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<document ID-DOI="10.1093/zoolinnean/zlac082" ID-GBIF-Dataset="f791f8ed-ae19-41a1-b5cc-b9b2cc89dec4" ID-ISSN="0024-4082" ID-Zenodo-Dep="7814127" ID-ZooBank="99842C3A-879F-4552-96B7-204D79CF3EDF" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" approvalRequired="31" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="13" approvalRequired_for_textStreams="11" approvalRequired_for_treatments="6" checkinTime="1680597737982" checkinUser="plazi" docAuthor="Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam &amp; Fikáček, Martin" docDate="2023" docId="03C2CE00FFEFFF8C3D940F63FB094054" docLanguage="en" docName="zlac082.pdf" docOrigin="Zoological Journal of the Linnean Society 197 (4)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlac082" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Karumbe pakaluki Arriaga-Varela, Tomaszewska, Szawaryn, Robertson, Seidel, Ślipiński &amp; Fikáček, 2023, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="1101" masterDocId="FFFBB678FFF9FF9B3E190B0AFFBD416E" masterDocTitle="The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence" masterLastPageNumber="1115" masterPageNumber="1078" pageNumber="1100" updateTime="1681148941709" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Arriaga-Varela, Emmanuel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tomaszewska, Wioletta</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Szawaryn, Karol</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Robertson, James</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seidel, Matthias</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ślipiński, Adam</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fikáček, Martin</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>197</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1078</mods:start>
<mods:end>1115</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1093/zoolinnean/zlac082</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1093/zoolinnean/zlac082</mods:identifier>
<mods:identifier type="GBIF-Dataset">f791f8ed-ae19-41a1-b5cc-b9b2cc89dec4</mods:identifier>
<mods:identifier type="ISSN">0024-4082</mods:identifier>
<mods:identifier type="Zenodo-Dep">7814127</mods:identifier>
<mods:identifier type="ZooBank">99842C3A-879F-4552-96B7-204D79CF3EDF</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7814193" ID-Zenodo-Dep="7814193" LSID="urn:lsid:plazi:treatment:03C2CE00FFEFFF8C3D940F63FB094054" httpUri="http://treatment.plazi.org/id/03C2CE00FFEFFF8C3D940F63FB094054" lastPageId="23" lastPageNumber="1101" pageId="22" pageNumber="1100">
<subSubSection pageId="22" pageNumber="1100" type="nomenclature">
<paragraph blockId="22.[909,1360,1129,1184]" box="[909,1360,1129,1154]" pageId="22" pageNumber="1100">
<heading box="[909,1360,1129,1154]" centered="true" fontSize="9" level="2" pageId="22" pageNumber="1100" reason="2">
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[909,1148,1129,1152]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1113" phylum="Arthropoda" rank="species" species="pakaluki">
<emphasis bold="true" box="[909,1148,1129,1152]" italics="true" pageId="22" pageNumber="1100">KARUMBE PAKALUKI</emphasis>
</taxonomicName>
ARRIAGA- VARELA,
</heading>
</paragraph>
<paragraph blockId="22.[909,1360,1129,1184]" box="[919,1350,1159,1184]" pageId="22" pageNumber="1100">
<heading box="[919,1350,1159,1184]" centered="true" fontSize="9" level="2" pageId="22" pageNumber="1100" reason="4">
TOMASZEWSKA &amp; SZAWARYN
<emphasis bold="true" box="[1251,1350,1159,1183]" pageId="22" pageNumber="1100">
<taxonomicNameLabel box="[1251,1350,1159,1183]" pageId="22" pageNumber="1100" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="1101" pageId="22" pageNumber="1100" type="description">
<paragraph blockId="22.[884,1385,1198,1223]" box="[884,1385,1198,1223]" pageId="22" pageNumber="1100">
(
<figureCitation box="[892,1011,1198,1223]" captionStart="Figure 6" captionStartId="15.[145,223,1742,1764]" captionTargetBox="[321,1262,200,1690]" captionTargetId="figure-73@15.[302,1268,197,1702]" captionTargetPageId="15" captionText="Figure 6. Karumbe gen. nov. A, B, Karumbe brethesi sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Karumbe geiseri sp. nov.: C, habitus dorsal. D, habitus lateral. E, F, Karumbe pakaluki sp. nov.: E, habitus dorsal. F, habitus lateral. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.7814141" httpUri="https://zenodo.org/record/7814141/files/figure.png" pageId="22" pageNumber="1100">FIGS 6E, F</figureCitation>
,
<figureCitation box="[1024,1214,1198,1223]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="22" pageNumber="1100">7A–E, G, J, K, M</figureCitation>
,
<figureCitation box="[1226,1288,1198,1223]" captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="22" pageNumber="1100">8D–F</figureCitation>
,
<figureCitation box="[1300,1377,1198,1223]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="22" pageNumber="1100">11A–F</figureCitation>
)
</paragraph>
<paragraph blockId="22.[827,1444,1238,1291]" pageId="22" pageNumber="1100">
<emphasis box="[827,1161,1238,1260]" italics="true" pageId="22" pageNumber="1100">Zoobank registration:</emphasis>
urn: lsid: zoobank. org:act:
<uuid box="[902,1430,1269,1291]" pageId="22" pageNumber="1100">349822CB-0F6B-4737-A717-D6CD788AD9CC</uuid>
</paragraph>
<paragraph blockId="22.[827,1443,1332,1476]" pageId="22" pageNumber="1100">
<emphasis box="[827,1006,1332,1354]" italics="true" pageId="22" pageNumber="1100">Type material:</emphasis>
<materialsCitation collectionCode="CMN" collectorName="S &amp; J Peck" country="Venezuela" elevation="400" location="Guatopo" pageId="22" pageNumber="1100" specimenCount="1" stateProvince="Miranda" typeStatus="holotype">
<typeStatus box="[1020,1131,1332,1354]" pageId="22" pageNumber="1100">Holotype</typeStatus>
, male,
<collectingCountry box="[1221,1361,1332,1353]" name="Venezuela" pageId="22" pageNumber="1100">VENEZUELA</collectingCountry>
: VEN:
<collectingRegion box="[827,933,1362,1384]" country="Venezuela" name="Miranda" pageId="22" pageNumber="1100">Miranda</collectingRegion>
:
<quantity box="[946,1022,1363,1384]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="1100" unit="m" value="400.0">
<elevation box="[946,1022,1363,1384]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="1100" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<locationDeviation box="[1036,1278,1362,1384]" location="Altagracia" pageId="22" pageNumber="1100">
<quantity box="[1036,1111,1362,1384]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="22" pageNumber="1100" unit="km" value="35.0">35 km</quantity>
N Altagracia
</locationDeviation>
,
<location LSID="urn:lsid:plazi:treatment:03C2CE00FFEFFF8C3D940F63FB094054:8EB429CDFFEFFF8D3B150E59FACC4406" box="[1292,1393,1363,1384]" country="Venezuela" name="Guatopo" pageId="22" pageNumber="1100" stateProvince="Miranda">Guatopo</location>
NP,
<collectorName box="[827,978,1393,1415]" pageId="22" pageNumber="1100">Agua Blanca</collectorName>
, 31.V-7.VI.87 -2,
<collectorName box="[1179,1293,1393,1415]" pageId="22" pageNumber="1100">S&amp;J Peck</collectorName>
, ravine FITs (
<collectionCode box="[834,901,1424,1445]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="22" pageNumber="1100" type="Museum">CMN</collectionCode>
)
</materialsCitation>
.
<materialsCitation collectionCode="MIZ" collectorName="S &amp; J Peck" country="Venezuela" elevation="400" location="Guatopo" pageId="22" pageNumber="1100" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Miranda" typeStatus="paratype">
<typeStatus box="[921,1042,1424,1445]" pageId="22" pageNumber="1100">Paratypes</typeStatus>
: same data as holotype (
<specimenCount box="[1336,1443,1424,1446]" pageId="22" pageNumber="1100" type="male">one male</specimenCount>
and 2 ex, CMN;
<specimenCount box="[1009,1132,1454,1476]" pageId="22" pageNumber="1100" type="female">one female</specimenCount>
and 1 ex,
<collectionCode box="[1248,1301,1455,1476]" pageId="22" pageNumber="1100">MIZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="22.[827,1444,1525,1884]" lastBlockId="23.[145,762,197,372]" lastPageId="23" lastPageNumber="1101" pageId="22" pageNumber="1100">
<emphasis box="[827,1156,1525,1547]" italics="true" pageId="22" pageNumber="1100">
Diagnosis:
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[955,1156,1525,1546]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1100" phylum="Arthropoda" rank="species" species="pakaluki">Karumbe pakaluki</taxonomicName>
</emphasis>
is most similar to
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[1346,1443,1526,1547]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1100" phylum="Arthropoda" rank="species" species="geiseri">
<emphasis box="[1346,1443,1526,1547]" italics="true" pageId="22" pageNumber="1100">K. geiseri</emphasis>
</taxonomicName>
but can be distinguished from it by having the narrower prosternal process being about 0.67 as wide as procoxal diameter (
<figureCitation box="[946,1043,1617,1639]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="22" pageNumber="1100">Fig. 11C</figureCitation>
) (
<quantity box="[1067,1133,1617,1639]" metricMagnitude="-2" metricUnit="m" metricValue="2.286" pageId="22" pageNumber="1100" unit="in" value="0.9">0.9 in</quantity>
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[1142,1248,1618,1639]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1100" phylum="Arthropoda" rank="species" species="geiseri">
<emphasis box="[1142,1248,1618,1639]" italics="true" pageId="22" pageNumber="1100">K. geiseri</emphasis>
</taxonomicName>
); pronotum with posterolateral indentation shorter and more rectangular (
<figureCitation box="[834,927,1678,1700]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="22" pageNumber="1100">Fig. 11B</figureCitation>
) (longer and more oblique in
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[1252,1353,1679,1700]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1100" phylum="Arthropoda" rank="species" species="geiseri">
<emphasis box="[1252,1353,1679,1700]" italics="true" pageId="22" pageNumber="1100">K. geiseri</emphasis>
</taxonomicName>
) and by the shape of the aedeagus (
<figureCitation box="[1135,1228,1709,1731]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="22" pageNumber="1100">Fig. 10E</figureCitation>
). From
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[1318,1437,1709,1731]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1100" phylum="Arthropoda" rank="species" species="brethesi">
<emphasis box="[1318,1437,1709,1731]" italics="true" pageId="22" pageNumber="1100">K. brethesi</emphasis>
</taxonomicName>
, the third species of the genus,
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[1164,1294,1740,1761]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1100" phylum="Arthropoda" rank="species" species="pakaluki">
<emphasis box="[1164,1294,1740,1761]" italics="true" pageId="22" pageNumber="1100">K. pakaluki</emphasis>
</taxonomicName>
can be easily distinguished by having dorsal surface covered with setiferous punctures and additional foveolate depressions without setae (
<figureCitation box="[980,1068,1832,1854]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="22" pageNumber="1100">Fig.11D</figureCitation>
) (in
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[1111,1222,1832,1853]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1100" phylum="Arthropoda" rank="species" species="brethesi">
<emphasis box="[1111,1222,1832,1853]" italics="true" pageId="22" pageNumber="1100">K.brethesi</emphasis>
</taxonomicName>
peculiar punctations as in
<figureCitation box="[883,958,1862,1884]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="22" pageNumber="1100">Fig. 9E</figureCitation>
), the central area of the metaventrite covered only with simple setiferous punctures (
<figureCitation box="[604,700,197,219]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="23" pageNumber="1101">Fig. 11F</figureCitation>
), the pronotal disc much less concave along the internal lateral carina (
<figureCitation box="[227,316,258,280]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="23" pageNumber="1101">Fig. 11B</figureCitation>
) and the terminal labial palpomere short and apically rounded, 1.2 longer than wide (
<figureCitation box="[635,716,289,311]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="23" pageNumber="1101">Fig. 7D</figureCitation>
) (in
<emphasis box="[145,268,320,341]" italics="true" pageId="23" pageNumber="1101">
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[145,262,320,341]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1101" phylum="Arthropoda" rank="species" species="brethesi">K. brethesi</taxonomicName>
:
</emphasis>
elongate and acuminate, 2.2 as long as wide as in
<figureCitation box="[202,278,350,372]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="23" pageNumber="1101">Fig. 9B</figureCitation>
).
</paragraph>
<paragraph blockId="23.[145,762,424,1917]" pageId="23" pageNumber="1101">
<emphasis box="[145,280,424,445]" italics="true" pageId="23" pageNumber="1101">Description:</emphasis>
Body: length
<quantity box="[441,601,424,446]" metricMagnitude="-4" metricUnit="m" metricValue="9.299999999999999" metricValueMax="9.5" metricValueMin="9.1" pageId="23" pageNumber="1101" unit="mm" value="0.9299999999999999" valueMax="0.95" valueMin="0.91">0.91–0.95 mm</quantity>
, 1.30 times as long as wide, 2.0 times as long as high, short oval and moderately convex, dark brown with yellowish legs, antennae and palpi (
<figureCitation box="[379,487,516,538]" captionStart="Figure 6" captionStartId="15.[145,223,1742,1764]" captionTargetBox="[321,1262,200,1690]" captionTargetId="figure-73@15.[302,1268,197,1702]" captionTargetPageId="15" captionText="Figure 6. Karumbe gen. nov. A, B, Karumbe brethesi sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Karumbe geiseri sp. nov.: C, habitus dorsal. D, habitus lateral. E, F, Karumbe pakaluki sp. nov.: E, habitus dorsal. F, habitus lateral. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.7814141" httpUri="https://zenodo.org/record/7814141/files/figure.png" pageId="23" pageNumber="1101">Fig. 6E, F</figureCitation>
).
</paragraph>
<paragraph blockId="23.[145,762,424,1917]" pageId="23" pageNumber="1101">
Antenna nearly 0.35 of length of body (
<figureCitation box="[604,690,546,568]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="23" pageNumber="1101">Figs 7A</figureCitation>
,
<figureCitation box="[702,747,547,568]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="23" pageNumber="1101">11A</figureCitation>
); antennomeres 1–5 longer than wide; antennomeres 6, 7, 9 subquadrate and antennomere 8 transverse. Apical labial palpomere short and rounded, 1.2 times as long as wide (
<figureCitation box="[244,326,669,691]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="23" pageNumber="1101">Fig. 7D</figureCitation>
).
</paragraph>
<paragraph blockId="23.[145,762,424,1917]" pageId="23" pageNumber="1101">
Pronotum 2.1 times as wide as long, 1.7 times wider at widest part than on front angles (
<figureCitation box="[648,745,730,752]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="23" pageNumber="1101">Fig. 11B</figureCitation>
). Anterior crenulate margin comparatively wide. Area between lateral edge and internal lateral carina weakly concave, wide and continuously weakly narrowing posteriorly; pronotal disc not concave along internal lateral carina. Basal pronotal bordering line shallowly and irregularly crenulate. Posterolateral indentations comparatively shallow, rectangular. Prosternal process wide (
<figureCitation box="[637,744,976,998]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="23" pageNumber="1101">Fig. 11C</figureCitation>
), with apex about 0.67 of width of procoxal cavity, narrowest near half-length, weakly widened apically, with lateral carinae raised, reaching almost anterior quarter of prosternum, central part between carinae slightly depressed.
</paragraph>
<paragraph blockId="23.[145,762,424,1917]" pageId="23" pageNumber="1101">
Elytra 0.70–
<quantity box="[341,457,1160,1182]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="23" pageNumber="1101" unit="mm" value="0.75">0.75 mm</quantity>
long, about as long a wide, 3.1 times as long and 1.4 times as wide as pronotum; lateral margins visible from above (
<figureCitation captionStart="Figure 6" captionStartId="15.[145,223,1742,1764]" captionTargetBox="[321,1262,200,1690]" captionTargetId="figure-73@15.[302,1268,197,1702]" captionTargetPageId="15" captionText="Figure 6. Karumbe gen. nov. A, B, Karumbe brethesi sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Karumbe geiseri sp. nov.: C, habitus dorsal. D, habitus lateral. E, F, Karumbe pakaluki sp. nov.: E, habitus dorsal. F, habitus lateral. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.7814141" httpUri="https://zenodo.org/record/7814141/files/figure.png" pageId="23" pageNumber="1101">Fig. 6E</figureCitation>
). Elytral dorsal surface covered with simple setiferous punctures and additional foveolate depressions without setae (
<figureCitation box="[477,578,1313,1335]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="23" pageNumber="1101">Fig. 11D</figureCitation>
). Metaventrite with fine setiferous punctures at centre (
<figureCitation box="[646,745,1344,1366]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="23" pageNumber="1101">Fig. 11F</figureCitation>
). Mesoventral process about 1.3 times as wide as mesocoxal diameter. Wings well developed.
</paragraph>
<paragraph blockId="23.[145,762,424,1917]" pageId="23" pageNumber="1101">Legs. Meso- and metatrochanters flattened, roundly produced posteriorly.</paragraph>
<paragraph blockId="23.[145,762,424,1917]" pageId="23" pageNumber="1101">
Abdomen. Ventrite 1 with rounded and laterally complete postcoxal lines (
<figureCitation box="[437,532,1528,1550]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="23" pageNumber="1101">Fig. 11F</figureCitation>
). Ventrite
<quantity box="[654,698,1527,1550]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="23" pageNumber="1101" unit="in" value="5.0">5 in</quantity>
male triangularly produced posteriorly at middle, in female straight. Male genital segment as in
<figureCitation box="[559,640,1589,1611]" captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="23" pageNumber="1101">Fig. 8D</figureCitation>
.
</paragraph>
<paragraph blockId="23.[145,762,424,1917]" pageId="23" pageNumber="1101">
Male genitalia (
<figureCitation box="[352,434,1620,1642]" captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="23" pageNumber="1101">Fig. 8E</figureCitation>
). Aedeagus with penis short and stout, narrow at base, widening towards curved, ramificate apex. Tegmen throne-shaped, long; tegminal strut short, reduced.
</paragraph>
<paragraph blockId="23.[145,762,424,1917]" pageId="23" pageNumber="1101">
Female genitalia. Bursa copulatrix large, with apical outlet of sperm duct. Spermatheca (
<figureCitation captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="23" pageNumber="1101">Fig. 8F</figureCitation>
) submembranous, moderately large, elongatesubcylindrical, oblong; sperm duct long, about as long as spermatheca; accessory gland small, membranous, of irregular shape.
</paragraph>
<paragraph blockId="23.[809,1426,197,249]" pageId="23" pageNumber="1101">
<emphasis box="[809,934,197,218]" italics="true" pageId="23" pageNumber="1101">Etymology:</emphasis>
This species is dedicated to the late James Pakaluk, who had keen interest in small brown beetles.
</paragraph>
<paragraph blockId="23.[809,1204,293,315]" box="[809,1204,293,315]" pageId="23" pageNumber="1101">
<emphasis box="[809,953,293,314]" italics="true" pageId="23" pageNumber="1101">Distribution:</emphasis>
<collectingCountry box="[965,1080,293,315]" name="Venezuela" pageId="23" pageNumber="1101">Venezuela</collectingCountry>
(
<figureCitation box="[1095,1189,293,315]" captionStart="Figure 20" captionStartId="34.[162,240,1081,1103]" captionTargetBox="[162,1442,200,1042]" captionTargetId="figure-327@34.[162,1442,200,1042]" captionTargetPageId="34" captionText="Figure 20. Geographical distribution of Cerasommatidiidae spp. A, distribution of species in eastern Caribbean region. B, distribution of species in southeastern Brazil. C, distribution of species in Madagascar." figureDoi="http://doi.org/10.5281/zenodo.7814187" httpUri="https://zenodo.org/record/7814187/files/figure.png" pageId="23" pageNumber="1101">Fig. 20A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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