245 lines
24 KiB
XML
245 lines
24 KiB
XML
<document ID-CLB-Dataset="209209" ID-DOI="http://dx.doi.org/10.3897/asp.80.e90570" ID-GBIF-Dataset="bac17de6-baf9-4fee-9ded-268adbb26a61" ID-Pensoft-Pub="1864-8312-80-649" ID-Pensoft-UUID="FAE6DFEF69735531A4CBA275E078320D" ID-ZooBank="AD0B1CC262414460B6B888E6B95F20D7" ModsDocID="1864-8312-80-649" checkinTime="1671114540108" checkinUser="pensoft" docAuthor="Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S. & Harms, Danilo" docDate="2022" docId="8415D05F4BBF5412AE4BF1FE8C7DC4CB" docLanguage="en" docName="ArthropodSystPhyl 80: 649-691" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-12-14" docSource="http://dx.doi.org/10.3897/asp.80.e90570" docTitle="Toliaranella griswoldi Lorenz & Loria & Harvey & Harms 2022, sp. nov." docType="treatment" docVersion="5" id="FAE6DFEF69735531A4CBA275E078320D" lastPageNumber="649" masterDocId="FAE6DFEF69735531A4CBA275E078320D" masterDocTitle="The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots" masterLastPageNumber="691" masterPageNumber="649" pageNumber="649" updateTime="1732733992417" updateUser="ExternalLinkService">
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<mods:title id="86855A3DC9537E94FEBDDEF64B4147AF">The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots</mods:title>
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<mods:namePart id="5E76D80BF382C90336B76D3616F03593">Lorenz, Michelle</mods:namePart>
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<mods:affiliation id="CB4CB8FD72FFC9615A28F21B7C9648BB">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany & University of Hamburg, Biology Department, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier id="ABB8737F0B86B8ACC81B5B3063CECFF1" type="email">michellelorenz@gmx.net</mods:nameIdentifier>
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<mods:namePart id="B9BBE87C4170FFF30A183C336CDE19E3">Loria, Stephanie F.</mods:namePart>
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<mods:affiliation id="A1A94EA5CA8D9D4DC37085BE9CFD4EB3">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:roleTerm id="A11796769014A9EF961D30286E49C00D">Author</mods:roleTerm>
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<mods:namePart id="14E641EDCC5965D8011F10234661A5A5">Harvey, Mark S.</mods:namePart>
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<mods:nameIdentifier id="B977293E85E969CD101BEB7C4F3DD456" type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier>
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<mods:affiliation id="DBC4E12D76596904395F9F676195F022">Western Australian Museum, Collections & Research, 49 Kew Street, Welshpool, WA 6106, Australia & University of Western Australia, School of Biological Sciences, Crawley WA 6009, Australia</mods:affiliation>
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<mods:namePart id="DE16D43A2EA72D2326AB2149F806A2B6">Harms, Danilo</mods:namePart>
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<mods:nameIdentifier id="30520F712DC37F7E11C3D1F4545B3ADC" type="ORCID">https://orcid.org/0000-0002-7189-5345</mods:nameIdentifier>
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<mods:affiliation id="C2E6497329D6CC0B6070889E9FEE39C8">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:title id="C92A3374B78541C89F60914B7F339C16">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:date id="60A9EB29A1FB1040F1E5CA130F2157D1">2022</mods:date>
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<mods:number id="D35078F7F7C1C7C3FD8B5C752240504F">2022-12-14</mods:number>
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<mods:identifier id="A0D8B0F7E3753B7E3F9D5EA34F789F2B" type="DOI">http://dx.doi.org/10.3897/asp.80.e90570</mods:identifier>
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<treatment id="8415D05F4BBF5412AE4BF1FE8C7DC4CB" ID-GBIF-Taxon="206273800" LSID="urn:lsid:zoobank.org:act:NomEnClAturAlACts/08C129BA-3575-4C44-94B4-64C8CF38E3FC" httpUri="http://treatment.plazi.org/id/8415D05F4BBF5412AE4BF1FE8C7DC4CB" lastPageNumber="649" pageId="0" pageNumber="649" scope_family="Feaellidae" scope_order="Pseudoscorpiones" scope_subFamily="Feaellinae" scope_superFamily="Feaelloidea">
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<subSubSection id="6509741A48CB32A5C0AFD92806AFF631" pageId="0" pageNumber="649" type="nomenclature">
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<paragraph id="37D65E367A5400F6BFF37CA691B48F5A" pageId="0" pageNumber="649">
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<taxonomicName id="862E7E0AE5A8BCD3FABB65BAF6B44C25" LSID="https://zoobank.org/NomenclaturalActs/08C129BA-3575-4C44-94B4-64C8CF38E3FC" authority="Lorenz & Loria & Harvey & Harms, 2022" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella griswoldi" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="griswoldi" status="sp. nov.">Toliaranella griswoldi</taxonomicName>
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<taxonomicNameLabel id="FBE7F991F383EC9426E08D7B4454BB37" pageId="0" pageNumber="649">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="B592D08010428464E89504C71F346AE3" pageId="0" pageNumber="649" type="type material">
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<paragraph id="A67C1DB892B497AF7F070801B4DA32FE" pageId="0" pageNumber="649">Type material examined.</paragraph>
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<paragraph id="61059EFCB6E56D5FD76ACBFF3CD98639" pageId="0" pageNumber="649">
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MADAGASCAR: Toliara Province [Anosy Region]: Holotype ♂ (CAS 9071500 [BLF4810]), allotype ♀ (ZMH-A0016739), paratypes: 11 ♂, 1 ♀ (CAS 9071500 [BLF4810]), 4 ♂, 2 ♀ (ZMH-A0016740 - ZMH-A0016745), Parc National
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<normalizedToken id="2B548266308541817248A2418DC24523" originalValue="d’Andohahela">d'Andohahela</normalizedToken>
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,
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<normalizedToken id="CDAE00DC3F7B0E6F594A120708DBCB08" originalValue="Forêt">Foret</normalizedToken>
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de Manantalinjo,
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<geoCoordinate id="01807443B2C1E7A992007EC1BCA277A4" degrees="24" direction="south" minutes="49" orientation="latitude" precision="15" seconds="1" value="-24.816946">24°49′1″S</geoCoordinate>
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<geoCoordinate id="FA8C43BC54FEF44B187B46626ABE32BC" degrees="46" direction="east" minutes="36" orientation="longitude" precision="15" seconds="36" value="46.609997">46°36′36″E</geoCoordinate>
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, 12-16 January 2002, B.L. Fisher, Griswold et al.
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</paragraph>
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</subSubSection>
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<subSubSection id="00F15A8A3CAC2C8E899C2AE5BFFABA1D" pageId="0" pageNumber="649" type="materials_examined">
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<paragraph id="7623B9E9220BE9EF9A5A9804984E9599" pageId="0" pageNumber="649">Additional material examined.</paragraph>
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<paragraph id="0E0CDC2FF88C7C16876187836F06586F" pageId="0" pageNumber="649">
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<materialsCitation id="45464758992532C23A71601E0E38E8E7" ID-GBIF-Occurrence="3990971326" collectingDate="2002-01-16" collectingDateMax="2002-01-20" collectingDateMin="2002-01-16" collectorName="Parc National d'Andohahela, Foret d'Ambohibory" country="MADAGASCAR" latitude="-24.93" location="Anosy Region" longLatPrecision="21" longitude="46.645557" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Toliara Province">
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<collectingCountry id="DCCB12608615B9AE1CEC33CCF73E53BE" name="Madagascar">MADAGASCAR</collectingCountry>
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:
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<collectingRegion id="F8E0AF90977E355DDA865AE71BA6D290" country="Madagascar" name="Toliara">Toliara Province</collectingRegion>
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[
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<location id="D5C6C485E298BF1AA00AA2B8E3C5EFC7" LSID="urn:lsid:plazi:treatment:8415D05F4BBF5412AE4BF1FE8C7DC4CB:D5C6C485E298BF1AA00AA2B8E3C5EFC7" country="MADAGASCAR" latitude="-24.93" longLatPrecision="21" longitude="46.645557" name="Anosy Region" stateProvince="Toliara Province">Anosy Region</location>
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]:
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<specimenCount id="03DF5C4F743F944D1C067E4AB50EB3A4" type="male">4 ♂</specimenCount>
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,
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<specimenCount id="67D197B6AFE0D4665A4A736C8B3C0124" type="female">4 ♀</specimenCount>
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, 9 nymphs (CAS 9071509 [BLF4984]),
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<collectorName id="52083DE1CC03DA03F8CA8CB1C9E8A3BA">
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Parc National
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<normalizedToken id="9FAE2C1C27D7B95717EEB0029142BAA9" originalValue="d’Andohahela">d'Andohahela</normalizedToken>
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</collectorName>
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,
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<collectorName id="0E020AFE3144197E12456CA4B31A5B07">
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<normalizedToken id="C74AB66BC307729AB440495D8EEB426E" originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken id="F544DD1BB55E2D531495D1D18F124918" originalValue="d’Ambohibory">d'Ambohibory</normalizedToken>
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</collectorName>
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,
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<geoCoordinate id="5485D57F990D269D58D154D2A47D36D2" degrees="24" direction="south" minutes="55" orientation="latitude" precision="15" seconds="48" value="-24.93">24°55′48″S</geoCoordinate>
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<geoCoordinate id="6BB30FCAD2570C73A685B80ABDFC348B" degrees="46" direction="east" minutes="38" orientation="longitude" precision="15" seconds="44" value="46.645557">46°38′44″E</geoCoordinate>
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,
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<collectingDate id="377781995D3C4B3584C32BB76A5372FF" value="2002-01-16" valueMax="2002-01-20" valueMin="2002-01-16">16-20 January 2002</collectingDate>
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, Fisher, Griswold et al
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</materialsCitation>
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.;
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<materialsCitation id="ABE18BA77F84B807F78C504623B5914D" ID-GBIF-Occurrence="3990971305" collectingDate="2002-01-12" collectingDateMax="2002-01-16" collectingDateMin="2002-01-12" collectorName="Parc National d'Andohahela, de Manantalinjo" country="Madagascar" latitude="-24.816946" location="Foret de Manantalinjo" longLatPrecision="21" longitude="46.609997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Toliara Province">
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<specimenCount id="AFC0AF81FC415BD23D251BCDD339D78D" type="male">1 ♂</specimenCount>
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,
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<specimenCount id="D5EA1638AB9A8281ADAE905F128AC957" type="female">1 ♀</specimenCount>
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(CAS 9071516 [BLF4815]),
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<collectorName id="9003B51D3A4FE5E239F369A49AF9F95A">
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Parc National
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<normalizedToken id="D0D7ECAECCEAABD111339355A33F4C69" originalValue="d’Andohahela">d'Andohahela</normalizedToken>
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</collectorName>
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,
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<location id="AF91E31B50A6A5E9106F1EA9EDB3C73B" LSID="urn:lsid:plazi:treatment:8415D05F4BBF5412AE4BF1FE8C7DC4CB:AF91E31B50A6A5E9106F1EA9EDB3C73B" country="Madagascar" latitude="-24.816946" longLatPrecision="21" longitude="46.609997" name="Foret de Manantalinjo" stateProvince="Toliara Province">
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Foret
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<collectorName id="DC2FBFBBBEE019A2735290CC20C1C90A">de Manantalinjo</collectorName>
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</location>
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,
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<geoCoordinate id="D4D242C4AA155DFBB6AD84662AC0133A" degrees="24" direction="south" minutes="49" orientation="latitude" precision="15" seconds="1" value="-24.816946">24°49′1″S</geoCoordinate>
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<geoCoordinate id="2E30EEF19279C1E525D0F49120180A91" degrees="46" direction="east" minutes="36" orientation="longitude" precision="15" seconds="36" value="46.609997">46°36′36″E</geoCoordinate>
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,
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<collectingDate id="FBE595B13F665CBACBC930CE02AE8E79" value="2002-01-12" valueMax="2002-01-16" valueMin="2002-01-12">12-16 January 2002</collectingDate>
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, Fisher, Griswold et al
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C4084BF017275037A21D43F18BAC6E27" pageId="0" pageNumber="649" type="diagnosis">
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<paragraph id="E2157EF4403CF8716407BC4A78DD6253" pageId="0" pageNumber="649">Diagnosis.</paragraph>
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<paragraph id="7EC59011417C802DB90128BD50DDDBE6" pageId="0" pageNumber="649">
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Differs from
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<taxonomicName id="D13B90CFB8C16F89A990D732878BF177" lsidName="T. pumila" pageId="0" pageNumber="649" rank="species" species="pumila">
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<emphasis id="1F37E6B1957C420568AAC60EFAC7EBCB" italics="true" pageId="0" pageNumber="649">T. pumila</emphasis>
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</taxonomicName>
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<emphasis id="A455B482D0FD0099786CF9AE8FD3EB6C" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
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in different cheotaxy:
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<emphasis id="E9DE97182E985F4FDA8F74452A0C5E1E" italics="true" pageId="0" pageNumber="649">est</emphasis>
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on fixed finger situated between
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<emphasis id="77C7A70A9B6E440C6E9D30F7AB726F46" italics="true" pageId="0" pageNumber="649">sb</emphasis>
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and
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<emphasis id="BAF0956B0D2902D094F1F16EF6F27ABE" italics="true" pageId="0" pageNumber="649">t</emphasis>
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of the movable finger on sagittal level (
|
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<emphasis id="0CD1AB344A9D08D8FD03C5B0FE8480A9" italics="true" pageId="0" pageNumber="649">est</emphasis>
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on same sagittal level with
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<emphasis id="5E66F899FE3F475788627790F6453899" italics="true" pageId="0" pageNumber="649">sb</emphasis>
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in
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<taxonomicName id="230F5B5F2531E0B33730A36732364D03" lsidName="T. pumila" pageId="0" pageNumber="649" rank="species" species="pumila">
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<emphasis id="2687AF69195A952A669A8F94647BD1F2" italics="true" pageId="0" pageNumber="649">T. pumila</emphasis>
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</taxonomicName>
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<emphasis id="E868005A5738D92EDC94EC08B81DC14A" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
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); most compact carapace of the southern feaellid members (1.27/1.23 ♂/♀ times longer than broad versus 1.31-1.37/1.31-1.39 ♂/♀).
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</paragraph>
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</subSubSection>
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<subSubSection id="87905D037482BD59D3902AC23DF51D3E" pageId="0" pageNumber="649" type="etymology">
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<paragraph id="561C346ED50875827FE7E9EF70C108B9" pageId="0" pageNumber="649">Etymology.</paragraph>
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<paragraph id="0CD60DE63E1044769AFDCBD0787D9E5A" pageId="0" pageNumber="649">This species is a patronym honoring Charles Griswold who collected many Malagasy feaellids.</paragraph>
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</subSubSection>
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<subSubSection id="25E4804BC1F73113A876EA8FB2D6F062" pageId="0" pageNumber="649" type="description">
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<paragraph id="A7DA523E4B7DC4E9C63FE38CE2689ED6" pageId="0" pageNumber="649">Description.</paragraph>
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<paragraph id="6B3ACA0C3278B7B3D8CD982659B51A0A" pageId="0" pageNumber="649">
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The following description is based on holotype and allotype. -
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<emphasis id="C63C42CC678FC73E85355E75BE2ACB59" italics="true" pageId="0" pageNumber="649">Carapace</emphasis>
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(Fig.
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<figureCitation id="504498532BB571E743E75CDB9A7BF84B" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Drawings of Toliaranella griswoldi sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure22" httpUri="https://binary.pensoft.net/fig/782014" pageId="0" pageNumber="649">22A</figureCitation>
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): Very compact, heavily granulate, 1.20-1.31 (♂), 1.18-1.31 (♀) times longer than broad. -
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<emphasis id="9B50C003577747DA98359379BB10D674" italics="true" pageId="0" pageNumber="649">Pedipalps</emphasis>
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(Fig.
|
||
<figureCitation id="5745F09DED630D9E21F3A6D73A70F0BA" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Drawings of Toliaranella griswoldi sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure22" httpUri="https://binary.pensoft.net/fig/782014" pageId="0" pageNumber="649">22C, D</figureCitation>
|
||
): Similar trichobothrial pattern as
|
||
<taxonomicName id="1C22B6228BBEE511F265B12E64DC9A79" lsidName="T. pumila" pageId="0" pageNumber="649" rank="species" species="pumila">
|
||
<emphasis id="4FF40F821217F8ABCFD7D75436BFB0E1" italics="true" pageId="0" pageNumber="649">T. pumila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="39640A7F8F28AFAB372611A4AC5281F9" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
||
but the following difference:
|
||
<emphasis id="80E0AEF98BA15191FE605263CB78DC23" italics="true" pageId="0" pageNumber="649">est</emphasis>
|
||
on fixed finger situated between
|
||
<emphasis id="54827073CD3DC73F7AEF0CB41FEC9566" italics="true" pageId="0" pageNumber="649">sb</emphasis>
|
||
and
|
||
<emphasis id="EB844E26ED41D0276E6BE69EBD24AE71" italics="true" pageId="0" pageNumber="649">t</emphasis>
|
||
of the movable finger on sagittal level. Chelal fixed finger with 9-10 chelal teeth in the OR, 12-14 in the MR and 9-11 in the IR; movable finger with 6-8 in the OR, 7-11 in the MR and 7-8 in the IR. Fixed finger with 6-7 terminal teeth equal in size and one large one, movable finger with 7-8 equally sized terminal teeth. -
|
||
<taxonomicName id="FD2183946132CE408B52790A7C1D7AA6" genus="Dimensions" lsidName="Dimensions" pageId="0" pageNumber="649" rank="genus">
|
||
<emphasis id="769C04A5ABCE649E2AF1186E8B65CD5D" italics="true" pageId="0" pageNumber="649">Dimensions</emphasis>
|
||
</taxonomicName>
|
||
(mm):
|
||
<emphasis id="F95CDDD71DA18F49B878C7526427E187" italics="true" pageId="0" pageNumber="649">Holotype</emphasis>
|
||
♂: Body length 1.62; abdomen length 1.13, width (with pleura) 1.05, width (without pleura) 0.91; carapace length 0.46, width 0.35. Pedipalp: trochanter 0.22; femur length 0.49, width 0.27; patella 0.35; chela (without pedicel) 0.35; hand length 0.10, width 0.11; movable finger 0.26. Leg I: trochanter 0.07; femur 0.14; patella 0.13; tibia 0.11; tarsus 0.21. Leg IV: trochanter 0.13; femur 0.10; patella 0.21; tibia 0.20; tarsus 0.26.
|
||
<emphasis id="DC9409F2E86B6E57946D40608D58489E" italics="true" pageId="0" pageNumber="649">Allotype</emphasis>
|
||
♀: Body length 2.04; abdomen length 1.47, width (with pleura) 1.33, width (without pleura) 1.11; carapace length 0.53, width 0.45. Pedipalp: trochanter 0.25; femur length 0.59, width 0.35; patella 0.46; chela (without pedicel) 0.44; hand length 0.10, width 1.14; movable finger 0.35. Leg I: trochanter 0.11; femur 0.16; patella 0.14; tibia 0.16; tarsus 0.25. Leg IV: trochanter 0.18; femur 0.14; patella 0.27; tibia 0.31; tarsus 0.30.
|
||
</paragraph>
|
||
<caption id="005ECF76DD914D8703C7943247EB4958" doi="10.3897/asp.80.e90570.figure22" httpUri="https://binary.pensoft.net/fig/782014" pageId="0" pageNumber="649" start="Figure 22" startId="F22">
|
||
<paragraph id="A04305257598DD167CDBED6EE69738E4" pageId="0" pageNumber="649">
|
||
<emphasis id="5C5F19D070695DB4EE700BE048572CE2" bold="true" pageId="0" pageNumber="649">Figure 22.</emphasis>
|
||
Drawings of
|
||
<taxonomicName id="58DBDB3C59554F69F2D93AB4275B85A9" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella griswoldi" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="griswoldi">
|
||
<emphasis id="4260A3345D85E820911392A0DFD0D448" italics="true" pageId="0" pageNumber="649">Toliaranella griswoldi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C771D8B8CD8CC0BBA271089B491F3901" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
||
:
|
||
<emphasis id="082D6AC38553732FDA85D2E18F2CFB0D" bold="true" pageId="0" pageNumber="649">A</emphasis>
|
||
carapace from dorsal;
|
||
<emphasis id="623F038C79F97A0965FE7E9485694604" bold="true" pageId="0" pageNumber="649">B</emphasis>
|
||
leg IV;
|
||
<emphasis id="90E91D06360E60BA0C4633E2CEA47896" bold="true" pageId="0" pageNumber="649">C</emphasis>
|
||
left chela from dorsal with trichobothrial pattern;
|
||
<emphasis id="BEC875FF63A57266D30F18DCC9AF1DA9" bold="true" pageId="0" pageNumber="649">D</emphasis>
|
||
right pedipalp from dorsal. Scale bars: 0.2 mm (
|
||
<emphasis id="DAA81DE968E90943148AA3F547CC36D5" bold="true" pageId="0" pageNumber="649">A</emphasis>
|
||
-
|
||
<emphasis id="5B45316795204570E8B967D47F31DBE0" bold="true" pageId="0" pageNumber="649">C</emphasis>
|
||
); 0.4 mm (
|
||
<emphasis id="28AB30DE145FC4DA0A4162D84AAA9ED1" bold="true" pageId="0" pageNumber="649">D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="BF350E6F9F76E3DA504114D93A70B682" pageId="0" pageNumber="649" type="variation">
|
||
<paragraph id="655E8D1644EB84A3B01E9DF9959F8BD4" pageId="0" pageNumber="649">Variations.</paragraph>
|
||
<paragraph id="7AE64ED0B6FF59B27B51F19C2F00E7E2" pageId="0" pageNumber="649">Body length 1.54-1.67 (♂), 1.96-2.24 (♀); abdomen length 1.08-1.18 (♂), 1.41-1.63 (♀); width (with pleura) 0.98-1.09 (♂), 1.27-1.41 (♀); width (without pleura) 0.84-0.95 (♂), 1.10-1.19 (♀); 1.20-1.29 (♂), 1.27-1.37 (♀) times longer than broad; carapace length 0.42-0.48 (♂), 0.53-0.57 (♀); width 0.35-0.38 (♂), 0.41-0.45 (♀); 1.20-1.31 (♂), 1.18-1.29 (♀) times longer than broad. Pedipalp: trochanter 0.19-0.22 (♂), 0.24-0.26 (♀); femur length 0.40-0.49 (♂), 0.54-0.59 (♀); width 0.25-0.28 (♂), 0.33-0.37 (♀); 1.60-1.81 (♂), 1.53-1.69 (♀) times longer than broad; patella 0.33-0.36 (♂), 0.45-0.48 (♀); chela (without pedicel) 0.35-0.38 (♂), 0.43-0.47 (♀); hand length 0.08-0.11 (♂), 0.10-0.11 (♀); width 0.11-0.13 (♂), 0.13-0.14 (♀); 0.73-0.91 (♂), 0.71-0.79 (♀) times longer than broad; movable finger 0.26-0.30 (♂), 0.35-0.37 (♀).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7D03FC18421335CE3692B8D2DB08ABD7" pageId="0" pageNumber="649" type="genbank code">
|
||
<paragraph id="58904304EDEDF24EF902D7C7683CAD4D" pageId="0" pageNumber="649">GenBank Code.</paragraph>
|
||
<paragraph id="087F08DD53AE12B4BB22AA0F365A0C4E" pageId="0" pageNumber="649">OP589959, OP589960. The species differs from all congeners by more than 3.7% in the H3 dataset.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4C030D0361846AD65EB9F1DEEBFF1A67" pageId="0" pageNumber="649" type="habitat">
|
||
<paragraph id="752D68699C57B3188208267655A9B45A" pageId="0" pageNumber="649">Habitat.</paragraph>
|
||
<paragraph id="EDF0C9036F0D247B61D4466C2C44C26E" pageId="0" pageNumber="649">Specimens were found in sifted litter of spiny forest thicket and tropical dry forest at an elevation of 150-300 m.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16D204F0BE19EC862735928B5CAB6D5C" pageId="0" pageNumber="649" type="distribution">
|
||
<paragraph id="8FBBA53C45778746FF339E696CC37861" pageId="0" pageNumber="649">Distribution.</paragraph>
|
||
<paragraph id="D09B547B3E4AD3E3439BF404C6271AF1" pageId="0" pageNumber="649">Currently known from the type locality (BLF4810) and two additional localities of (BLF4984 and BLF4815) in the Anosy Region (former Toliara Province) in southern Madagascar.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |