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<document id="817E6B6DDDA718468A4E9C4CBF45501B" ID-DOI="10.37828/em.2021.48.7" ID-ISSN="2336-9744" ID-Zenodo-Dep="13250168" ID-ZooBank="urn:lsid:zoobank.org:pub:243E3971-5340-4129-AFA8-B6A9449B9D3F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722975665387" checkinUser="felipe" docAuthor="Volynkin, Anton V." docDate="2021" docId="03BCE764FFBDFFF10FA6FDB4C92D1728" docLanguage="en" docName="EcolMontenegrina.48.39-42.pdf" docOrigin="Ecologica Montenegrina 48" docSource="http://dx.doi.org/10.37828/em.2021.48.7" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Cyana (Cryptanaema) halmahera Volynkin, 2021, sp. n." docType="treatment" docUuid="E1A2D3CA-58DC-45EE-AE72-9325778DAA00" docUuidSource="ZooBank" docVersion="2" lastPageNumber="42" masterDocId="FF859F1CFFBCFFF20F31FF83CD21160A" masterDocTitle="Cyana (Cryptanaema) halmahera, a new species from the Maluku Islands (Lepidoptera: Erebidae: Arctiinae: Lithosiini)" masterLastPageNumber="42" masterPageNumber="39" pageNumber="40" updateTime="1722985426343" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9C914EC7277B2F2AC2DD92A859143381">Cyana (Cryptanaema) halmahera, a new species from the Maluku Islands (Lepidoptera: Erebidae: Arctiinae: Lithosiini)</mods:title>
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<mods:namePart id="C286C86AB3BC7EB9AE74A0315B06912D">Volynkin, Anton V.</mods:namePart>
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<mods:title id="E3821B57829F3A0D041617ADEC438094">Ecologica Montenegrina</mods:title>
</mods:titleInfo>
<mods:part id="FAE41A506FAEC04A7FC0B74A899F1323">
<mods:date id="9CE1BC3A8BB26295B46313561A0CA89A">2021</mods:date>
<mods:detail id="779B61F39D84B9ACDAA32D544964625F" type="pubDate">
<mods:number id="F224D0678D5EF1BE0358499CFB2DEB95">2021-11-25</mods:number>
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<mods:number id="40E00748D27229D9B90B0CDF7E7CAD2C">48</mods:number>
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<mods:classification id="AF7ACD178246651CD33885FFC8115D49">journal article</mods:classification>
<mods:identifier id="411D41B4F69F92EC1512FB8EDBA16199" type="DOI">10.37828/em.2021.48.7</mods:identifier>
<mods:identifier id="38848B6E1CC358E3553FB64EDE95E78B" type="ISSN">2336-9744</mods:identifier>
<mods:identifier id="24FD11FFC59CD264BE4FCD6FF6146427" type="Zenodo-Dep">13250168</mods:identifier>
<mods:identifier id="B2C7FAA73E74312A11359DFF9C5898AF" type="ZooBank">urn:lsid:zoobank.org:pub:243E3971-5340-4129-AFA8-B6A9449B9D3F</mods:identifier>
</mods:mods>
<treatment id="03BCE764FFBDFFF10FA6FDB4C92D1728" LSID="urn:lsid:zoobank.org:act:E1A2D3CA-58DC-45EE-AE72-9325778DAA00" httpUri="http://treatment.plazi.org/id/03BCE764FFBDFFF10FA6FDB4C92D1728" lastPageId="3" lastPageNumber="42" pageId="1" pageNumber="40">
<subSubSection id="C30F05F9FFBDFFF30FA6FDB4C9A1147A" pageId="1" pageNumber="40" type="nomenclature">
<paragraph id="8BAA5672FFBDFFF30FA6FDB4CF58145B" blockId="1.[151,1152,567,658]" box="[151,633,567,593]" pageId="1" pageNumber="40">
<heading id="D0E2E11EFFBDFFF30FA6FDB4CF58145B" box="[151,633,567,593]" fontSize="11" level="2" pageId="1" pageNumber="40" reason="2">
<emphasis id="B9618A60FFBDFFF30FA6FDB4CF58145B" bold="true" box="[151,633,567,593]" pageId="1" pageNumber="40">
<taxonomicName id="4C152DF1FFBDFFF30FA6FDB4CF0E145B" ID-CoL="BFNRR" authorityName="Volynkin" authorityYear="2021" box="[151,559,567,593]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="halmahera" status="sp. nov." subGenus="Cryptanaema">
<emphasis id="B9618A60FFBDFFF30FA6FDB4CDC4145B" bold="true" box="[151,229,567,593]" italics="true" pageId="1" pageNumber="40">Cyana</emphasis>
(
<emphasis id="B9618A60FFBDFFF30FC6FDB4CCBB145B" bold="true" box="[247,410,567,593]" italics="true" pageId="1" pageNumber="40">Cryptanaema</emphasis>
)
<emphasis id="B9618A60FFBDFFF30E9AFDB4CF0E145B" bold="true" box="[427,559,567,593]" italics="true" pageId="1" pageNumber="40">halmahera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252371BFFBDFFF30D07FDB4CF58145B" box="[566,633,567,593]" pageId="1" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA5672FFBDFFF30FA6FDDBC9A1147A" blockId="1.[151,1152,567,658]" box="[151,1152,600,624]" pageId="1" pageNumber="40">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="C30F05F9FFBDFFF30FA6FDF4CC301498" box="[151,273,631,658]" pageId="1" pageNumber="40" type="description">
<paragraph id="8BAA5672FFBDFFF30FA6FDF4CC301498" blockId="1.[151,1152,567,658]" box="[151,273,631,658]" pageId="1" pageNumber="40">
(
<figureCitation id="132E4AF7FFBDFFF30FAEFDF4CC261498" box="[159,263,631,658]" captionStart="Figures 1–4" captionStartId="2.[151,237,1968,1992]" captionTargetBox="[176,1424,190,1916]" captionTargetId="figure-18@2.[151,1436,189,1928]" captionTargetPageId="2" captionText="Figures 1–4. Cyana (Cryptanaema) spp.: adults (1, 2) and male genitalia (3, 4). Depositories of the specimens and the slides: 1 and 3 in MWM/ZSM; 2 and 4 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13250170" httpUri="https://zenodo.org/record/13250170/files/figure.png" pageId="1" pageNumber="40">Figs 1, 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F05F9FFBDFFF30FA6FD38C9171556" pageId="1" pageNumber="40" type="materials_examined">
<paragraph id="8BAA5672FFBDFFF30FA6FD38CE431531" blockId="1.[151,1437,698,2007]" pageId="1" pageNumber="40">
<emphasis id="B9618A60FFBDFFF30FA6FD38CC7514DF" bold="true" box="[151,340,699,725]" pageId="1" pageNumber="40">Type material</emphasis>
.
<materialsCitation id="3B7D5C2FFFBDFFF30E5FFD38CE431531" collectingDate="1996-01-31" collectionCode="ZSM" collectorName="Tarasov &amp; No. &amp; Arct. &amp; Volynkin" country="Indonesia" county="Halmahera Mt." elevation="600" latitude="1.2" location="Primarwald" longLatPrecision="1308" longitude="127.53333" municipality="Mt. Talagaranu" pageId="1" pageNumber="40" specimenCount="1" stateProvince="Maluku Utara" typeStatus="holotype">
<emphasis id="B9618A60FFBDFFF30E5FFD38CCC114DF" bold="true" box="[366,480,699,725]" pageId="1" pageNumber="40">
<typeStatus id="54AEE8D0FFBDFFF30E5FFD38CCC114DF" box="[366,480,699,725]" pageId="1" pageNumber="40" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="132E4AF7FFBDFFF30ECAFD39CF5D14DF" box="[507,636,698,725]" captionStart="Figures 1–4" captionStartId="2.[151,237,1968,1992]" captionTargetBox="[176,1424,190,1916]" captionTargetId="figure-18@2.[151,1436,189,1928]" captionTargetPageId="2" captionText="Figures 1–4. Cyana (Cryptanaema) spp.: adults (1, 2) and male genitalia (3, 4). Depositories of the specimens and the slides: 1 and 3 in MWM/ZSM; 2 and 4 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13250170" httpUri="https://zenodo.org/record/13250170/files/figure.png" pageId="1" pageNumber="40">Figs 1, 3</figureCitation>
): male, [
<collectingCountry id="F30216E2FFBDFFF30DC8FD39CE5014DF" box="[761,881,698,725]" name="Indonesia" pageId="1" pageNumber="40">Indonesia</collectingCountry>
,
<collectingRegion id="49D19890FFBDFFF30CB9FD39C99B14DF" box="[904,1210,698,725]" country="Indonesia" name="Maluku Utara" pageId="1" pageNumber="40">North Maluku Province</collectingRegion>
, Maluku Islands] “
<collectingCountry id="F30216E2FFBDFFF30F90FD5FCC3E14FD" box="[161,287,732,759]" name="Indonesia" pageId="1" pageNumber="40">Molukken</collectingCountry>
,
<collectingCounty id="62CB2EFEFFBDFFF30E1FFD5FCCC314FD" box="[302,482,732,759]" pageId="1" pageNumber="40">
Halmahera
<collectingMunicipality id="6BCECC08FFBDFFF30E89FD5ECCC314FD" box="[440,482,733,759]" pageId="1" pageNumber="40">Mt.</collectingMunicipality>
</collectingCounty>
Talagaranu,
<quantity id="4CEDFB97FFBDFFF30DB0FD5FCFE214FD" box="[641,707,732,759]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="40" unit="m" value="600.0">
<elevation id="0038B141FFBDFFF30DB0FD5FCFE214FD" box="[641,707,732,759]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="40" unit="m" value="600.0">600m</elevation>
</quantity>
<locationDeviation id="9A184F50FFBDFFF30DFCFD5FCE5E14FD" box="[717,895,732,759]" location="Baru" pageId="1" pageNumber="40">
<quantity id="4CEDFB97FFBDFFF30DFCFD5FCE2E14FD" box="[717,783,732,759]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="40" unit="km" value="15.0">15km</quantity>
SE Baru
</locationDeviation>
(
<location id="8ECA00A9FFBDFFF30CB7FD5FC93014FD" LSID="urn:lsid:plazi:treatment:03BCE764FFBDFFF10FA6FDB4C92D1728:8ECA00A9FFBDFFF30CB7FD5FC93014FD" box="[902,1041,732,759]" country="Indonesia" county="Halmahera Mt." latitude="1.2" longLatPrecision="1308" longitude="127.53333" municipality="Mt. Talagaranu" name="Primarwald" pageId="1" pageNumber="40" stateProvince="Maluku Utara">Primärwald</location>
[primary forest]), 22.–
<date id="FFAB70B2FFBDFFF30A16FD5FC8BD14FD" box="[1319,1436,732,759]" pageId="1" pageNumber="40" value="1996-01-31">
<collectingDate id="EFEF895AFFBDFFF30A16FD5FC8BD14FD" box="[1319,1436,732,759]" pageId="1" pageNumber="40" value="1996-01-31">31.1.1996</collectingDate>
</date>
1’12’N – 127’32’E [
<geoCoordinate id="EE2130B5FFBDFFF30EA4FD7DCCCA1513" box="[405,491,766,793]" degrees="1" direction="north" minutes="12" orientation="latitude" pageId="1" pageNumber="40" precision="925" value="1.2">1°12'N</geoCoordinate>
,
<geoCoordinate id="EE2130B5FFBDFFF30ECDFD7DCF4C1513" box="[508,621,766,793]" degrees="127" direction="east" minutes="32" orientation="longitude" pageId="1" pageNumber="40" precision="925" value="127.53333">127°32'E</geoCoordinate>
], lgSinjaev &amp; Tarasov [Sinyaev &amp;
<collectorName id="26E033A4FFBDFFF30B15FD7CC9A31513" box="[1060,1154,767,793]" pageId="1" pageNumber="40">Tarasov</collectorName>
leg.]”, gen. prep.
<collectorName id="26E033A4FFBDFFF30A59FD7CC8B61513" box="[1384,1431,767,793]" pageId="1" pageNumber="40">No.</collectorName>
:
<collectionCode id="ED04CEB7FFBDFFF30FA6FCA3CDF21531" box="[151,211,800,827]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="40" type="Museum">ZSM</collectionCode>
<collectorName id="26E033A4FFBDFFF30FEAFCA3CC361531" box="[219,279,800,827]" pageId="1" pageNumber="40">Arct.</collectorName>
2021-024 (prepared by
<collectorName id="26E033A4FFBDFFF30D03FCA3CF821531" box="[562,675,800,827]" pageId="1" pageNumber="40">Volynkin</collectorName>
) (MWM/
<collectionCode id="ED04CEB7FFBDFFF30C26FCA3CE771531" box="[791,854,800,827]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="40" type="Museum">ZSM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAA5672FFBDFFF30FC7FCC1C9171556" blockId="1.[151,1437,698,2007]" box="[246,1078,833,860]" pageId="1" pageNumber="40">
<materialsCitation id="3B7D5C2FFFBDFFF30FC7FCC1C9131556" box="[246,1074,833,860]" collectionCode="ZSM" pageId="1" pageNumber="40" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B9618A60FFBDFFF30FC7FCC1CC491556" bold="true" box="[246,360,834,860]" pageId="1" pageNumber="40">
<typeStatus id="54AEE8D0FFBDFFF30FC7FCC1CC491556" box="[246,360,834,860]" pageId="1" pageNumber="40" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCount id="9D139DFBFFBDFFF30E46FCC2CCE71556" box="[375,454,833,860]" count="1" pageId="1" pageNumber="40" type="male">1 male</specimenCount>
with the same data as in the
<typeStatus id="54AEE8D0FFBDFFF30C28FCC2CE5E1556" box="[793,895,833,860]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
(MWM/
<collectionCode id="ED04CEB7FFBDFFF30CDDFCC2C90A1556" box="[1004,1067,833,860]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="40" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F05F9FFBDFFF30FC7FCE0CC29135C" pageId="1" pageNumber="40" type="diagnosis">
<paragraph id="8BAA5672FFBDFFF30FC7FCE0CC29135C" blockId="1.[151,1437,698,2007]" pageId="1" pageNumber="40">
<emphasis id="B9618A60FFBDFFF30FC7FCE0CC4E1577" bold="true" box="[246,367,867,893]" pageId="1" pageNumber="40">Diagnosis</emphasis>
. The new species (
<figureCitation id="132E4AF7FFBDFFF30D64FCE0CFBB1574" box="[597,666,867,894]" captionStart="Figures 1–4" captionStartId="2.[151,237,1968,1992]" captionTargetBox="[176,1424,190,1916]" captionTargetId="figure-18@2.[151,1436,189,1928]" captionTargetPageId="2" captionText="Figures 1–4. Cyana (Cryptanaema) spp.: adults (1, 2) and male genitalia (3, 4). Depositories of the specimens and the slides: 1 and 3 in MWM/ZSM; 2 and 4 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13250170" httpUri="https://zenodo.org/record/13250170/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
) is externally reminiscent of
<taxonomicName id="4C152DF1FFBDFFF30CCDFCE7C9491574" authorityName="de Vos" authorityYear="2017" baseAuthorityName="Hampson" baseAuthorityYear="1900" box="[1020,1128,867,894]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="B9618A60FFBDFFF30CCDFCE7C9491574" box="[1020,1128,867,894]" italics="true" pageId="1" pageNumber="40">C. fumea</emphasis>
</taxonomicName>
(
<figureCitation id="132E4AF7FFBDFFF30B48FCE0C9E01574" box="[1145,1217,867,894]" captionStart="Figures 1–4" captionStartId="2.[151,237,1968,1992]" captionTargetBox="[176,1424,190,1916]" captionTargetId="figure-18@2.[151,1436,189,1928]" captionTargetPageId="2" captionText="Figures 1–4. Cyana (Cryptanaema) spp.: adults (1, 2) and male genitalia (3, 4). Depositories of the specimens and the slides: 1 and 3 in MWM/ZSM; 2 and 4 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13250170" httpUri="https://zenodo.org/record/13250170/files/figure.png" pageId="1" pageNumber="40">Fig. 2</figureCitation>
) but is recognised by the more elongate forewing and the indistinct transverse lines. Compared to
<taxonomicName id="4C152DF1FFBDFFF30B57FC05C9E51595" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1904" box="[1126,1220,902,928]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="aroa">
<emphasis id="B9618A60FFBDFFF30B57FC05C9E51595" box="[1126,1220,902,928]" italics="true" pageId="1" pageNumber="40">C. aroa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C152DF1FFBDFFF30A32FC05C8BD15AA" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1904" box="[1283,1436,901,928]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="charybdis">
<emphasis id="B9618A60FFBDFFF30A32FC05C8BD15AA" box="[1283,1436,901,928]" italics="true" pageId="1" pageNumber="40">C. charybdis</emphasis>
</taxonomicName>
(illustrated by
<bibRefCitation id="EF842B83FFBDFFF30E72FC25CCD615CB" author="References de Vos" box="[323,503,934,961]" pageId="1" pageNumber="40" pagination="126 - 143" refId="ref1485" refString="References de Vos, R. (2017) The Cyana Walker, 1854 species of New Guinea, with description of a new subgenus, eight new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Lithosiini). In: Telnov, D., Barclay, M. V. L. &amp; Pauwels, O. S. G. (Eds.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea. Volume III. The Entomological Society of Latvia, Riga, pp. 587 - 611, pls. 126 - 143." type="book chapter" year="2017">de Vos (2017))</bibRefCitation>
,
<taxonomicName id="4C152DF1FFBDFFF30D35FC24CF8715CB" authorityName="Volynkin" authorityYear="2021" box="[516,678,934,961]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="halmahera" status="sp. n.">
<emphasis id="B9618A60FFBDFFF30D35FC24CF8715CB" box="[516,678,934,961]" italics="true" pageId="1" pageNumber="40">C. halmahera</emphasis>
</taxonomicName>
<emphasis id="B9618A60FFBDFFF30D9CFC24CFD115CB" bold="true" box="[685,752,935,961]" pageId="1" pageNumber="40">
<taxonomicNameLabel id="A252371BFFBDFFF30D9CFC24CFD115CB" box="[685,752,935,961]" pageId="1" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has a pale creamy basal and medial areas of the hindwing whereas the hindwing of
<taxonomicName id="4C152DF1FFBDFFF30EFAFC4ACF0915E8" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1904" box="[459,552,969,995]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="aroa">
<emphasis id="B9618A60FFBDFFF30EFAFC4ACF0915E8" box="[459,552,969,995]" italics="true" pageId="1" pageNumber="40">C. aroa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C152DF1FFBDFFF30D56FC4ACFDE15E9" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1904" box="[615,767,968,995]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="charybdis">
<emphasis id="B9618A60FFBDFFF30D56FC4ACFDE15E9" box="[615,767,968,995]" italics="true" pageId="1" pageNumber="40">C. charybdis</emphasis>
</taxonomicName>
is monotonous brown. The male genital capsule of the new species (
<figureCitation id="132E4AF7FFBDFFF30E04FC69CC5A120F" box="[309,379,1002,1029]" captionStart="Figures 1–4" captionStartId="2.[151,237,1968,1992]" captionTargetBox="[176,1424,190,1916]" captionTargetId="figure-18@2.[151,1436,189,1928]" captionTargetPageId="2" captionText="Figures 1–4. Cyana (Cryptanaema) spp.: adults (1, 2) and male genitalia (3, 4). Depositories of the specimens and the slides: 1 and 3 in MWM/ZSM; 2 and 4 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13250170" httpUri="https://zenodo.org/record/13250170/files/figure.png" pageId="1" pageNumber="40">Fig. 3</figureCitation>
) differs from those of the similar congeners in the apically rectangular vinculum whereas it is U-shaped in
<taxonomicName id="4C152DF1FFBDFFF30E52FB8FCCEC122C" authorityName="de Vos" authorityYear="2017" baseAuthorityName="Hampson" baseAuthorityYear="1900" box="[355,461,1035,1062]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="B9618A60FFBDFFF30E52FB8FCCEC122C" box="[355,461,1035,1062]" italics="true" pageId="1" pageNumber="40">C. fumea</emphasis>
</taxonomicName>
(
<figureCitation id="132E4AF7FFBDFFF30EEFFB88CF06122C" box="[478,551,1035,1062]" captionStart="Figures 1–4" captionStartId="2.[151,237,1968,1992]" captionTargetBox="[176,1424,190,1916]" captionTargetId="figure-18@2.[151,1436,189,1928]" captionTargetPageId="2" captionText="Figures 1–4. Cyana (Cryptanaema) spp.: adults (1, 2) and male genitalia (3, 4). Depositories of the specimens and the slides: 1 and 3 in MWM/ZSM; 2 and 4 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13250170" httpUri="https://zenodo.org/record/13250170/files/figure.png" pageId="1" pageNumber="40">Fig. 4</figureCitation>
),
<taxonomicName id="4C152DF1FFBDFFF30D0FFB8FCFB9122C" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1904" box="[574,664,1036,1062]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="aroa">
<emphasis id="B9618A60FFBDFFF30D0FFB8FCFB9122C" box="[574,664,1036,1062]" italics="true" pageId="1" pageNumber="40">C. aroa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C152DF1FFBDFFF30DE2FB8FCE4B122C" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1904" box="[723,874,1035,1062]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="charybdis">
<emphasis id="B9618A60FFBDFFF30DE2FB8FCE4B122C" box="[723,874,1035,1062]" italics="true" pageId="1" pageNumber="40">C. charybdis</emphasis>
</taxonomicName>
(illustrated by
<bibRefCitation id="EF842B83FFBDFFF30B2CFB88C9F4122C" author="References de Vos" box="[1053,1237,1035,1062]" pageId="1" pageNumber="40" pagination="126 - 143" refId="ref1485" refString="References de Vos, R. (2017) The Cyana Walker, 1854 species of New Guinea, with description of a new subgenus, eight new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Lithosiini). In: Telnov, D., Barclay, M. V. L. &amp; Pauwels, O. S. G. (Eds.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea. Volume III. The Entomological Society of Latvia, Riga, pp. 587 - 611, pls. 126 - 143." type="book chapter" year="2017">de Vos (2017))</bibRefCitation>
. Compared to
<taxonomicName id="4C152DF1FFBDFFF30AB3FB8FCDFE1242" authorityName="de Vos" authorityYear="2017" baseAuthorityName="Hampson" baseAuthorityYear="1900" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="B9618A60FFBDFFF30AB3FB8FCDFE1242" italics="true" pageId="1" pageNumber="40">C. fumea</emphasis>
</taxonomicName>
, the male genital capsule of
<taxonomicName id="4C152DF1FFBDFFF30D72FBADCFC61242" authorityName="Volynkin" authorityYear="2021" box="[579,743,1069,1096]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="halmahera" status="sp. n.">
<emphasis id="B9618A60FFBDFFF30D72FBADCFC61242" box="[579,743,1069,1096]" italics="true" pageId="1" pageNumber="40">C. halmahera</emphasis>
</taxonomicName>
<emphasis id="B9618A60FFBDFFF30DC3FBADCE161242" bold="true" box="[754,823,1070,1096]" pageId="1" pageNumber="40">
<taxonomicNameLabel id="A252371BFFBDFFF30DC3FBADCE161242" box="[754,823,1070,1096]" pageId="1" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has a basally thinner uncus, a medially wider and apically narrower valva, and a longer, thinner, and more upcurved distal saccular process. The phallus of the new species differs from that of
<taxonomicName id="4C152DF1FFBDFFF30D16FBF1CFB41286" authorityName="de Vos" authorityYear="2017" baseAuthorityName="Hampson" baseAuthorityYear="1900" box="[551,661,1137,1164]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="B9618A60FFBDFFF30D16FBF1CFB41286" box="[551,661,1137,1164]" italics="true" pageId="1" pageNumber="40">C. fumea</emphasis>
</taxonomicName>
in the flat and less serrulate dorsal surface of the distal section whereas it is swollen and covered with larger denticles in
<taxonomicName id="4C152DF1FFBDFFF30C5FFB10CEFC12A7" authorityName="de Vos" authorityYear="2017" baseAuthorityName="Hampson" baseAuthorityYear="1900" box="[878,989,1170,1197]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="B9618A60FFBDFFF30C5FFB10CEFC12A7" box="[878,989,1170,1197]" italics="true" pageId="1" pageNumber="40">C. fumea</emphasis>
</taxonomicName>
. The vesicae of the two species are similar but in
<taxonomicName id="4C152DF1FFBDFFF30E0EFB36CCC312C5" authorityName="Volynkin" authorityYear="2021" box="[319,482,1204,1231]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="halmahera" status="sp. n.">
<emphasis id="B9618A60FFBDFFF30E0EFB36CCC312C5" box="[319,482,1204,1231]" italics="true" pageId="1" pageNumber="40">C. halmahera</emphasis>
</taxonomicName>
<emphasis id="B9618A60FFBDFFF30EDBFB36CF0F12C5" bold="true" box="[490,558,1205,1231]" pageId="1" pageNumber="40">
<taxonomicNameLabel id="A252371BFFBDFFF30EDBFB36CF0F12C5" box="[490,558,1205,1231]" pageId="1" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, the subbasal ventral diverticulum is narrower and membranous (whereas it is granulated in
<taxonomicName id="4C152DF1FFBDFFF30E42FB54CCFF12FB" authorityName="de Vos" authorityYear="2017" baseAuthorityName="Hampson" baseAuthorityYear="1900" box="[371,478,1238,1265]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="B9618A60FFBDFFF30E42FB54CCFF12FB" box="[371,478,1238,1265]" italics="true" pageId="1" pageNumber="40">C. fumea</emphasis>
</taxonomicName>
), the dorsal subbasal diverticulum bears a wider area of granulation, the row of cornuti is more or less straight (whereas it is distally curved in
<taxonomicName id="4C152DF1FFBDFFF30C9DFB7BC93B1319" authorityName="de Vos" authorityYear="2017" baseAuthorityName="Hampson" baseAuthorityYear="1900" box="[940,1050,1272,1299]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="B9618A60FFBDFFF30C9DFB7BC93B1319" box="[940,1050,1272,1299]" italics="true" pageId="1" pageNumber="40">C. fumea</emphasis>
</taxonomicName>
), and the distal diverticulum is membranous whereas it bears an area of granulation and a medial cluster of very short but robust cornuti in
<taxonomicName id="4C152DF1FFBDFFF30FA6FABFCC20135C" authorityName="de Vos" authorityYear="2017" baseAuthorityName="Hampson" baseAuthorityYear="1900" box="[151,257,1339,1366]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="B9618A60FFBDFFF30FA6FABFCC20135C" box="[151,257,1339,1366]" italics="true" pageId="1" pageNumber="40">C. fumea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F05F9FFBDFFF30FC7FADDCCE911DD" pageId="1" pageNumber="40" type="description">
<paragraph id="8BAA5672FFBDFFF30FC7FADDCE1A11BF" blockId="1.[151,1437,698,2007]" pageId="1" pageNumber="40">
<emphasis id="B9618A60FFBDFFF30FC7FADDCCA61372" bold="true" box="[246,391,1374,1400]" pageId="1" pageNumber="40">Description</emphasis>
.
<emphasis id="B9618A60FFBDFFF30EA9FADDCF041372" bold="true" box="[408,549,1374,1400]" pageId="1" pageNumber="40">Adult male</emphasis>
(
<figureCitation id="132E4AF7FFBDFFF30D09FADECF5E1372" box="[568,639,1373,1400]" captionStart="Figures 1–4" captionStartId="2.[151,237,1968,1992]" captionTargetBox="[176,1424,190,1916]" captionTargetId="figure-18@2.[151,1436,189,1928]" captionTargetPageId="2" captionText="Figures 1–4. Cyana (Cryptanaema) spp.: adults (1, 2) and male genitalia (3, 4). Depositories of the specimens and the slides: 1 and 3 in MWM/ZSM; 2 and 4 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13250170" httpUri="https://zenodo.org/record/13250170/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
). Forewing length
<quantity id="4CEDFB97FFBDFFF30C59FADECEE11372" box="[872,960,1373,1400]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="40" unit="mm" value="13.0">13 mm</quantity>
. Head and thorax dark brown. Antenna dark brown, ciliate. Forewing with postmedially convex costal margin and elongate and broad longitudinal cluster of androconial scales along costal margin medially. Forewing ground colour dark brown. Ante- and postmedial transverse lines sinuous, indistinct. Cilia dark brown. Hindwing pale creamy subbasally and medially, with areas of intense brownish-grey suffusion along costal and outer margins dilated at apex. Cilia brownish-grey. Abdomen pale brown, covered with ochreous-brown scales apically.
<emphasis id="B9618A60FFBDFFF30BB3F985C815102A" bold="true" box="[1154,1332,1542,1568]" pageId="1" pageNumber="40">Male genitalia</emphasis>
(
<figureCitation id="132E4AF7FFBDFFF30A77F985C8AC102B" box="[1350,1421,1542,1569]" captionStart="Figures 1–4" captionStartId="2.[151,237,1968,1992]" captionTargetBox="[176,1424,190,1916]" captionTargetId="figure-18@2.[151,1436,189,1928]" captionTargetPageId="2" captionText="Figures 1–4. Cyana (Cryptanaema) spp.: adults (1, 2) and male genitalia (3, 4). Depositories of the specimens and the slides: 1 and 3 in MWM/ZSM; 2 and 4 in CKC." figureDoi="http://doi.org/10.5281/zenodo.13250170" httpUri="https://zenodo.org/record/13250170/files/figure.png" pageId="1" pageNumber="40">Fig. 3</figureCitation>
). Uncus long and thin, downcurved and apically pointed. Tegumen relatively short with narrow arms. Vinculum more or less equal in length to tegumen, heavilu sclerotised, with laterally convex arms, apically rectangular. Valva lobular, dilated medially and tapered distally, apically rounded. Ventral plate of costa weakly sclerotised, with short thorn-like ampulla directed ventro-inwards. Sacculus relatively narrow, with long and robust, strongly upcurved and apically pointed distal process. Juxta trapezoid with weakly sclerotised anterio-medial section. Phallus tubular, somewhat dilated proximally, with very short and apically rounded coecum, distal section with cluster of serrulation dorsally. Vesica sack-like, directed ventrad, bearing long longitudinal row of robust triangular cornuti of various sizes from base of distal to tip of ventro-distal diverticulum. Dorsal subbasal diverticulum short but broad, granulate. Ventral subbasal diverticulum semiglobular, membranous. Distal diverticulum conical and apically rounded, membranous. Ventro-distal diverticulum broadly conical and apically rounded, membranous. Distal plate of vesica short and thin but heavily sclerotised. Vesica ejaculatorius directed ventrad.
</paragraph>
<paragraph id="8BAA5672FFBDFFF30FC7F83FCCE911DD" blockId="1.[151,1437,698,2007]" box="[246,456,1980,2007]" pageId="1" pageNumber="40">Female unknown.</paragraph>
</subSubSection>
<caption id="DF6A06FAFFBEFFF00FA6F833CFB711EC" ID-DOI="http://doi.org/10.5281/zenodo.13250170" ID-Zenodo-Dep="13250170" httpUri="https://zenodo.org/record/13250170/files/figure.png" pageId="2" pageNumber="41" startId="2.[151,237,1968,1992]" targetBox="[176,1424,190,1916]" targetPageId="2" targetType="figure">
<paragraph id="8BAA5672FFBEFFF00FA6F833CFB711EC" blockId="2.[151,1435,1967,2022]" pageId="2" pageNumber="41">
<emphasis id="B9618A60FFBEFFF00FA6F833CC3C11C2" bold="true" box="[151,285,1968,1992]" pageId="2" pageNumber="41">Figures 1–4</emphasis>
.
<taxonomicName id="4C152DF1FFBEFFF00E1DF833CF3D11CD" authorityName="de Vos" authorityYear="2017" box="[300,540,1967,1991]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Cryptanaema">
<emphasis id="B9618A60FFBEFFF00E1DF833CC5011CD" box="[300,369,1968,1991]" italics="true" pageId="2" pageNumber="41">Cyana</emphasis>
(
<emphasis id="B9618A60FFBEFFF00EB3F833CF3211CD" box="[386,531,1968,1991]" italics="true" pageId="2" pageNumber="41">Cryptanaema</emphasis>
)
</taxonomicName>
spp.: adults (1, 2) and male genitalia (3, 4). Depositories of the specimens and the slides: 1 and 3 in MWM/ZSM; 2 and 4 in CKC.
</paragraph>
</caption>
<subSubSection id="C30F05F9FFBFFFF10FC7FF47CF43170A" pageId="3" pageNumber="42" type="distribution">
<paragraph id="8BAA5672FFBFFFF10FC7FF47CF43170A" blockId="3.[151,1436,195,290]" pageId="3" pageNumber="42">
<emphasis id="B9618A60FFBFFFF10FC7FF47CCB116D4" bold="true" box="[246,400,196,222]" pageId="3" pageNumber="42">Distribution</emphasis>
. The new species is known only from the island of Halmahera, the
<collectingCountry id="F30216E2FFBFFFF10BD3FF40C8BD16D4" box="[1250,1436,195,222]" name="Indonesia" pageId="3" pageNumber="42">Maluku Islands</collectingCountry>
(
<collectingRegion id="49D19890FFBFFFF10FAEFF66CC96170A" box="[159,439,229,256]" country="Indonesia" name="Maluku Utara" pageId="3" pageNumber="42">North Maluku Province</collectingRegion>
of
<collectingCountry id="F30216E2FFBFFFF10EEFFF66CF77170A" box="[478,598,229,256]" name="Indonesia" pageId="3" pageNumber="42">Indonesia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F05F9FFBFFFF10FC7FE84C92D1728" box="[246,1036,263,290]" pageId="3" pageNumber="42" type="etymology">
<paragraph id="8BAA5672FFBFFFF10FC7FE84C92D1728" blockId="3.[151,1436,195,290]" box="[246,1036,263,290]" pageId="3" pageNumber="42">
<emphasis id="B9618A60FFBFFFF10FC7FE84CC5F172B" bold="true" box="[246,382,263,289]" pageId="3" pageNumber="42">Etymology</emphasis>
. The specific epithet is a homonym of the
<typeStatus id="54AEE8D0FFBFFFF10C43FE8BCE851728" box="[882,932,264,290]" pageId="3" pageNumber="42">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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