotype">Holotype</typeStatus>
</emphasis>
male, Prov. Fianarantsoa,
<quantity id="4C9DECFFFF8B1E64FDDAF00FEEA05636" box="[637,698,786,812]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="60" unit="km" value="7.0">7 km</quantity>
W Ranomafana,
<quantity id="4C9DECFFFF8B1E64FC28F00FEFF75636" box="[911,1005,786,812]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="60" unit="m" value="1100.0">1100 m</quantity>
,
<date id="FFDB67DAFF8B1E64FC5DF00FE8645636" box="[1018,1150,786,812]" pageId="6" pageNumber="60" value="1988-09-14">14.ix.1988</date>
, black light in montane forest [C. Kremen] (
<collectionCode id="ED74D9DFFF8B1E64FE23F025EDF85648" box="[388,482,824,850]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="60">NMNH</collectionCode>
, alcohol).
</paragraph>
<paragraph id="8BDA411AFF8B1E64FF63F040EEA9566D" blockId="6.[151,1437,152,962]" box="[196,691,861,887]" pageId="6" pageNumber="60">
<emphasis id="B9119D08FF8B1E64FF63F040ED25566D" bold="true" box="[196,319,861,887]" pageId="6" pageNumber="60">
<typeStatus id="54DEFFB8FF8B1E64FF63F040ED21566D" box="[196,315,861,887]" pageId="6" pageNumber="60" type="paratype">Paratype</typeStatus>
:
</emphasis>
Same data as
<typeStatus id="54DEFFB8FF8B1E64FE40F040EE4A566D" box="[487,592,861,887]" pageId="6" pageNumber="60" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D638A93FF8B1E64FDFBF040EEB5566D" box="[604,687,861,887]" pageId="6" pageNumber="60" type="male">1 male</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F1291FF8B1E64FF63F09FEF3B5686" box="[196,801,898,924]" pageId="6" pageNumber="60" type="distribution">
<paragraph id="8BDA411AFF8B1E64FF63F09FEF3B5686" blockId="6.[151,1437,152,962]" box="[196,801,898,924]" pageId="6" pageNumber="60">
<emphasis id="B9119D08FF8B1E64FF63F09FED7E5686" bold="true" box="[196,356,898,924]" pageId="6" pageNumber="60">Distribution.</emphasis>
<collectingCountry id="F372018AFF8B1E64FECBF09FEDE05686" box="[364,506,898,924]" name="Madagascar" pageId="6" pageNumber="60">Madagascar</collectingCountry>
(Province Fianarantsoa).
</paragraph>
</subSubSection>
<subSubSection id="C37F1291FF8B1E64FF63F0BAEFA656D8" box="[196,956,935,962]" pageId="6" pageNumber="60" type="etymology">
<paragraph id="8BDA411AFF8B1E64FF63F0BAEFA656D8" blockId="6.[151,1437,152,962]" box="[196,956,935,962]" pageId="6" pageNumber="60">
<emphasis id="B9119D08FF8B1E64FF63F0BAED4B56DB" bold="true" box="[196,337,935,961]" pageId="6" pageNumber="60">Etymology.</emphasis>
The name,
<taxonomicName id="4C653A99FF8B1E64FE7CF0B5EE4556DB" box="[475,607,936,961]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="60" phylum="Arthropoda" rank="species" species="madagassa">
<emphasis id="B9119D08FF8B1E64FE7CF0B5EE4556DB" box="[475,607,936,961]" italics="true" pageId="6" pageNumber="60">madagassa</emphasis>
</taxonomicName>
, is derived from
<collectingCountry id="F372018AFF8B1E64FC8FF0B5EFA256D8" box="[808,952,936,962]" name="Madagascar" pageId="6" pageNumber="60">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>