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<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2022" docId="CAF1EE5E3DF75B39AD0355F922D83BAA" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Rhaphaulus perakensis Smith 1898" docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:title id="D62D851EA06E80D954109B0C4A4D9C49">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:namePart id="A5ECB17D4A43DAEF4410B4DB0FF6EADA">Jirapatrasilp, Parin</mods:namePart>
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<mods:namePart id="C4BDA947094AA6872C7EC208B853748D">Sutcharit, Chirasak</mods:namePart>
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<treatment id="CAF1EE5E3DF75B39AD0355F922D83BAA" ID-GBIF-Taxon="199839053" LSID="urn:lsid:plazi:treatment:CAF1EE5E3DF75B39AD0355F922D83BAA" httpUri="http://treatment.plazi.org/id/CAF1EE5E3DF75B39AD0355F922D83BAA" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupinellinae" scope_superFamily="Cyclophoroidea">
<subSubSection id="296DAAE7F722658F84B57F75146B9297" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="1230E5CA01FE4182B592FE64E15BA69A" pageId="0" pageNumber="1">
<taxonomicName id="228BE2C91AEF32260E485F85DF7A5C61" ID-CoL="4S3QM" LSID="CAF1EE5E-3DF7-5B39-AD03-55F922D83BAA" authority="Smith, 1898" authorityName="Smith" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">Rhaphaulus perakensis Smith, 1898</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="50CBD58012FAE3767D98EA43DEAC04AB" pageId="0" pageNumber="1" type="description">
<paragraph id="EC82DEBF012E3C09455723AFA0854C6D" pageId="0" pageNumber="1">
<figureCitation id="E048EFD09FE2C15D61FED46F7767EBA0" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A, B Rhaphaulus lorraini: A syntype NHMUK 20130454 from Pulo Penang and B specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga C Rhaphaulus perakensis, syntype NHMUK 1897.3.15.41 from Larut, Perak D Rhaphaulus ascendens, syntype UMZC I. 100025 from Patalung, Malay Peninsula E, F Rhaphaulus tonkinensis E holotype HNHM 98757 and F specimen CUMZ 12163 / 1 from Luang Cave, Chiang Rai. Photo: B. Pall-Gergely (E)" figureDoi="10.3897/zookeys.1119.85400.figure13" httpUri="https://binary.pensoft.net/fig/735374" pageId="0" pageNumber="1">Fig. 13C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5566A4B3A8D475482943C5DEE9E844E0" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="1FD0809FC6AC87A60E92DC98AB188DC9" pageId="0" pageNumber="1">
<taxonomicName id="34422ECC79BC67BBE5D04A82437CDAEA" authorityName="Smith" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">Rhaphaulus perakensis</taxonomicName>
Smith, 1898: 17, figs 1, 2. Type locality:
<normalizedToken id="D0717BF3170D6558FF2C0D9DFA358C73" originalValue="Maxwells">Maxwell's</normalizedToken>
Hill, Larut [Bukit Larut], Perak.
<bibRefCitation id="BC44D6D4629BB8C513C26FB393E3FF33" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
: 276, 277.
<bibRefCitation id="C9CC43B955583D96846E83DF35B1B586" author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
: 32, 33.
<bibRefCitation id="EDE2A6725C140B18D2CE925AE5D44294" author="van Benthem Jutting, WSS" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="1" pagination="50 - 77" refId="B180" refString="van Benthem Jutting, WSS, 1949. On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells. Bulletin of the Raffles Museum 19: 50 - 77" title="On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells." volume="19" year="1949">van Benthem Jutting 1949</bibRefCitation>
: 57, Kuala Kenering;
<normalizedToken id="4D5AE512D624835FAF801B5B0C23ED26" originalValue="Maxwells">Maxwell's</normalizedToken>
Hill, Perak; Dusun Tua, Selangor [Malaysia].
<bibRefCitation id="4D880018DBEA7767FFA05AA22A301A23" author="Habe, T" journalOrPublisher="Nature and Life in Southeast Asia" pageId="0" pageNumber="1" pagination="111 - 127" refId="B63" refString="Habe, T, 1965. Operculated land molluscs from Southeast Asia. Nature and Life in Southeast Asia 4: 111 - 127" title="Operculated land molluscs from Southeast Asia." volume="4" year="1965">Habe 1965</bibRefCitation>
: 115, 116, as a synonym of
<taxonomicName id="B8AD8EDD2CC554E45642C04B89B40001" baseAuthorityName="Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus chrysalis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysalis">Rhaphaulus chrysalis</taxonomicName>
.
<bibRefCitation id="33BDF741CFB3981F497406A8000823A4" author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie" pageId="0" pageNumber="1" pagination="35 - 70" refId="B66" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
: 40, Thailand.
<bibRefCitation id="94CDECB859D1BEED0425123EC4E47084" author="Pall-Gergely, B" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="565 - 573" refId="B111" refString="Pall-Gergely, B, Hunyadi, A, Maassen, WJM, 2014. Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae). Journal of Conchology 41: 565 - 573" title="Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae)." volume="41" year="2014">
<normalizedToken id="74C94955CF3087999D754E555B4CFB02" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2014
</bibRefCitation>
: 572, fig. 12, western Malaysia.
<bibRefCitation id="E6E649F5EBC69588D737D3169A722B39" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
: 97.
</paragraph>
<paragraph id="4EDB6E674B4CA81C30709D95C05CBDD5" pageId="0" pageNumber="1">
<taxonomicName id="94B93D4C61FDEEFFF6F96DD15FE9C324" authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus perakensis subsp. var. var. jalorensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="perakensis" subSpecies="var." variety="jalorensis">Rhaphaulus perakensis var. jalorensis</taxonomicName>
Sykes, 1903: 197, pl. 20, figs 9, 10. Type locality: Bukit Bisar, on the borders of Jalor [Khao Yai National Reserved Forest, Namtok Sai Khao National Park, Mueang Yala District, Yala Province, Thailand].
</paragraph>
<paragraph id="96986503BE1F7E6F2450BBC5B91B66E5" pageId="0" pageNumber="1">
<taxonomicName id="69B3D17F7BDC70C45343DD53B6181E65" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus perakensis subsp. var. var. ialorensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="perakensis" subSpecies="var." variety="ialorensis">Rhaphaulus perakensis var. ialorensis</taxonomicName>
[sic]-
<bibRefCitation id="A27CCEE761679D7878AA9A29E2236439" author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
: 33.
</paragraph>
<paragraph id="613B3B0C79392E4AA2A29E8989CCB6BF" pageId="0" pageNumber="1">
<taxonomicName id="17BF7F882901426A360EFE7499EBF205" authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus perakensis subsp. jalorensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perakensis" subSpecies="jalorensis">Rhaphaulus perakensis jalorensis</taxonomicName>
-
<bibRefCitation id="F69481FA8EDFAE48DB6F9CA189123A31" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
: 42.
</paragraph>
<paragraph id="7D7202E18F54456429D9E9D6D023577A" pageId="0" pageNumber="1">
<taxonomicName id="B43AD24F035D371DD9BDDA846BFD7FAB" authority="perakensis" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus perakensis subsp. perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perakensis" subSpecies="perakensis">Rhaphaulus perakensis perakensis</taxonomicName>
-
<bibRefCitation id="70373999A9FA3F3258A88E367D89D192" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
: 42.
</paragraph>
<paragraph id="E2D610DE32F16D26CA1D1E4710FCC917" pageId="0" pageNumber="1">
<taxonomicName id="26E72E5D44FE8CB20E608F3CC10B83CD" authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus jalorensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jalorensis">Rhaphaulus jalorensis</taxonomicName>
-
<bibRefCitation id="CE0CADD3F41A566CDB9BE26C656791D9" author="Pall-Gergely, B" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="565 - 573" refId="B111" refString="Pall-Gergely, B, Hunyadi, A, Maassen, WJM, 2014. Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae). Journal of Conchology 41: 565 - 573" title="Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae)." volume="41" year="2014">
<normalizedToken id="ABECC1A7FE61C29F09710F21280752C1" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2014
</bibRefCitation>
: 572, western Malaysia.
<bibRefCitation id="47D54747D91CEF15969852CC1E157E4F" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
: 96.
<bibRefCitation id="3D30844CA4E82D60F43491696E07B1B8" author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
: 157, fig. 5-13l.
</paragraph>
</subSubSection>
<subSubSection id="56B4BA3FC5EAD455FCAEA8DCA6AF800F" pageId="0" pageNumber="1" type="type material">
<paragraph id="C25191D8B6C8E57705D6D889B4C5578F" pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph id="87B96CDE0921D72F8E272455BAC462DE" pageId="0" pageNumber="1">
<emphasis id="91FFE66121283C17429F1FB71B3C3BDA" bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
of
<taxonomicName id="47FCBB9D15BA519A33E3CA574B25DCA4" authorityName="Smith" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">
<emphasis id="D7EF2CB0A13CC72C924F21D2E1B9B2BF" italics="true" pageId="0" pageNumber="1">Rhaphaulus perakensis</emphasis>
</taxonomicName>
NHMUK 1897.3.15.41-2 (2 shells; Fig.
<figureCitation id="A61EBC34A3E99DDA5B6310CB618E15E7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A, B Rhaphaulus lorraini: A syntype NHMUK 20130454 from Pulo Penang and B specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga C Rhaphaulus perakensis, syntype NHMUK 1897.3.15.41 from Larut, Perak D Rhaphaulus ascendens, syntype UMZC I. 100025 from Patalung, Malay Peninsula E, F Rhaphaulus tonkinensis E holotype HNHM 98757 and F specimen CUMZ 12163 / 1 from Luang Cave, Chiang Rai. Photo: B. Pall-Gergely (E)" figureDoi="10.3897/zookeys.1119.85400.figure13" httpUri="https://binary.pensoft.net/fig/735374" pageId="0" pageNumber="1">13C</figureCitation>
) from Larut, Perak.
</paragraph>
</subSubSection>
<subSubSection id="32ABF7C9E6A2B860761CAD1DD5F370D5" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="78FC060157CC696FE78F1F3E4D000EC5" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="3533DEEF805DE4FC0703D35E09FCED95" pageId="0" pageNumber="1">Shell elongate ovate; body whorls slightly bulging. Tube cylindrical, pointing diagonally downward and backward.</paragraph>
</subSubSection>
<subSubSection id="F3B9A78939C85CB98EA8C59C77271F20" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="88715CE6CA323CBCE63CEB52A2BDA28F" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="645376E2C96FDDA01D16AA1ACB4B046A" pageId="0" pageNumber="1">
<taxonomicName id="4D005B4F15998C6AA4465B5F2AE062C1" authorityName="Smith" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">
<emphasis id="65C222D326A5891D797A1656C5BF3071" italics="true" pageId="0" pageNumber="1">Rhaphaulus perakensis</emphasis>
</taxonomicName>
can be distinguished from all other species from mainland Southeast Asia by a cylindrical tube pointing diagonally downward and backward.
</paragraph>
</subSubSection>
<subSubSection id="A82C4A04C8F97F2BBA4BB6C30BC25E27" pageId="0" pageNumber="1" type="distribution">
<paragraph id="FE4826D43C31F8800B4F26757F802F01" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="5FD2A7EDDBEF90B2CDD556F304AF670B" pageId="0" pageNumber="1">
Northern Peninsular Malaysia and southern Thailand (
<bibRefCitation id="741D72BB459A8F4CADE5764200D43D82" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
;
<bibRefCitation id="129A1945F07E7320512651C8C0F368B0" author="Pall-Gergely, B" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="565 - 573" refId="B111" refString="Pall-Gergely, B, Hunyadi, A, Maassen, WJM, 2014. Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae). Journal of Conchology 41: 565 - 573" title="Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae)." volume="41" year="2014">
<normalizedToken id="1B7013A436A549704EA73757CDD5F080" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F3C09A460BC345F80B1D83D6641DC89A" pageId="0" pageNumber="1" type="remarks">
<paragraph id="579C721819AC4C8FCDF7B8AEE2A1969B" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="A81663DB7F9DB4A78375F79617FCD5CF" pageId="0" pageNumber="1">
No material of this species was found during this survey.
<bibRefCitation id="221BDE85E9BA992F12CD1CC9910B9A6B" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen (2001)</bibRefCitation>
treated
<taxonomicName id="65CBB39E6347D10F47825E997B749415" lsidName="R. perakensis subsp. jalorensis" pageId="0" pageNumber="1" rank="subSpecies" species="perakensis" subSpecies="jalorensis">
<emphasis id="FE20E5E3B02DF2C559A6B966E30CA066" italics="true" pageId="0" pageNumber="1">R. perakensis jalorensis</emphasis>
</taxonomicName>
as a junior subjective synonym of
<taxonomicName id="652109A82010CB9F18DF146EA378B453" lsidName="R. p. subsp. perakensis" pageId="0" pageNumber="1" rank="subSpecies" species="p." subSpecies="perakensis">
<emphasis id="FCBFA6E8083450C5E053DEC4865D5D75" italics="true" pageId="0" pageNumber="1">R. p. perakensis</emphasis>
</taxonomicName>
without apparent reason, whereas
<bibRefCitation id="292BA8D4FB6B4E90209AFC20CA58DCC5" author="Pall-Gergely, B" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="565 - 573" refId="B111" refString="Pall-Gergely, B, Hunyadi, A, Maassen, WJM, 2014. Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae). Journal of Conchology 41: 565 - 573" title="Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae)." volume="41" year="2014">
<normalizedToken id="56C6C55EBAD422C5ED5CE38C0B5CE30D" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. (2014)
</bibRefCitation>
listed this subspecies as a valid species following the opinion of
<bibRefCitation id="6A4051287D3F0BB04B3E6CA016AB3B50" author="Sykes, ER" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="194 - 199" refId="B160" refString="Sykes, ER, 1903. On the land operculate mollusks collected during the &quot;Skeat Expedition&quot; to the Malay peninsula in 1899-1900. Proceedings of the Zoological Society of London 1903: 194 - 199" title="On the land operculate mollusks collected during the &quot; Skeat Expedition &quot; to the Malay peninsula in 1899 - 1900." volume="1903" year="1903">Sykes (1903)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>