143 lines
20 KiB
XML
143 lines
20 KiB
XML
<document id="4210C79C44E23FE307F1BEF40FE34493" ID-CLB-Dataset="302295" ID-DOI="10.11646/zootaxa.5501.3.1" ID-GBIF-Dataset="3de6936e-301e-4a57-ada2-a261af89bb7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="13628165" ID-ZooBank="6A91B5D0-0251-4F5C-AA32-D70CAAE55F90" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1725278913235" checkinUser="plazi" docAuthor="Telnov, Dmitry" docDate="2024" docId="03C887C4361A34120D84D79CB83DBC2D" docLanguage="en" docName="zootaxa.5501.3.1.pdf" docOrigin="Zootaxa 5501 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5501.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copobaeninae Abdullah 1969" docType="treatment" docVersion="7" lastPageNumber="404" masterDocId="FFF1FFBC361934110D13D70BBC3AB943" masterDocTitle="On the taxonomy of some South American and Australo-Pacific Anthicidae Latreille, 1819 (Coleoptera) with new subfamily and tribe-rank synonymies" masterLastPageNumber="424" masterPageNumber="401" pageNumber="404" updateTime="1725641563685" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B3004D52F42C13F6D3B9BF3E83A926D9">On the taxonomy of some South American and Australo-Pacific Anthicidae Latreille, 1819 (Coleoptera) with new subfamily and tribe-rank synonymies</mods:title>
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<mods:namePart id="A303A7CA3F1026AC4F42657FDA492069">Telnov, Dmitry</mods:namePart>
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<mods:affiliation id="9FA7BA4B2F6A5E1F8B24A3A60CBFD4AD">Department of Life Sciences, Natural History Museum, SW 7 5 BD London, United Kingdom & Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV- 5401 Daugavpils, Latvia & Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004 Rīga, Latvia</mods:affiliation>
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<treatment id="03C887C4361A34120D84D79CB83DBC2D" ID-DOI="http://doi.org/10.5281/zenodo.13712904" ID-GBIF-Taxon="240378952" ID-Zenodo-Dep="13712904" LSID="urn:lsid:plazi:treatment:03C887C4361A34120D84D79CB83DBC2D" httpUri="http://treatment.plazi.org/id/03C887C4361A34120D84D79CB83DBC2D" lastPageNumber="404" pageId="3" pageNumber="404">
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<paragraph id="8BDE36D2361A34120D84D79CBE35B9F2" blockId="3.[151,527,151,213]" box="[151,527,151,177]" pageId="3" pageNumber="404">
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<heading id="D09681BE361A34120D84D79CBE35B9F2" bold="true" box="[151,527,151,177]" fontSize="11" level="1" pageId="3" pageNumber="404" reason="1">
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<taxonomicName id="4C614D51361A34120D84D79CBE35B9F2" authority="Abdullah, 1969" authorityName="Abdullah" authorityYear="1969" box="[151,527,151,177]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subFamily" subFamily="Copobaeninae">
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<emphasis id="B915EAC0361A34120D84D79CBE35B9F2" bold="true" box="[151,527,151,177]" pageId="3" pageNumber="404">
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Copobaeninae
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<bibRefCitation id="EFF04B23361A34120C43D79CBE35B9F2" author="Abdullah, M." box="[336,527,151,177]" pageId="3" pageNumber="404" pagination="323 - 366" refId="ref11957" refString="Abdullah, M. (1969) The natural classification of the family Anthicidae with some ecological and ethological observations (Coleoptera). Deutsche Entomologische Zeitschrift, 16 (iv / v), 323 - 366. https: // doi. org / 10.1002 / mmnd. 4810160404" type="journal article" year="1969">Abdullah, 1969</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BDE36D2361A34120D84D7B0BDABB996" blockId="3.[151,527,151,213]" box="[151,401,187,213]" pageId="3" pageNumber="404">
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<bibRefCitation id="EFF04B23361A34120D84D7B0BDB7B996" author="Abdullah, M." box="[151,397,187,213]" pageId="3" pageNumber="404" pagination="323 - 366" refId="ref11957" refString="Abdullah, M. (1969) The natural classification of the family Anthicidae with some ecological and ethological observations (Coleoptera). Deutsche Entomologische Zeitschrift, 16 (iv / v), 323 - 366. https: // doi. org / 10.1002 / mmnd. 4810160404" type="journal article" year="1969">Abdullah (1969: 334)</bibRefCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37B6559361A34120D84D60BBD49B875" pageId="3" pageNumber="404" type="reference_group">
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||
<paragraph id="8BDE36D2361A34120D84D60BBE11B85B" blockId="3.[151,555,256,310]" box="[151,555,256,280]" pageId="3" pageNumber="404">
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=
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<taxonomicName id="4C614D51361A34120DBFD60BBDF2B85B" authority="Matthews, 1987" authorityName="Matthews" authorityYear="1987" box="[172,456,256,280]" pageId="3" pageNumber="402" rank="subFamily" status="syn. nov." subFamily="Lemodinae">
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Lemodinae
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<bibRefCitation id="EFF04B23361A34120C37D60BBDF2B85B" author="Matthews, E. G." box="[292,456,256,280]" pageId="3" pageNumber="404" pagination="1 - 67" refId="ref13287" refString="Matthews, E. G. (1987) A guide to the genera of beetles of South Australia. Part 5. Polyphaga: Tenebrionoidea. South Australian Museum, Special Educational Bulletin, 8, i - v + 1 - 67. https: // doi. org / 10.1163 / 9789004627727" type="journal article" year="1987">Matthews, 1987</bibRefCitation>
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</taxonomicName>
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||
<emphasis id="B915EAC0361A34120CDDD60BBE11B85B" bold="true" box="[462,555,256,280]" pageId="3" pageNumber="404">
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<taxonomicNameLabel id="A22657BB361A34120CDDD60BBE11B85B" box="[462,555,256,280]" pageId="3" pageNumber="404">syn. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8BDE36D2361A34120D84D615BD49B875" blockId="3.[151,555,256,310]" box="[151,371,286,310]" pageId="3" pageNumber="404">
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<bibRefCitation id="EFF04B23361A34120D84D615BD55B875" author="Matthews, E. G." box="[151,367,286,310]" pageId="3" pageNumber="404" pagination="1 - 67" refId="ref13287" refString="Matthews, E. G. (1987) A guide to the genera of beetles of South Australia. Part 5. Polyphaga: Tenebrionoidea. South Australian Museum, Special Educational Bulletin, 8, i - v + 1 - 67. https: // doi. org / 10.1163 / 9789004627727" type="journal article" year="1987">Matthews (1987: 40)</bibRefCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37B6559361A34120D84D66FB83DBC2D" pageId="3" pageNumber="404" type="description">
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<paragraph id="8BDE36D2361A34120D84D66FBF77BBDD" blockId="3.[151,1437,356,1391]" pageId="3" pageNumber="404">
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<bibRefCitation id="EFF04B23361A34120D84D66FBD6CB83D" author="Abdullah, M." box="[151,342,356,382]" pageId="3" pageNumber="404" pagination="323 - 366" refId="ref11957" refString="Abdullah, M. (1969) The natural classification of the family Anthicidae with some ecological and ethological observations (Coleoptera). Deutsche Entomologische Zeitschrift, 16 (iv / v), 323 - 366. https: // doi. org / 10.1002 / mmnd. 4810160404" type="journal article" year="1969">Abdullah (1969)</bibRefCitation>
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erected a new anthicid subfamily
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<taxonomicName id="4C614D51361A34120FE6D66FBFAFB83D" authority="Abdullah, 1969" authorityName="Abdullah" authorityYear="1969" box="[757,917,356,382]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subFamily" subFamily="Copobaeninae">Copobaeninae</taxonomicName>
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for the South American genus
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<taxonomicName id="4C614D51361A34120803D66FBDF0B8E1" authority="Fairmaire & Germain, 1863" authorityName="Fairmaire & Germain" authorityYear="1863" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAC0361A34120803D66FB9A6B83D" box="[1296,1436,356,382]" italics="true" pageId="3" pageNumber="404">Copobaenus</emphasis>
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<bibRefCitation id="EFF04B23361A34120D84D683BDF0B8E1" author="Fairmaire, L. & Germain, P." box="[151,458,392,419]" pageId="3" pageNumber="404" pagination="225 - 284" refId="ref12651" refString="Fairmaire, L. & Germain, P. (1863) Revision des Coleopteres du Chili. Suite. Annales de la Societe Entomologique de France, 3 (4), 225 - 284. https: // doi. org / 10.1080 / 00379271.1863.11755447" type="journal article" year="1863">Fairmaire & Germain, 1863</bibRefCitation>
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</taxonomicName>
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based on (sequence as in
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<bibRefCitation id="EFF04B23361A34120FCCD683BFA3B8E1" author="Abdullah, M." box="[735,921,392,418]" pageId="3" pageNumber="404" pagination="323 - 366" refId="ref11957" refString="Abdullah, M. (1969) The natural classification of the family Anthicidae with some ecological and ethological observations (Coleoptera). Deutsche Entomologische Zeitschrift, 16 (iv / v), 323 - 366. https: // doi. org / 10.1002 / mmnd. 4810160404" type="journal article" year="1969">Abdullah (1969)</bibRefCitation>
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, supplemented by me) entire (non-emarginate) compound eyes, wide cranial neck, procoxal cavity closed internally open externally behind, narrowly separated metacoxa (‘subcontiguous’ as of
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<bibRefCitation id="EFF04B23361A34120F05D6DBBEE7B8A9" author="Abdullah, M." box="[534,733,464,490]" pageId="3" pageNumber="404" pagination="323 - 366" refId="ref11957" refString="Abdullah, M. (1969) The natural classification of the family Anthicidae with some ecological and ethological observations (Coleoptera). Deutsche Entomologische Zeitschrift, 16 (iv / v), 323 - 366. https: // doi. org / 10.1002 / mmnd. 4810160404" type="journal article" year="1969">Abdullah (1969))</bibRefCitation>
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with internal carina reduced ‘to a narrow-based apophysis’, simple, basally subdentate pretarsal claws, metendosternite with long stalk which is as long as arms, well-developed laminae and anterior tendons present on arms above their junction point with laminae, metathoracic wing with ‘radial cell closed, anal cell absent’, aedeagus with parameres fused, ovipositor with ‘stylus borne on second segment of coxite’. Additionally, in his key to
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<taxonomicName id="4C614D51361A34120F30D56BBEA4BB39" authorityName="Latreille" authorityYear="1819" box="[547,670,608,634]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="family">Anthicidae</taxonomicName>
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subfamilies in a couplet with
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<taxonomicName id="4C614D51361A34120EEBD56BB8A1BB39" box="[1016,1179,608,634]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subFamily" subFamily="Copobaeninae">Copobaeninae</taxonomicName>
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,
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<bibRefCitation id="EFF04B23361A341209B4D56BB9A6BB38" author="Abdullah, M." box="[1191,1436,608,635]" pageId="3" pageNumber="404" pagination="323 - 366" refId="ref11957" refString="Abdullah, M. (1969) The natural classification of the family Anthicidae with some ecological and ethological observations (Coleoptera). Deutsche Entomologische Zeitschrift, 16 (iv / v), 323 - 366. https: // doi. org / 10.1002 / mmnd. 4810160404" type="journal article" year="1969">Abdullah (1969: 330)</bibRefCitation>
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specifically mentioned ‘ovipositor with 2-segmented coxites’.
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</paragraph>
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<paragraph id="8BDE36D2361A34120DD4D5A3BD0ABA11" blockId="3.[151,1437,356,1391]" pageId="3" pageNumber="404">
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Morphology and taxonomy of the
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<taxonomicName id="4C614D51361A34120F2DD5A3BE86BB81" authorityName="Matthews" authorityYear="1987" box="[574,700,680,706]" pageId="3" pageNumber="402" rank="subFamily" subFamily="Lemodinae">Lemodinae</taxonomicName>
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was recently treated in detail by Telnov & Degiovanni (2021: 104– 107) and is not repeated here.Adult
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<taxonomicName id="4C614D51361A34120F35D5C7B878BBA5" authority="Fairmaire & Germain, 1863" authorityName="Fairmaire & Germain" authorityYear="1863" box="[550,1090,716,743]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="nobilis">
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<emphasis id="B915EAC0361A34120F35D5C7BF3EBBA5" box="[550,772,716,742]" italics="true" pageId="3" pageNumber="404">Copobaenus nobilis</emphasis>
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<bibRefCitation id="EFF04B23361A34120E19D5C7B878BBA5" author="Fairmaire, L. & Germain, P." box="[778,1090,716,743]" pageId="3" pageNumber="404" pagination="225 - 284" refId="ref12651" refString="Fairmaire, L. & Germain, P. (1863) Revision des Coleopteres du Chili. Suite. Annales de la Societe Entomologique de France, 3 (4), 225 - 284. https: // doi. org / 10.1080 / 00379271.1863.11755447" type="journal article" year="1863">Fairmaire & Germain, 1863</bibRefCitation>
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</taxonomicName>
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,
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<typeStatus id="54DA8870361A34120958D5C6B841BBA4" box="[1099,1147,717,743]" pageId="3" pageNumber="404">type</typeStatus>
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species of
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<taxonomicName id="4C614D51361A341209E6D5C7B9A2BBA5" box="[1269,1432,716,742]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subFamily" subFamily="Copobaeninae">Copobaeninae</taxonomicName>
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, is comprised of the same set of morphological features as described for
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<taxonomicName id="4C614D51361A34120EBDD5FBB816BA49" authorityName="Matthews" authorityYear="1987" box="[942,1068,752,778]" pageId="3" pageNumber="402" rank="subFamily" subFamily="Lemodinae">Lemodinae</taxonomicName>
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by Telnov & Degiovanni (2021). So,
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<taxonomicName id="4C614D51361A34120DD1D41FBD7ABA6D" authorityName="Matthews" authorityYear="1987" box="[194,320,788,814]" pageId="3" pageNumber="402" rank="subFamily" subFamily="Lemodinae">Lemodinae</taxonomicName>
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is found to be identical with
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<taxonomicName id="4C614D51361A34120F96D41FBF1FBA6D" box="[645,805,788,814]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subFamily" subFamily="Copobaeninae">Copobaeninae</taxonomicName>
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in the present study and a new subfamily-rank synonym is introduced.
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</paragraph>
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<paragraph id="8BDE36D2361A34120DD4D457BFA0BAD9" blockId="3.[151,1437,356,1391]" pageId="3" pageNumber="404">
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Additional features of the
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<taxonomicName id="4C614D51361A34120CE6D457BEAFBA35" box="[501,661,860,886]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subFamily" subFamily="Copobaeninae">Copobaeninae</taxonomicName>
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supplementary to the list in Telnov & Degiovanni (2021) are given below (the numeration follows and supplements the former authors):
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</paragraph>
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<paragraph id="8BDE36D2361A34120DD4D4AFBD7ABAA0" blockId="3.[151,1437,356,1391]" pageId="3" pageNumber="404">i.xiii.i. Maxilla—galea and lacinia separated, galea well-developed, apically more or less widened and rounded, densely setose;</paragraph>
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<paragraph id="8BDE36D2361A34120DD4D4E7BF9BBD68" blockId="3.[151,1437,356,1391]" pageId="3" pageNumber="404">iv.ii. Mesoventrite, mesoventral intercoxal process—projecting along almost 2/3 of mesocoxal length, apically slightly bilobate (to lock with vertical ridge of metaventrite) or acute;</paragraph>
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<paragraph id="8BDE36D2361A34120DD4D33FBD0ABD31" blockId="3.[151,1437,356,1391]" pageId="3" pageNumber="404">iv.ii.i. Mesoventrite—mesothoracic glands present, ducts connect to a single pore at the anterior margin of mesoventrite;</paragraph>
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<paragraph id="8BDE36D2361A34120DD4D377BECBBDF8" blockId="3.[151,1437,356,1391]" pageId="3" pageNumber="404">
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vi.vi.i. Metathoracic wing—R cell basally open, a short remnant of RP3+4 present, almost no RP base and no wedge cell in the medial field (
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<bibRefCitation id="EFF04B23361A34120CE7D3AABEDFBDF8" author="Lawrence, J. F. & Zhou, Y. - L. & Lemann, C. & Sinclair, B. & Slipinski, A." box="[500,741,1184,1211]" pageId="3" pageNumber="404" pagination="433 - 755" refId="ref13103" refString="Lawrence, J. F., Zhou, Y. - L., Lemann, C., Sinclair, B. & Slipinski, A. (2022) The hind wing of Coleoptera (Insecta): morphology, nomenclature and phylogenetic significance: Part 2. Further discussion, Histeroidea, Bostrichoidea to Curculionoidea. Annales zoologici, 72 (3), 433 - 755. https: // doi. org / 10.3161 / 00034541 ANZ 2022.72.3.004" type="journal article" year="2022">
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Lawrence
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<emphasis id="B915EAC0361A34120F7AD3AABEA0BDF9" box="[617,666,1184,1210]" italics="true" pageId="3" pageNumber="404">et al</emphasis>
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. 2022
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</bibRefCitation>
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||
);
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</paragraph>
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<paragraph id="8BDE36D2361A34120DD4D3CFBEC5BC40" blockId="3.[151,1437,356,1391]" pageId="3" pageNumber="404">viii.iii. Abdomen, morphological sternite IX (with spiculum gastrale)—Y-shaped (misinterpreted I-shaped by Telnov & Degiovanni (2021)), with long or short stem;</paragraph>
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<paragraph id="8BDE36D2361A34120DD4D207BE35BC09" blockId="3.[151,1437,356,1391]" pageId="3" pageNumber="404">x.i. Male genital organs—phallobase not present or short and/or fused with tegmen, penis longer than tegmen, extending anteriorly beyond of it.</paragraph>
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<paragraph id="8BDE36D2361A34120DD4D25FB83DBC2D" blockId="3.[151,1437,356,1391]" box="[199,1031,1364,1391]" pageId="3" pageNumber="404">x.ii. Ovipositor—styli small, borne on incompletely bi-segmented coxites.</paragraph>
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</subSubSection>
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</treatment>
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</document> |