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<document id="8801B6610ACBC170703EE9712208BE5E" ID-DOI="10.5852/ejt.2024.957.2651" ID-ISSN="2118-9773" ID-Zenodo-Dep="13773385" ID-ZooBank="9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726582870537" checkinUser="plazi" docAuthor="Selnekovič, Dávid, Jäch, Manfred A. &amp; Kodada, Ján" docDate="2024" docId="038B5001DA7AE739FDD1FA7FFDE72584" docLanguage="en" docName="EJT.2024.957.1-229.pdf" docOrigin="European Journal of Taxonomy 957" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Elmomorphus bispinosus Selnekovič, Jäch &amp; Kodada, 2024, sp. nov." docType="treatment" docUuid="B9F8E28F-FD2D-4D67-99E4-580CD3200EBA" docUuidSource="ZooBank" docVersion="2" lastPageNumber="119" masterDocId="FFB22879DA08E74FFF9EFFBAFFE32440" masterDocTitle="Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)" masterLastPageNumber="229" masterPageNumber="1" pageNumber="115" updateTime="1726584008522" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1185A6D6F4A362CF88B45EA9C384796E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D0366E7B84283890B8E800521C727F96">
<mods:title id="BF1BCBF1893982DE7578974D94503CC4">Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)</mods:title>
</mods:titleInfo>
<mods:name id="E53D2447B72111158B0AC7716BCE5E5C" type="personal">
<mods:role id="BFE84100CFBC65B1742FBBF2A2617ECB">
<mods:roleTerm id="CCDFC62472E53C154F2F8109A49DF4F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="20C87404D60B2C158043314AFDC1C087">Selnekovič, Dávid</mods:namePart>
<mods:nameIdentifier id="F1102B23254E9A97D2219335495A25B8" type="LSID">D930D757-D929-4248-A29C-A857E0505345</mods:nameIdentifier>
<mods:affiliation id="D7D1370C60A9FEDA7A14F5F08758A6E5">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="1F8B7B18811524321FF8564B41E86F50" type="email">david.selnekovic@uniba.sk</mods:nameIdentifier>
</mods:name>
<mods:name id="93B3BA6692BC785AEDCD1768A9EBEDF4" type="personal">
<mods:role id="2A645A7D3A8FA694B5CDE4B3EBF85402">
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<mods:namePart id="1AD30E71C3D3B3C9B4D4C1164C03981F">Jäch, Manfred A.</mods:namePart>
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<mods:affiliation id="EB32E9E2851DF84F5BF1B3FDABD32064">Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria.</mods:affiliation>
<mods:nameIdentifier id="2A6AD2D427BFCB39E8B5B57D35955D78" type="email">manfred.jaech@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:namePart id="6C5ECF043060561ABF00AEB98F15D61E">Kodada, Ján</mods:namePart>
<mods:nameIdentifier id="44040DD4A66D07BE15D3217ECD336B0E" type="LSID">6E88BFBB-8769-44EC-8285-29E357CEC064</mods:nameIdentifier>
<mods:affiliation id="398DC287CE4A43DCEEDC302071D492AE">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="977B5917832F21E9D07CF6CA61D9D924" type="email">jan.kodada@uniba.sk</mods:nameIdentifier>
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<mods:title id="6CE50668011CECE979716F9FF2F53704">European Journal of Taxonomy</mods:title>
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<mods:part id="C2F8EAF31A2DC0AEFD003A0049997DDA">
<mods:date id="BC6776354E1BD6524419B56BDDDAF9F5">2024</mods:date>
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<mods:number id="17BBA148235F8408215187ABA0A5145F">2024-09-11</mods:number>
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<mods:number id="44A80A18114989504DD61A0716F830B2">957</mods:number>
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<treatment id="038B5001DA7AE739FDD1FA7FFDE72584" ID-DOI="http://doi.org/10.5281/zenodo.13773545" ID-Zenodo-Dep="13773545" LSID="urn:lsid:zoobank.org:act:B9F8E28F-FD2D-4D67-99E4-580CD3200EBA" httpUri="http://treatment.plazi.org/id/038B5001DA7AE739FDD1FA7FFDE72584" lastPageId="118" lastPageNumber="119" pageId="114" pageNumber="115">
<subSubSection id="C338B29CDA7AE73DFDD1FA7FFB282243" pageId="114" pageNumber="115" type="nomenclature">
<paragraph id="8B9DE117DA7AE73DFDD1FA7FFC0621A0" blockId="114.[363,1227,1477,1575]" box="[591,997,1477,1504]" pageId="114" pageNumber="115">
<heading id="D0D5567BDA7AE73DFDD1FA7FFC0621A0" box="[591,997,1477,1504]" centered="true" fontSize="11" level="2" pageId="114" pageNumber="115" reason="2">
<taxonomicName id="4C229A94DA7AE73DFDD1FA7FFC6121A0" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[591,898,1477,1504]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="bispinosus" status="sp. nov.">
<emphasis id="B9563D05DA7AE73DFDD1FA7FFC6121A0" bold="true" box="[591,898,1477,1504]" italics="true" pageId="114" pageNumber="115">Elmomorphus bispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7AE73DFC14FA7CFC0621A0" box="[906,997,1478,1504]" pageId="114" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B9DE117DA7AE73DFEF5FA52FB282243" blockId="114.[363,1227,1477,1575]" box="[363,1227,1512,1539]" pageId="114" pageNumber="115">
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urn:lsid:zoobank.org:act:
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<subSubSection id="C338B29CDA7AE73DFD39F9B6FC6C2266" box="[679,911,1548,1575]" pageId="114" pageNumber="115" type="description">
<paragraph id="8B9DE117DA7AE73DFD39F9B6FC6C2266" blockId="114.[363,1227,1477,1575]" box="[679,911,1548,1575]" pageId="114" pageNumber="115">
<figureCitation id="1319FD92DA7AE73DFD39F9B6FCE22267" box="[679,769,1548,1575]" captionStart="Fig" captionStartId="115.[189,232,1957,1983]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@115.[189,1399,265,1928]" captionTargetPageId="115" captionText="Fig. 54. Elmomorphus bispinosus sp. nov., holotype, female (IAECAS), TL: 3.54 mm." figureDoi="http://doi.org/10.5281/zenodo.13773534" httpUri="https://zenodo.org/record/13773534/files/figure.png" pageId="114" pageNumber="115">Figs 54</figureCitation>
,
<figureCitation id="1319FD92DA7AE73DFC93F9B6FCA12267" box="[781,834,1548,1575]" captionStart="Fig" captionStartId="116.[189,232,1891,1917]" captionTargetBox="[235,1362,269,1864]" captionTargetId="figure-56@116.[213,1376,265,1864]" captionTargetPageId="116" captionText="Fig. 55. Bursa copulatrix and vagina. A. Elmomorphus parvulus sp. nov., paratype from type locality (CKB). B. E. yunnanensis Kodada, Selnekovič &amp; Jäch sp. nov., paratype from type locality (CKB). C. E. longitarsis sp. nov., paratype from type locality (NMW). D. E. bispinosus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773540" httpUri="https://zenodo.org/record/13773540/files/figure.png" pageId="114" pageNumber="115">55D</figureCitation>
,
<figureCitation id="1319FD92DA7AE73DFCD0F9B6FC6C2266" box="[846,911,1548,1574]" captionStart="Fig" captionStartId="220.[189,232,1871,1897]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-44@220.[189,1399,265,1830]" captionTargetPageId="220" captionText="Fig. 112. Distribution maps. A. Elmomorphus bispinosus sp. nov. B. E. nepalensis Satô, 1981. C. E. punctulatus sp. nov. D. E. simplipes sp. nov. E. E. donatus Kodada, Selnekovič &amp; Jäch sp. nov. F. E. paradonatus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13773718" httpUri="https://zenodo.org/record/13773718/files/figure.png" pageId="114" pageNumber="115">112A</figureCitation>
</paragraph>
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<subSubSection id="C338B29CDA7AE73AFF23F9F7FC512528" lastPageId="117" lastPageNumber="118" pageId="114" pageNumber="115" type="diagnosis">
<paragraph id="8B9DE117DA7AE73DFF23F9F7FE2A2227" blockId="114.[189,457,1613,1639]" box="[189,457,1613,1639]" pageId="114" pageNumber="115">
<heading id="D0D5567BDA7AE73DFF23F9F7FE2A2227" bold="true" box="[189,457,1613,1639]" fontSize="11" level="3" pageId="114" pageNumber="115" reason="3">
<emphasis id="B9563D05DA7AE73DFF23F9F7FE2A2227" bold="true" box="[189,457,1613,1639]" pageId="114" pageNumber="115">Differential diagnosis</emphasis>
</heading>
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<taxonomicName id="4C229A94DA7AE73DFF23F9C6FE0522D6" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[189,486,1660,1686]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="bispinosus" status="sp. nov.">
<emphasis id="B9563D05DA7AE73DFF23F9C6FE0522D6" box="[189,486,1660,1686]" italics="true" pageId="114" pageNumber="115">Elmomorphus bispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7AE73DFE71F9C6FDA922D6" box="[495,586,1660,1686]" pageId="114" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1319FD92DA7AE73DFDC4F9C6FD5222D6" box="[602,689,1659,1686]" captionStart="Fig" captionStartId="115.[189,232,1957,1983]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@115.[189,1399,265,1928]" captionTargetPageId="115" captionText="Fig. 54. Elmomorphus bispinosus sp. nov., holotype, female (IAECAS), TL: 3.54 mm." figureDoi="http://doi.org/10.5281/zenodo.13773534" httpUri="https://zenodo.org/record/13773534/files/figure.png" pageId="114" pageNumber="115">Fig. 54</figureCitation>
) is characterised by the presence of a plastron on the dorsal cranial surface (except on the anteriorly narrowed area on the vertex) and on the lateral sides of the pronotum and elytra. Elytral punctures are scattered, and striae are absent. Similar characters are also present in
<taxonomicName id="4C229A94DA7AE73DFEACF95CFE7D22BF" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[306,414,1766,1791]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="sausai" status="sp. nov.">
<emphasis id="B9563D05DA7AE73DFEACF95CFE7D22BF" box="[306,414,1766,1791]" italics="true" pageId="114" pageNumber="115">E. sausai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7AE73DFE3CF95CFE1D2340" box="[418,510,1766,1792]" pageId="114" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C229A94DA7AE73DFD96F95CFD2022BF" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[520,707,1765,1791]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="umphangicus" status="sp. nov.">
<emphasis id="B9563D05DA7AE73DFD96F95CFD2022BF" box="[520,707,1765,1791]" italics="true" pageId="114" pageNumber="115">E. umphangicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7AE73DFD57F95CFCC72340" box="[713,804,1766,1792]" pageId="114" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C229A94DA7AE73DFCB1F95CFC0322BF" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[815,992,1766,1791]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="yunnanensis" status="sp. nov.">
<emphasis id="B9563D05DA7AE73DFCB1F95CFC0322BF" box="[815,992,1766,1791]" italics="true" pageId="114" pageNumber="115">E. yunnanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7AE73DFC7AF95CFBA32340" box="[996,1088,1766,1792]" pageId="114" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C229A94DA7AE73DFBE4F95CFAFA22BF" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[1146,1305,1765,1791]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="longitarsis" status="sp. nov.">
<emphasis id="B9563D05DA7AE73DFBE4F95CFAFA22BF" box="[1146,1305,1765,1791]" italics="true" pageId="114" pageNumber="115">E. longitarsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7AE73DFA82F95CFA962340" box="[1308,1397,1766,1792]" pageId="114" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="4C229A94DA7AE73DFF25F8B2FE502362" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[187,435,1800,1826]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="sausai">
<emphasis id="B9563D05DA7AE73DFF25F8B2FE502362" box="[187,435,1800,1826]" italics="true" pageId="114" pageNumber="115">Elmomorphus sausai</emphasis>
</taxonomicName>
differs in the distinctly wider elytral plastron bands, each band occupying ca two thirds of elytron width, while in
<taxonomicName id="4C229A94DA7AE73DFDDEF896FD3D2306" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[576,734,1836,1862]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B9563D05DA7AE73DFDDEF896FD3D2306" box="[576,734,1836,1862]" italics="true" pageId="114" pageNumber="115">E. bispinosus</emphasis>
</taxonomicName>
it is ca one-third.
<taxonomicName id="4C229A94DA7AE73DFC27F896FAE12306" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[953,1282,1836,1862]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="umphangicus">
<emphasis id="B9563D05DA7AE73DFC27F896FAE12306" box="[953,1282,1836,1862]" italics="true" pageId="114" pageNumber="115">Elmomorphus umphangicus</emphasis>
</taxonomicName>
differs in smaller body dimensions (TL in
<taxonomicName id="4C229A94DA7AE73DFDD0F8EAFCED2329" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[590,782,1871,1897]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="umphangicus">
<emphasis id="B9563D05DA7AE73DFDD0F8EAFCED2329" box="[590,782,1871,1897]" italics="true" pageId="114" pageNumber="115">E. umphangicus</emphasis>
</taxonomicName>
:
<specimenCount id="9D242A9EDA7AE73DFCBCF8F5FCAD2329" box="[802,846,1871,1897]" count="1" pageId="114" pageNumber="115" type="male">♂♂</specimenCount>
<quantity id="4CDA4CF2DA7AE73DFCC7F8F5FBE42329" box="[857,1031,1871,1897]" metricMagnitude="-3" metricUnit="m" metricValue="2.77" metricValueMax="2.9" metricValueMin="2.64" pageId="114" pageNumber="115" unit="mm" value="2.77" valueMax="2.9" valueMin="2.64">2.64–2.90 mm</quantity>
(2.76 ±0.10, n= 5),
<specimenCount id="9D242A9EDA7AE73DFB63F8F5FACA2329" box="[1277,1321,1871,1897]" count="1" pageId="114" pageNumber="115" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA7AE73DFAAAF8F5FECB23CD" metricMagnitude="-3" metricUnit="m" metricValue="2.79" metricValueMax="3.12" metricValueMin="2.46" pageId="114" pageNumber="115" unit="mm" value="2.79" valueMax="3.12" valueMin="2.46">2.46– 3.12 mm</quantity>
(2.83 ±0.30, n=6), in
<taxonomicName id="4C229A94DA7AE73DFDA4F8C9FD3A23CC" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[570,729,1906,1932]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B9563D05DA7AE73DFDA4F8C9FD3A23CC" box="[570,729,1906,1932]" italics="true" pageId="114" pageNumber="115">E. bispinosus</emphasis>
</taxonomicName>
:
<specimenCount id="9D242A9EDA7AE73DFD74F8C9FCF523CD" box="[746,790,1907,1933]" count="1" pageId="114" pageNumber="115" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA7AE73DFCBEF8C8FC2E23CD" box="[800,973,1906,1933]" metricMagnitude="-3" metricUnit="m" metricValue="3.46" metricValueMax="3.54" metricValueMin="3.38" pageId="114" pageNumber="115" unit="mm" value="3.46" valueMax="3.54" valueMin="3.38">3.38–3.54 mm</quantity>
(n =3)) and the elytra being widest behind midlength.
<taxonomicName id="4C229A94DA7AE73DFE05F82FFD2023EF" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[411,707,1941,1967]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B9563D05DA7AE73DFE05F82FFD2023EF" box="[411,707,1941,1967]" italics="true" pageId="114" pageNumber="115">Elmomorphus bispinosus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C229A94DA7AE73DFCFEF82CFBF123EF" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[864,1042,1942,1967]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B9563D05DA7AE73DFCFEF82CFBF123EF" box="[864,1042,1942,1967]" italics="true" pageId="114" pageNumber="115">E. yunnanensis</emphasis>
</taxonomicName>
in its less convex body and in the presence of two microsclerites on the bursa copulatrix versus several microsclerites in
<taxonomicName id="4C229A94DA7AE73DFB59F803FA952392" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[1223,1398,1977,2002]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B9563D05DA7AE73DFB59F803FA952392" box="[1223,1398,1977,2002]" italics="true" pageId="114" pageNumber="115">E. yunnanensis</emphasis>
</taxonomicName>
(
<figureCitation id="1319FD92DA7DE73AFF58FEB3FED12564" box="[198,306,265,292]" captionStart="Fig" captionStartId="116.[189,232,1891,1917]" captionTargetBox="[235,1362,269,1864]" captionTargetId="figure-56@116.[213,1376,265,1864]" captionTargetPageId="116" captionText="Fig. 55. Bursa copulatrix and vagina. A. Elmomorphus parvulus sp. nov., paratype from type locality (CKB). B. E. yunnanensis Kodada, Selnekovič &amp; Jäch sp. nov., paratype from type locality (CKB). C. E. longitarsis sp. nov., paratype from type locality (NMW). D. E. bispinosus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773540" httpUri="https://zenodo.org/record/13773540/files/figure.png" pageId="117" pageNumber="118">Fig. 55D</figureCitation>
). From
<taxonomicName id="4C229A94DA7DE73AFE12FEB0FDC92563" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[396,554,265,291]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="longitarsis">
<emphasis id="B9563D05DA7DE73AFE12FEB0FDC92563" box="[396,554,265,291]" italics="true" pageId="117" pageNumber="118">E. longitarsis</emphasis>
</taxonomicName>
it differs in the larger (TL in
<taxonomicName id="4C229A94DA7DE73AFCE5FEB0FBF42563" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[891,1047,265,291]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B9563D05DA7DE73AFCE5FEB0FBF42563" box="[891,1047,265,291]" italics="true" pageId="117" pageNumber="118">E. bispinosus</emphasis>
</taxonomicName>
:
<specimenCount id="9D242A9EDA7DE73AFBB8FEB0FBB12564" box="[1062,1106,266,292]" count="1" pageId="117" pageNumber="118" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA7DE73AFBC7FEB3FAE02564" box="[1113,1283,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="3.46" metricValueMax="3.54" metricValueMin="3.38" pageId="117" pageNumber="118" unit="mm" value="3.46" valueMax="3.54" valueMin="3.38">3.38–3.54 mm</quantity>
(n=3), in
<taxonomicName id="4C229A94DA7DE73AFF23FE96FEB92505" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[189,346,299,325]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="longitarsis">
<emphasis id="B9563D05DA7DE73AFF23FE96FEB92505" box="[189,346,299,325]" italics="true" pageId="117" pageNumber="118">E. longitarsis</emphasis>
</taxonomicName>
:
<specimenCount id="9D242A9EDA7DE73AFEF7FE96FE762506" box="[361,405,300,326]" count="1" pageId="117" pageNumber="118" type="male">♂♂</specimenCount>
<quantity id="4CDA4CF2DA7DE73AFE02FE96FDA52506" box="[412,582,300,326]" metricMagnitude="-3" metricUnit="m" metricValue="3.01" metricValueMax="3.09" metricValueMin="2.93" pageId="117" pageNumber="118" unit="mm" value="3.01" valueMax="3.09" valueMin="2.93">2.93–3.09 mm</quantity>
(n =3),
<specimenCount id="9D242A9EDA7DE73AFD3CFE96FD2D2506" box="[674,718,300,326]" count="1" pageId="117" pageNumber="118" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA7DE73AFD4BFE91FC9C2506" box="[725,895,299,326]" metricMagnitude="-3" metricUnit="m" metricValue="3.235" metricValueMax="3.32" metricValueMin="3.15" pageId="117" pageNumber="118" unit="mm" value="3.235" valueMax="3.32" valueMin="3.15">3.15–3.32 mm</quantity>
(n= 2)), broader, and less convex body and the presence of two spiny microsclerites on the bursa copulatrix.
</paragraph>
</subSubSection>
<caption id="DF5DB19FDA7BE73CFF23F81FFB572380" ID-DOI="http://doi.org/10.5281/zenodo.13773534" ID-Zenodo-Dep="13773534" box="[189,1204,1957,1984]" httpUri="https://zenodo.org/record/13773534/files/figure.png" pageId="115" pageNumber="116" startId="115.[189,232,1957,1983]" targetBox="[189,1399,265,1928]" targetPageId="115">
<paragraph id="8B9DE117DA7BE73CFF23F81FFB572380" blockId="115.[189,1204,1957,1984]" box="[189,1204,1957,1984]" pageId="115" pageNumber="116">
<emphasis id="B9563D05DA7BE73CFF23F81FFEF923FF" bold="true" box="[189,282,1957,1983]" pageId="115" pageNumber="116">Fig. 54.</emphasis>
<taxonomicName id="4C229A94DA7BE73CFEBEF81FFDAB23FF" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[288,584,1957,1983]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="bispinosus" status="sp. nov.">
<emphasis id="B9563D05DA7BE73CFEBEF81FFDAB23FF" box="[288,584,1957,1983]" italics="true" pageId="115" pageNumber="116">Elmomorphus bispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7BE73CFDCEF81CFD4E2380" box="[592,685,1958,1984]" pageId="115" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, female (IAECAS), TL: 3.54 mm.
</paragraph>
</caption>
<caption id="DF5DB19FDA7CE73BFF23F8D9FDC523A8" ID-DOI="http://doi.org/10.5281/zenodo.13773540" ID-Zenodo-Dep="13773540" httpUri="https://zenodo.org/record/13773540/files/figure.png" pageId="116" pageNumber="117" startId="116.[189,232,1891,1917]" targetBox="[235,1362,269,1864]" targetPageId="116">
<paragraph id="8B9DE117DA7CE73BFF23F8D9FDC523A8" blockId="116.[189,1399,1891,2024]" pageId="116" pageNumber="117">
<emphasis id="B9563D05DA7CE73BFF23F8D9FEF8233E" bold="true" box="[189,283,1891,1918]" pageId="116" pageNumber="117">Fig. 55.</emphasis>
Bursa copulatrix and vagina.
<emphasis id="B9563D05DA7CE73BFD1CF8D9FD74233D" bold="true" box="[642,663,1891,1917]" pageId="116" pageNumber="117">A</emphasis>
.
<taxonomicName id="4C229A94DA7CE73BFD36F8DEFC5E233E" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[680,957,1892,1918]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov.">
<emphasis id="B9563D05DA7CE73BFD36F8DEFC5E233E" box="[680,957,1892,1918]" italics="true" pageId="116" pageNumber="117">Elmomorphus parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7CE73BFC58F8DEFBC6233E" box="[966,1061,1892,1918]" pageId="116" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
, paratype from type locality (CKB).
<emphasis id="B9563D05DA7CE73BFE81F83DFED023E1" bold="true" box="[287,307,1927,1953]" pageId="116" pageNumber="117">B</emphasis>
.
<taxonomicName id="4C229A94DA7CE73BFEDAF832FCB023E1" authority="Kodada, Selnekovic &amp; Jach" authorityName="Kodada, Selnekovic &amp; Jach" authorityYear="2024" box="[324,851,1927,1953]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="yunnanensis" status="sp. nov.">
<emphasis id="B9563D05DA7CE73BFEDAF832FE1A23E1" box="[324,505,1928,1953]" italics="true" pageId="116" pageNumber="117">E. yunnanensis</emphasis>
Kodada, Selnekovič &amp; Jäch
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7CE73BFCC3F83DFC5D23E1" box="[861,958,1927,1953]" pageId="116" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
, paratype from type locality (CKB).
<emphasis id="B9563D05DA7CE73BFF23F810FF312384" bold="true" box="[189,210,1962,1988]" pageId="116" pageNumber="117">C</emphasis>
.
<taxonomicName id="4C229A94DA7CE73BFF7BF811FE6B2384" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[229,392,1962,1988]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="longitarsis" status="sp. nov.">
<emphasis id="B9563D05DA7CE73BFF7BF811FE6B2384" box="[229,392,1962,1988]" italics="true" pageId="116" pageNumber="117">E. longitarsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7CE73BFE0CF811FE172385" box="[402,500,1963,1989]" pageId="116" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
, paratype from type locality (NMW).
<emphasis id="B9563D05DA7CE73BFC55F810FC032384" bold="true" box="[971,992,1962,1988]" pageId="116" pageNumber="117">D</emphasis>
.
<taxonomicName id="4C229A94DA7CE73BFC6DF811FB702384" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[1011,1171,1962,1988]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="bispinosus" status="sp. nov.">
<emphasis id="B9563D05DA7CE73BFC6DF811FB702384" box="[1011,1171,1962,1988]" italics="true" pageId="116" pageNumber="117">E. bispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA7CE73BFB01F811FAE32385" box="[1183,1280,1963,1989]" pageId="116" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (IAECAS). Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C338B29CDA7DE73AFF23FE35FD9D25B9" pageId="117" pageNumber="118" type="etymology">
<paragraph id="8B9DE117DA7DE73AFF23FE35FEA725E9" blockId="117.[189,324,399,425]" box="[189,324,399,425]" pageId="117" pageNumber="118">
<heading id="D0D5567BDA7DE73AFF23FE35FEA725E9" bold="true" box="[189,324,399,425]" fontSize="11" level="3" pageId="117" pageNumber="118" reason="3">
<emphasis id="B9563D05DA7DE73AFF23FE35FEA725E9" bold="true" box="[189,324,399,425]" pageId="117" pageNumber="118">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF23FE07FD9D25B9" blockId="117.[189,1399,445,506]" pageId="117" pageNumber="118">The epithet is a Latin adjective in the nominative singular, referring to the presence of two spiny microsclerites on the bursa copulatrix.</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA7DE73AFF23FD9AFDAF27A7" pageId="117" pageNumber="118" type="materials_examined">
<paragraph id="8B9DE117DA7DE73AFF23FD9AFE8F267A" blockId="117.[189,364,544,570]" box="[189,364,544,570]" pageId="117" pageNumber="118">
<heading id="D0D5567BDA7DE73AFF23FD9AFE8F267A" bold="true" box="[189,364,544,570]" fontSize="11" level="3" pageId="117" pageNumber="118" reason="3">
<emphasis id="B9563D05DA7DE73AFF23FD9AFE8F267A" bold="true" box="[189,364,544,570]" pageId="117" pageNumber="118">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF74FDF4FEBF2628" blockId="117.[189,1399,590,686]" box="[234,348,590,616]" pageId="117" pageNumber="118">
<emphasis id="B9563D05DA7DE73AFF74FDF4FEBF2628" bold="true" box="[234,348,590,616]" pageId="117" pageNumber="118">
<typeStatus id="54995FB5DA7DE73AFF74FDF4FEBF2628" box="[234,348,590,616]" pageId="117" pageNumber="118" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF23FDCAFD3026ED" blockId="117.[189,1399,590,686]" pageId="117" pageNumber="118">
<materialsCitation id="3B4AEB4ADA7DE73AFF23FDCAFD3026ED" collectionCode="IAECAS" collectorName="Schonmann" country="China" elevation="400" location="Hunan Province" pageId="117" pageNumber="118" specimenCount="1" specimenCount-female="1" stateProvince="Hunan" typeStatus="holotype">
<collectingCountry id="F335A187DA7DE73AFF23FDCAFEFB26CA" box="[189,280,624,650]" name="China" pageId="117" pageNumber="118">CHINA</collectingCountry>
–
<emphasis id="B9563D05DA7DE73AFEACFDCAFE1F26CA" bold="true" box="[306,508,624,650]" pageId="117" pageNumber="118">
<collectingRegion id="49E62FF5DA7DE73AFEACFDCAFE1F26CA" box="[306,508,624,650]" country="China" name="Hunan" pageId="117" pageNumber="118">Hunan Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D242A9EDA7DE73AFD8AFDCBFDC826CB" box="[532,555,625,651]" count="1" pageId="117" pageNumber="118" type="female">♀</specimenCount>
; “
<collectingCountry id="F335A187DA7DE73AFDDDFDCAFD4426CA" box="[579,679,624,650]" name="China" pageId="117" pageNumber="118">CHINA</collectingCountry>
: SW
<collectingRegion id="49E62FF5DA7DE73AFD7CFDCBFCD326CB" box="[738,816,625,651]" country="China" name="Hunan" pageId="117" pageNumber="118">Hunan</collectingRegion>
1993 SW Huitong, 7. 11. Umg. Guangping,
<quantity id="4CDA4CF2DA7DE73AFAAAFDCAFA9426CA" box="[1332,1399,624,650]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="117" pageNumber="118" unit="m" value="400.0">
<elevation id="000F0624DA7DE73AFAAAFDCAFA9426CA" box="[1332,1399,624,650]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="117" pageNumber="118" unit="m" value="400.0">400m</elevation>
</quantity>
leg.
<collectorName id="26D784C1DA7DE73AFF71FD29FE9926ED" box="[239,378,659,685]" pageId="117" pageNumber="118">Schönmann</collectorName>
(14) [CWBS 35]”;
<collectionCode id="ED3379D2DA7DE73AFDFFFD29FD2C26ED" box="[609,719,659,685]" pageId="117" pageNumber="118">IAECAS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF74FD6EFE8426AE" blockId="117.[189,1398,724,819]" box="[234,359,724,750]" pageId="117" pageNumber="118">
<emphasis id="B9563D05DA7DE73AFF74FD6EFE8426AE" bold="true" box="[234,359,724,750]" pageId="117" pageNumber="118">
<typeStatus id="54995FB5DA7DE73AFF74FD6EFE8426AE" box="[234,359,724,750]" pageId="117" pageNumber="118" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF23FD4CFEEE2773" blockId="117.[189,1398,724,819]" pageId="117" pageNumber="118">
<materialsCitation id="3B4AEB4ADA7DE73AFF23FD4CFEEA2773" collectionCode="NMW" collectorName="Schonmann" country="China" elevation="400" location="Hunan Province" pageId="117" pageNumber="118" specimenCount="3" specimenCount-female="2" stateProvince="Hunan" typeStatus="paratype">
<collectingCountry id="F335A187DA7DE73AFF23FD4CFEFB2750" box="[189,280,758,784]" name="China" pageId="117" pageNumber="118">CHINA</collectingCountry>
–
<emphasis id="B9563D05DA7DE73AFEAAFD4CFE1C2750" bold="true" box="[308,511,758,784]" pageId="117" pageNumber="118">
<collectingRegion id="49E62FF5DA7DE73AFEAAFD4CFE1C2750" box="[308,511,758,784]" country="China" name="Hunan" pageId="117" pageNumber="118">Hunan Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D242A9EDA7DE73AFD86FD4CFDB92751" box="[536,602,758,785]" count="2" pageId="117" pageNumber="118" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype, but
<specimenCount id="9D242A9EDA7DE73AFBD8FD4DFB042750" box="[1094,1255,758,785]" count="1" pageId="117" pageNumber="118" type="generic">one specimen</specimenCount>
“leg. L. JI”;
<collectionCode id="ED3379D2DA7DE73AFF23FCA3FEEA2773" box="[189,265,793,819]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="117" pageNumber="118">NMW</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF23FCE3FEBE2733" blockId="117.[189,349,857,883]" box="[189,349,857,883]" pageId="117" pageNumber="118">
<heading id="D0D5567BDA7DE73AFF23FCE3FEBE2733" bold="true" box="[189,349,857,883]" fontSize="11" level="3" pageId="117" pageNumber="118" reason="3">
<emphasis id="B9563D05DA7DE73AFF23FCE3FEBE2733" bold="true" box="[189,349,857,883]" pageId="117" pageNumber="118">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF23FC3DFDAF27A7" blockId="117.[189,1399,903,999]" pageId="117" pageNumber="118">
<collectingCountry id="F335A187DA7DE73AFF23FC3DFEE527E1" box="[189,262,903,929]" name="China" pageId="117" pageNumber="118">China</collectingCountry>
,
<collectingRegion id="49E62FF5DA7DE73AFE86FC32FE3C27E1" box="[280,479,903,930]" country="China" name="Hunan" pageId="117" pageNumber="118">Hunan Province</collectingRegion>
, Huaihua Prefecture, Huitong county, Guangping Township, near Moshao village, small stream, 0.5–1.0 m wide, partly canalised, partly vanishing beneath the gravel, ca
<quantity id="4CDA4CF2DA7DE73AFAB5FC10FA942784" box="[1323,1399,938,964]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="117" pageNumber="118" unit="m" value="400.0">400 m</quantity>
a.s.l. (
<collectionCode id="ED3379D2DA7DE73AFE9DFC76FEBB27A6" box="[259,344,972,998]" pageId="117" pageNumber="118">CWBS</collectionCode>
35;
<bibRefCitation id="EFB39CE6DA7DE73AFE12FC76FDDC27A7" author="Jach M. A. &amp; Ji L." box="[396,575,972,999]" pageId="117" pageNumber="118" pagination="5 - 32" refId="ref99754" refString="Jach M. A. &amp; Ji L. 1995. Introduction. In: Jach M. A. &amp; Ji L. (eds) Water Beetles of China. Vol. I: 5 - 32. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Vienna." type="book chapter" year="1995">Jäch &amp; Ji 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA7DE739FF23FBB7FE952512" lastPageId="118" lastPageNumber="119" pageId="117" pageNumber="118" type="description">
<paragraph id="8B9DE117DA7DE73AFF23FBB7FEAD2067" blockId="117.[189,334,1037,1063]" box="[189,334,1037,1063]" pageId="117" pageNumber="118">
<heading id="D0D5567BDA7DE73AFF23FBB7FEAD2067" bold="true" box="[189,334,1037,1063]" fontSize="11" level="3" pageId="117" pageNumber="118" reason="3">
<emphasis id="B9563D05DA7DE73AFF23FBB7FEAD2067" bold="true" box="[189,334,1037,1063]" pageId="117" pageNumber="118">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF23FB81FCD62037" blockId="117.[189,1399,1082,1144]" pageId="117" pageNumber="118">
Measurements (mm): TL:
<specimenCount id="9D242A9EDA7DE73AFD9DFB81FDCC2015" box="[515,559,1083,1109]" count="1" pageId="117" pageNumber="118" type="female">♀♀</specimenCount>
3.38–3.54 (n =3); PL:
<specimenCount id="9D242A9EDA7DE73AFCCAFB81FC632015" box="[852,896,1083,1109]" count="1" pageId="117" pageNumber="118" type="female">♀♀</specimenCount>
0.75 (n =3); PW:
<specimenCount id="9D242A9EDA7DE73AFBF4FB81FB752015" box="[1130,1174,1083,1109]" count="1" pageId="117" pageNumber="118" type="female">♀♀</specimenCount>
1.44–1.48 (n =3); EL:
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2.67–2.86 (n= 3); EW:
<specimenCount id="9D242A9EDA7DE73AFDABFBE4FD822038" box="[565,609,1118,1144]" count="1" pageId="117" pageNumber="118" type="female">♀♀</specimenCount>
1.72–1.82 (n =3).
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF23FB24FAFB2160" blockId="117.[189,1399,1182,1312]" pageId="117" pageNumber="118">
Body oval, moderately convex dorsally (
<figureCitation id="1319FD92DA7DE73AFD34FB24FCE620F9" box="[682,773,1182,1209]" captionStart="Fig" captionStartId="115.[189,232,1957,1983]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@115.[189,1399,265,1928]" captionTargetPageId="115" captionText="Fig. 54. Elmomorphus bispinosus sp. nov., holotype, female (IAECAS), TL: 3.54 mm." figureDoi="http://doi.org/10.5281/zenodo.13773534" httpUri="https://zenodo.org/record/13773534/files/figure.png" pageId="117" pageNumber="118">Fig. 54</figureCitation>
). Integument black; mouthparts, antennae, and tarsi reddish brown. Pubescence consisting of short yellowish setae. Plastron present on cranial surface except anteriorly narrowed area on vertex, on lateral sides of pronotum and elytra, and on ventral surface except for prosternal process, median part of metaventrite and median part of ventrites 1 and 2.
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF23FAFCFEEF21AB" blockId="117.[189,1399,1350,1515]" pageId="117" pageNumber="118">
Head with round punctures separated by 0.5–1.0 × puncture diameter. Plastron present on entire cranial surface except on anteriorly narrowed area on vertex. Labrum transverse, anterior edge straight, exposed portion microreticulate; setae concentrated along anterior margin. Anterior margin of clypeus straight. Eyes oval, moderately protruding, ID:
<specimenCount id="9D242A9EDA7DE73AFD05FA14FD242188" box="[667,711,1454,1480]" count="1" pageId="117" pageNumber="118" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA7DE73AFD4DFA14FC612188" box="[723,898,1454,1481]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="117" pageNumber="118" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.50 mm</quantity>
(n =3). Antennae 10-segmented, densely setose.
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF23F9A8FD642322" blockId="117.[189,1399,1553,1890]" pageId="117" pageNumber="118">
Pronotum transverse, widest at base, moderately convex, PW/PL:
<specimenCount id="9D242A9EDA7DE73AFC5CF9A8FC0D226C" box="[962,1006,1554,1580]" count="1" pageId="117" pageNumber="118" type="female">♀♀</specimenCount>
1.92–1.97 (n =3); anterior angles protruding, acute; lateral sides convergent, rounded; surface smooth, with round setiferous punctures; plastron bands present along entire lateral sides. Lateral and posterior edges of prosternal process rounded; lateral portions raised; median keel arcuate. Lateral sides of metaventral process weakly raised. Disc of metaventrite weakly convex, without clusters of long setae. Elytra oval, moderately convex, widest around midlength, EL/EW:
<specimenCount id="9D242A9EDA7DE73AFD5FF904FD0E2298" box="[705,749,1726,1752]" count="1" pageId="117" pageNumber="118" type="female">♀♀</specimenCount>
1.55–1.61 (n =3); surface weakly microreticulate with irregular meshes, with weak bronze lustre; round setiferous punctures scattered; lateral plastron bands widened posteriorly, covering about one-third of elytron width at elytral midlength. Tibiae straight, protibia ca 1.2 × as long as protarsus, PrTL/PL:
<specimenCount id="9D242A9EDA7DE73AFC9AF89CFCD32300" box="[772,816,1830,1856]" count="1" pageId="117" pageNumber="118" type="female">♀♀</specimenCount>
1.16–1.26 (n =3). Terminal protarsomere longer than all preceding segments combined.
</paragraph>
<paragraph id="8B9DE117DA7DE73AFF23F832FEA323A9" blockId="117.[189,1399,1928,2025]" pageId="117" pageNumber="118">
Ventrites covered with plastron, except on middle of ventrites 1 and 2. Ventrite 5 with short longitudinal apical keel. Bursa copulatrix on each side with a rather large microsclerite bearing one prominent spine (
<figureCitation id="1319FD92DA7DE73AFF58F874FED023A9" box="[198,307,1998,2025]" captionStart="Fig" captionStartId="116.[189,232,1891,1917]" captionTargetBox="[235,1362,269,1864]" captionTargetId="figure-56@116.[213,1376,265,1864]" captionTargetPageId="116" captionText="Fig. 55. Bursa copulatrix and vagina. A. Elmomorphus parvulus sp. nov., paratype from type locality (CKB). B. E. yunnanensis Kodada, Selnekovič &amp; Jäch sp. nov., paratype from type locality (CKB). C. E. longitarsis sp. nov., paratype from type locality (NMW). D. E. bispinosus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773540" httpUri="https://zenodo.org/record/13773540/files/figure.png" pageId="117" pageNumber="118">Fig. 55D</figureCitation>
).
</paragraph>
<paragraph id="8B9DE117DA7EE739FF23FEB2FDD42562" blockId="118.[189,567,264,291]" box="[189,567,264,291]" pageId="118" pageNumber="119">
<heading id="D0D5567BDA7EE739FF23FEB2FDD42562" bold="true" box="[189,567,264,291]" fontSize="11" level="3" pageId="118" pageNumber="119" reason="3">
<emphasis id="B9563D05DA7EE739FF23FEB2FDD42562" bold="true" box="[189,567,264,291]" pageId="118" pageNumber="119">Secondary sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA7EE739FF23FE82FE952512" blockId="118.[189,374,312,338]" box="[189,374,312,338]" pageId="118" pageNumber="119">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA7EE739FF23FEC0FDE72584" box="[189,516,378,452]" pageId="118" pageNumber="119" type="distribution">
<paragraph id="8B9DE117DA7EE739FF23FEC0FEB525D4" blockId="118.[189,342,378,404]" box="[189,342,378,404]" pageId="118" pageNumber="119">
<heading id="D0D5567BDA7EE739FF23FEC0FEB525D4" bold="true" box="[189,342,378,404]" fontSize="11" level="3" pageId="118" pageNumber="119" reason="3">
<emphasis id="B9563D05DA7EE739FF23FEC0FEB525D4" bold="true" box="[189,342,378,404]" pageId="118" pageNumber="119">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA7EE739FF23FE13FDE72584" blockId="118.[189,516,425,452]" box="[189,516,425,452]" pageId="118" pageNumber="119">
<collectingCountry id="F335A187DA7EE739FF23FE13FEE02583" box="[189,259,425,451]" name="China" pageId="118" pageNumber="119">China</collectingCountry>
(
<collectingRegion id="49E62FF5DA7EE739FE8CFE10FE872584" box="[274,356,426,452]" country="China" name="Hunan" pageId="118" pageNumber="119">Hunan</collectingRegion>
) (
<figureCitation id="1319FD92DA7EE739FEE2FE13FE1B2584" box="[380,504,425,452]" captionStart="Fig" captionStartId="220.[189,232,1871,1897]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-44@220.[189,1399,265,1830]" captionTargetPageId="220" captionText="Fig. 112. Distribution maps. A. Elmomorphus bispinosus sp. nov. B. E. nepalensis Satô, 1981. C. E. punctulatus sp. nov. D. E. simplipes sp. nov. E. E. donatus Kodada, Selnekovič &amp; Jäch sp. nov. F. E. paradonatus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13773718" httpUri="https://zenodo.org/record/13773718/files/figure.png" pageId="118" pageNumber="119">Fig. 112A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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