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<document id="7AF6D2E3793699602116460619864318" ID-DOI="10.11646/zootaxa.5415.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10695701" ID-ZooBank="0667714E-7612-4836-A9BA-08331FB28F43" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1708677971549" checkinUser="plazi" docAuthor="Hamer, Matthew T. &amp; Turner, Clive R." docDate="2024" docId="6E175657855849652FFD6EA4FA120AAA" docLanguage="en" docName="zootaxa.5415.4.6.pdf" docOrigin="Zootaxa 5415 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5415.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Strumigenys species" docType="key" docVersion="1" lastPageNumber="575" masterDocId="922E2E2F855C49612F6A6E33FF8F0D65" masterDocTitle="Strumigenys emmae (Emery, 1890) (Myrmicinae) new to Britain, with an updated key to the known Strumigenys of the West Palaearctic" masterLastPageNumber="576" masterPageNumber="570" pageNumber="574" updateTime="1708976891317" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="1A94034822E6E7E75236E9FDAE836C9B">Strumigenys emmae (Emery, 1890) (Myrmicinae) new to Britain, with an updated key to the known Strumigenys of the West Palaearctic</mods:title>
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<mods:namePart id="B21B56BEC21D0B012A87C4B4ECC2C325">Hamer, Matthew T.</mods:namePart>
<mods:affiliation id="8EEA2394E7BC7200CB7CAEDB8DE38195">School of Biological Sciences, The University of Hong Kong, Kadoorie Biological Sciences Building, Pok Fu Lam Road, Hong Kong SAR, China.</mods:affiliation>
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<mods:namePart id="D4D1E9CD060BBA2EBA18F17BEC9D9460">Turner, Clive R.</mods:namePart>
<mods:affiliation id="8DA3EE58E4FBB2EA849CF8C2D482D307">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD.</mods:affiliation>
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<mods:date id="536F39898A857E52A79B0B936E37079E">2024</mods:date>
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<mods:url id="4CCD3F0E17B0DCEDC63C899E220D01A8">http://dx.doi.org/10.11646/zootaxa.5415.4.6</mods:url>
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<mods:classification id="7F1AC8D8923835E72B3148F0ADBF48BA">journal article</mods:classification>
<mods:identifier id="1D897F38FA04A94D6AE6EFEA736830D3" type="DOI">10.11646/zootaxa.5415.4.6</mods:identifier>
<mods:identifier id="196D7A72E62EEF30AC373E34FB062A5C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2CD55C26EF616A28EE49214384DE3B46" type="Zenodo-Dep">10695701</mods:identifier>
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<treatment id="6E175657855849652FFD6EA4FA120AAA" LSID="urn:lsid:plazi:treatment:6E175657855849652FFD6EA4FA120AAA" httpUri="http://treatment.plazi.org/id/6E175657855849652FFD6EA4FA120AAA" lastPageNumber="575" pageId="4" pageNumber="574">
<subSubSection id="AEA4B4CA855849652FFD6EA4FB320DD4" box="[151,1213,151,177]" pageId="4" pageNumber="574" type="nomenclature">
<paragraph id="E601E741855849652FFD6EA4FB320DD4" blockId="4.[151,1213,151,177]" box="[151,1213,151,177]" pageId="4" pageNumber="574">
<heading id="BD49502D855849652FFD6EA4FB320DD4" bold="true" box="[151,1213,151,177]" fontSize="11" level="1" pageId="4" pageNumber="574" reason="1">
<emphasis id="D4CA3B53855849652FFD6EA4FB320DD4" bold="true" box="[151,1213,151,177]" pageId="4" pageNumber="574">
Taxonomic key of the native and introduced
<taxonomicName id="21BE9CC2855849652DAE6EA4FC360DD4" box="[708,953,151,177]" pageId="4" pageNumber="574">
<emphasis id="D4CA3B53855849652DAE6EA4FCD30DD4" bold="true" box="[708,860,151,177]" italics="true" pageId="4" pageNumber="574">Strumigenys</emphasis>
species
</taxonomicName>
known from Europe
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AEA4B4CA855849652FFD6EECFE0D0C24" pageId="4" pageNumber="574" type="description">
<paragraph id="E601E741855849652FFD6EECFE0D0C24" blockId="4.[151,1437,223,322]" pageId="4" pageNumber="574">
This key is adapted from Hamer
<emphasis id="D4CA3B53855849652D4C6ED3FDD20D9C" box="[550,605,223,249]" italics="true" pageId="4" pageNumber="574">et al</emphasis>
., (2021). An additional couplet (couplet 3) is provided here to enable identification of
<taxonomicName id="21BE9CC2855849652E386F30FE330C78" authorityName="Myrmicinae" baseAuthorityName="Emery" baseAuthorityYear="1890" box="[338,444,259,285]" class="Insecta" family="Formicidae" genus="Strumigenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="emmae">
<emphasis id="D4CA3B53855849652E386F30FE330C78" box="[338,444,259,285]" italics="true" pageId="4" pageNumber="574">S. emmae</emphasis>
</taxonomicName>
within the European
<taxonomicName id="21BE9CC2855849652DC76F30FCB80C78" box="[685,823,259,285]" class="Insecta" family="Formicidae" genus="Strumigenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="D4CA3B53855849652DC76F30FCB80C78" box="[685,823,259,285]" italics="true" pageId="4" pageNumber="574">Strumigenys</emphasis>
</taxonomicName>
fauna. Italicised figure citations refer to figures within Hamer
<emphasis id="D4CA3B53855849652F806F1BFE930C24" box="[234,284,295,321]" italics="true" pageId="4" pageNumber="574">et al</emphasis>
., (2021).
</paragraph>
</subSubSection>
<key id="922FB070855849642FFD6F5FFA120C38" lastPageId="5" lastPageNumber="575" pageId="4" pageNumber="574">
<subSubSection id="AEA4B4CA855849652FFD6F5FFA120AAA" pageId="4" pageNumber="574" type="key">
<keyStep id="5D4AFFE4855849652FFD6F5FFA130F54" pageId="4" pageNumber="574">
<paragraph id="E601E741855849652FFD6F5FFA130CD9" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6F5FFA130CD9" pageId="4" pageNumber="574">
<emphasis id="D4CA3B53855849652FFD6F5FFF2C0CE4" bold="true" box="[151,163,364,385]" pageId="4" pageNumber="574">1</emphasis>
Mandibles narrow, MI 24–58, linear or bowed. At full closure mandibles engage only at their extreme apices and enclose an open space between them.Apicodorsal tooth of mandible long and spiniform, strongly crossing over the tooth from the opposite mandible when closed. Inner margin of mandible with 1–4 teeth or denticles (
<emphasis id="D4CA3B53855849652CA76F94FBC60CD9" box="[973,1097,423,444]" italics="true" pageId="4" pageNumber="574">figure 6 A–D</emphasis>
)..........................
<emphasis id="D4CA3B53855849652AFA6F94FA130CD9" bold="true" box="[1424,1436,423,444]" pageId="4" pageNumber="574">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="E601E741855849652FFD6FF6FA130F54" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6FF6FA130F54" pageId="4" pageNumber="574">
- Mandibles triangular, MI 15–20. At full closure mandibles engage throughout their visible length, without an open space between them. Tooth at apex of mandible small and inconspicuous, not spiniform, not strongly crossing over the tooth from the opposite mandible when closed. Inner margin of mandible with 12–14 teeth and denticles, some of which may be minute (
<emphasis id="D4CA3B53855849652A0E6FCCFEAF0F57" italics="true" pageId="4" pageNumber="574">figure 6 E–H</emphasis>
).............................................................................................
<emphasis id="D4CA3B53855849652AFA6C2FFA130F54" bold="true" box="[1424,1436,540,561]" pageId="4" pageNumber="574">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5D4AFFE4855849652FFD6C0AFA130E79" pageId="4" pageNumber="574">
<paragraph id="E601E741855849652FFD6C0AFA120FA0" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6C0AFA120FA0" pageId="4" pageNumber="574">
<emphasis id="D4CA3B53855849652FFD6C0AFF2C0F2B" bold="true" box="[151,163,569,590]" pageId="4" pageNumber="574">2</emphasis>
Pronotal humeral hair absent. Dorsum of mesosoma without standing hairs (
<emphasis id="D4CA3B53855849652CA46C0AFBBE0F2A" box="[974,1073,569,591]" italics="true" pageId="4" pageNumber="574">figure 7 A</emphasis>
). Mandibles each with 3–4 preapical denticles. When mandibles fully closed the labral lobes are visible between them as a pair of elongate narrow triangles (
<emphasis id="D4CA3B53855849652A396C64FF7E0FEF" italics="true" pageId="4" pageNumber="574">figure 7 B</emphasis>
). (
<collectingCountry id="9EA9A7D1855849652E606C47FEDA0FEF" box="[266,341,628,650]" name="Austria" pageId="4" pageNumber="574">Austria</collectingCountry>
, Azerbaijan,
<collectingCountry id="9EA9A7D1855849652EBA6C47FDAA0FEF" box="[464,549,628,650]" name="Bulgaria" pageId="4" pageNumber="574">Bulgaria</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652D476C47FDF90FEF" box="[557,630,628,650]" name="Croatia" pageId="4" pageNumber="574">Croatia</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652D146C47FC970FEF" box="[638,792,628,650]" name="Czech Republic" pageId="4" pageNumber="574">Czech Republic</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652C4B6C46FCED0FEF" box="[801,866,629,650]" name="France" pageId="4" pageNumber="574">France</collectingCountry>
(southern mainland &amp;
<collectingCountry id="9EA9A7D1855849652B566C47FB060FEF" box="[1084,1161,628,650]" name="France" pageId="4" pageNumber="574">Corsica</collectingCountry>
), Georgia,
<collectingCountry id="9EA9A7D1855849652B9A6C47FAC50FEC" box="[1264,1354,628,649]" name="Germany" pageId="4" pageNumber="574">Germany</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652A386C47FA170FEC" box="[1362,1432,628,649]" name="Greece" pageId="4" pageNumber="574">Greece</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652F886CA1FEB80FC2" box="[226,311,658,679]" name="Hungary" pageId="4" pageNumber="574">Hungary</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652E2B6CA2FEF60FC2" box="[321,377,657,679]" name="Israel" pageId="4" pageNumber="574">Israel</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652EE86CA2FE210FC2" box="[386,430,657,679]" name="Italy" pageId="4" pageNumber="574">Italy</collectingCountry>
(mainland,
<collectingRegion id="247A29A3855849652D496CA2FDFC0FC2" box="[547,627,657,679]" country="Italy" name="Sardegna" pageId="4" pageNumber="574">Sardinia</collectingRegion>
&amp;
<collectingRegion id="247A29A3855849652DF86CA2FD420FC2" box="[658,717,657,679]" country="Italy" name="Sicilia" pageId="4" pageNumber="574">Sicily</collectingRegion>
),
<collectingCountry id="9EA9A7D1855849652DB56CA2FC960FC2" box="[735,793,657,679]" name="Malta" pageId="4" pageNumber="574">Malta</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652C496CA1FCF30FC2" box="[803,892,658,679]" name="Morocco" pageId="4" pageNumber="574">Morocco</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652CEC6CA2FC490FC2" box="[902,966,657,679]" name="Russia" pageId="4" pageNumber="574">Russia</collectingCountry>
(Caucasus),
<collectingCountry id="9EA9A7D1855849652B2E6CA2FB0B0FC2" box="[1092,1156,657,679]" name="Serbia" pageId="4" pageNumber="574">Serbia</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652BE76CA2FB490FC2" box="[1165,1222,657,679]" name="Spain" pageId="4" pageNumber="574">Spain</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652BBA6CA2FACA0FC2" box="[1232,1349,657,679]" name="Switzerland" pageId="4" pageNumber="574">Switzerland</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652A256CA2FA160FC2" box="[1359,1433,657,679]" name="Tunisia" pageId="4" pageNumber="574">Tunisia</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652F886C9CFEA70FA0" box="[226,296,687,709]" name="Turkey" pageId="4" pageNumber="574">Turkey</collectingCountry>
)............................................................................... ..
<emphasis id="D4CA3B53855849652A6F6C9DFAC30FA1" bold="true" box="[1285,1356,686,708]" italics="true" pageId="4" pageNumber="574">argiola</emphasis>
(native)
</keyLead>
</paragraph>
<paragraph id="E601E741855849652FFD6CFEFA130E79" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6CFEFA130E79" pageId="4" pageNumber="574">
- Pronotal humeral hair present (
<emphasis id="D4CA3B53855849652D7E6CFFFDF60F84" box="[532,633,716,737]" italics="true" pageId="4" pageNumber="574">figure 7 C</emphasis>
). Dorsum of mesosoma with at least one pair of standing hairs (
<emphasis id="D4CA3B53855849652B936CFFFA000F87" box="[1273,1423,716,738]" italics="true" pageId="4" pageNumber="574">figure 7 C &amp; E</emphasis>
). Mandibles each with 1–2 preapical teeth. When mandibles fully closed the labral lobes are not visible or at most appear between them as a pair of minute points basally (
<emphasis id="D4CA3B53855849652D096D34FD7C0E79" box="[611,755,775,796]" italics="true" pageId="4" pageNumber="574">figure 7 D &amp; F</emphasis>
).......................................................
<emphasis id="D4CA3B53855849652AFA6D34FA130E79" bold="true" box="[1424,1436,775,796]" pageId="4" pageNumber="574">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5D4AFFE4855849652FFD6D17FA130ECA" pageId="4" pageNumber="574">
<paragraph id="E601E741855849652FFD6D17FA120E10" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6D17FA120E10" pageId="4" pageNumber="574">
<emphasis id="D4CA3B53855849652FFD6D17FF2C0E5C" bold="true" box="[151,163,804,825]" pageId="4" pageNumber="574">3</emphasis>
Head dorsum covered in orbicular hairs (
<figureCitation id="7E85FBC4855849652D046D17FD5A0E5C" box="[622,725,804,826]" captionStart="FIGURE 3" captionStartId="3.[152,255,1277,1301]" captionTargetBox="[270,1318,181,1253]" captionTargetId="figure-31@3.[270,1318,181,1253]" captionTargetPageId="3" captionText="FIGURE 3. Strumigenys emmae worker and queen specimens.A) S. emmae worker full face view, B) S. emmae worker lateral view, C) S. emmae worker dorsal view. D) S. emmae queen full face view, E) S. emmae queen lateral view, F) S. emmae queen dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10695707" httpUri="https://zenodo.org/record/10695707/files/figure.png" pageId="4" pageNumber="574">figure 3 A</figureCitation>
); antennae with four segments (including scape) (
<figureCitation id="7E85FBC4855849652BDD6D17FA910E5C" box="[1207,1310,804,826]" captionStart="FIGURE 3" captionStartId="3.[152,255,1277,1301]" captionTargetBox="[270,1318,181,1253]" captionTargetId="figure-31@3.[270,1318,181,1253]" captionTargetPageId="3" captionText="FIGURE 3. Strumigenys emmae worker and queen specimens.A) S. emmae worker full face view, B) S. emmae worker lateral view, C) S. emmae worker dorsal view. D) S. emmae queen full face view, E) S. emmae queen lateral view, F) S. emmae queen dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10695707" httpUri="https://zenodo.org/record/10695707/files/figure.png" pageId="4" pageNumber="574">figure 3 A</figureCitation>
); scape short and expanded into convex lobe in median third of length (
<figureCitation id="7E85FBC4855849652C7B6D72FCF80E32" box="[785,887,833,855]" captionStart="FIGURE 3" captionStartId="3.[152,255,1277,1301]" captionTargetBox="[270,1318,181,1253]" captionTargetId="figure-31@3.[270,1318,181,1253]" captionTargetPageId="3" captionText="FIGURE 3. Strumigenys emmae worker and queen specimens.A) S. emmae worker full face view, B) S. emmae worker lateral view, C) S. emmae worker dorsal view. D) S. emmae queen full face view, E) S. emmae queen lateral view, F) S. emmae queen dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10695707" httpUri="https://zenodo.org/record/10695707/files/figure.png" pageId="4" pageNumber="574">figure 3 A</figureCitation>
); leading edge of scape with orbicular hairs (
<figureCitation id="7E85FBC4855849652A406D72FA1F0E32" box="[1322,1424,833,855]" captionStart="FIGURE 3" captionStartId="3.[152,255,1277,1301]" captionTargetBox="[270,1318,181,1253]" captionTargetId="figure-31@3.[270,1318,181,1253]" captionTargetPageId="3" captionText="FIGURE 3. Strumigenys emmae worker and queen specimens.A) S. emmae worker full face view, B) S. emmae worker lateral view, C) S. emmae worker dorsal view. D) S. emmae queen full face view, E) S. emmae queen lateral view, F) S. emmae queen dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10695707" httpUri="https://zenodo.org/record/10695707/files/figure.png" pageId="4" pageNumber="574">figure 3 A</figureCitation>
); mandibles averaging shorter, MI 24-35 (
<collectingCountry id="9EA9A7D1855849652D0E6D6CFC850E10" box="[612,778,863,885]" name="United Kingdom" pageId="4" pageNumber="574">United Kingdom</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652C7E6D6CFCD60E10" box="[788,857,863,885]" name="Poland" pageId="4" pageNumber="574">Poland</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652C086D6CFB8B0E10" box="[866,1028,863,885]" name="Netherlands" pageId="4" pageNumber="574">The Netherlands</collectingCountry>
)..........
<taxonomicName id="21BE9CC2855849652BEB6D6CFB470E11" authorityName="Myrmicinae" baseAuthorityName="Emery" baseAuthorityYear="1890" box="[1153,1224,863,884]" class="Insecta" family="Formicidae" genus="Strumigenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="emmae">
<emphasis id="D4CA3B53855849652BEB6D6CFB470E11" bold="true" box="[1153,1224,863,884]" italics="true" pageId="4" pageNumber="574">emmae</emphasis>
</taxonomicName>
(non-native[indoors])
</keyLead>
</paragraph>
<paragraph id="E601E741855849652FFD6D4EFA130ECA" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6D4EFA130ECA" pageId="4" pageNumber="574">- Head dorsum without orbicular hairs; antennae with six segments (including scape); scape long and linear, lacking convex median lobe; leading edge of scape without orbicular hairs; mandibles averaging longer, MI 34-57..................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="5D4AFFE4855849652FFD6D84FA13086A" pageId="4" pageNumber="574">
<paragraph id="E601E741855849652FFD6D84FA12093A" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6D84FA12093A" pageId="4" pageNumber="574">
<emphasis id="D4CA3B53855849652FFD6D84FF2C0EA9" bold="true" box="[151,163,951,972]" pageId="4" pageNumber="574">4</emphasis>
Ventrolateral margin of head interrupted by a deep, strongly incised preocular notch; in dorsal view anterior portion of eye is detached from side of head (
<emphasis id="D4CA3B53855849652D656DE7FD100E8F" box="[527,671,980,1002]" italics="true" pageId="4" pageNumber="574">figure 8 A &amp; B</emphasis>
). With head in ventral view the preocular notch forms the apex of a transverse impression in the ventral surface of the head capsule that extends toward the midline, well behind the postbuccal impression. First gastral sternite without a basal spongiform pad. Apicoscrobal hair absent. Pronotal humeral hair flagellate (
<emphasis id="D4CA3B53855849652A416A3CFA010941" box="[1323,1422,1039,1060]" italics="true" pageId="4" pageNumber="574">figure 7 C</emphasis>
). Mandibles each with 2 short preapical teeth, located close to the apicodorsal tooth, and the proximal preapical tooth longer than the distal (
<emphasis id="D4CA3B53855849652E2D6A7AFE28093A" box="[327,423,1097,1119]" italics="true" pageId="4" pageNumber="574">figure 6 B</emphasis>
) (
<collectingCountry id="9EA9A7D1855849652ED16A7AFDEE093A" box="[443,609,1097,1119]" name="United Kingdom" pageId="4" pageNumber="574">United Kingdom</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652D006A79FD4B093A" box="[618,708,1098,1119]" name="Germany" pageId="4" pageNumber="574">Germany</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652DA46A7AFCA5093A" box="[718,810,1097,1119]" name="Denmark" pageId="4" pageNumber="574">Denmark</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652C596A79FC0D093A" box="[819,898,1098,1119]" name="Norway" pageId="4" pageNumber="574">Norway</collectingCountry>
)....................
<taxonomicName id="21BE9CC2855849652BE66A7AFB47093B" authorityName="Emery" authorityYear="1890" box="[1164,1224,1097,1118]" class="Insecta" family="Formicidae" genus="Strumigenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="rogeri">
<emphasis id="D4CA3B53855849652BE66A7AFB47093B" bold="true" box="[1164,1224,1097,1118]" italics="true" pageId="4" pageNumber="574">rogeri</emphasis>
</taxonomicName>
(non-native[indoors])
</keyLead>
</paragraph>
<paragraph id="E601E741855849652FFD6A54FA13086A" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6A54FA13086A" pageId="4" pageNumber="574">
- Ventrolateral margin of head uninterrupted to anterior margin of eye; in dorsal view anterior portion of eye is not detached from side of head (
<emphasis id="D4CA3B53855849652EFC6AB7FDAC09FF" box="[406,547,1156,1178]" italics="true" pageId="4" pageNumber="574">figure 8 C &amp; D</emphasis>
). With head in ventral view without a transverse impression in the ventral surface of the head capsule posterior to the postbuccal impression. First gastral sternite with a basal spongiform pad. Apicoscrobal hair present. Pronotal humeral hair straight and simple. Mandibles either with 1 spiniform preapical tooth, located close to the apicodorsal tooth, or also with a denticle close to the midlength which is much shorter than the spiniform preapical tooth (
<emphasis id="D4CA3B53855849652A686AEFFA1D0997" box="[1282,1426,1244,1266]" italics="true" pageId="4" pageNumber="574">figure 6 C &amp; D</emphasis>
) ...................................................................................................
<emphasis id="D4CA3B53855849652AFA6AC9FA13086A" bold="true" box="[1424,1436,1274,1295]" pageId="4" pageNumber="574">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5D4AFFE4855849652FFD6B24FA120B50" pageId="4" pageNumber="574">
<paragraph id="E601E741855849652FFD6B24FA1208C7" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6B24FA1208C7" pageId="4" pageNumber="574">
<emphasis id="D4CA3B53855849652FFD6B24FF2C0849" bold="true" box="[151,163,1303,1324]" pageId="4" pageNumber="574">5</emphasis>
Leading edge of scape with all hairs curved toward the apex of the scape (
<emphasis id="D4CA3B53855849652CB06B24FBCE0849" box="[986,1089,1303,1324]" italics="true" pageId="4" pageNumber="574">figure 9 B</emphasis>
). Ventral surface of petiole with a conspicuous strip or curtain of spongiform material (
<emphasis id="D4CA3B53855849652DBC6B07FCBC082C" box="[726,819,1332,1353]" italics="true" pageId="4" pageNumber="574">figure 9 A</emphasis>
). Mandible with a single spiniform preapical tooth, located close to the apicodorsal tooth, without an additional denticle close to the midlength (
<emphasis id="D4CA3B53855849652CA36B62FBA90802" box="[969,1062,1361,1383]" italics="true" pageId="4" pageNumber="574">figure 9 B</emphasis>
). Dorsum of mesosoma obviously with more than one pair of standing hairs, on pronotum and mesonotum. Mandibles relatively slightly shorter, MI 34–48. (
<collectingCountry id="9EA9A7D1855849652A556B5CFA1208E1" box="[1343,1437,1391,1412]" name="Guernsey" pageId="4" pageNumber="574">Guernsey</collectingCountry>
[British Channel Islands])......................................................
<taxonomicName id="21BE9CC2855849652B0C6BBFFB3408C7" baseAuthorityName="Smith" baseAuthorityYear="1876" box="[1126,1211,1420,1442]" class="Insecta" family="Formicidae" genus="Strumigenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="D4CA3B53855849652B0C6BBFFB3408C7" bold="true" box="[1126,1211,1420,1442]" italics="true" pageId="4" pageNumber="574">perplexa</emphasis>
</taxonomicName>
(non-native[outdoors])
</keyLead>
</paragraph>
<paragraph id="E601E741855849652FFD6B99FA120B50" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6B99FA120B50" pageId="4" pageNumber="574">
- Leading edge of scape with 2 or more hairs curved toward the base of the scape (
<emphasis id="D4CA3B53855849652C826B9AFBC608DA" box="[1000,1097,1449,1471]" italics="true" pageId="4" pageNumber="574">figure 9 D</emphasis>
). Ventral surface of petiole entirely lacks spongiform material (
<emphasis id="D4CA3B53855849652E826BF4FDC808B9" box="[488,583,1479,1500]" italics="true" pageId="4" pageNumber="574">figure 9 C</emphasis>
). Mandible with a single spiniform preapical tooth, located close to the apicodorsal tooth, and also with an additional denticle close to the midlength (
<emphasis id="D4CA3B53855849652C7E6BD7FCFC089F" box="[788,883,1508,1530]" italics="true" pageId="4" pageNumber="574">figure 9 D</emphasis>
). Dorsum of mesosoma with only a single pair of standing hairs, on the mesonotum. Mandibles relatively slightly longer, MI 50–57 (
<collectingCountry id="9EA9A7D1855849652CCF6832FC770B72" box="[933,1016,1537,1559]" name="Portugal" pageId="4" pageNumber="574">Portugal</collectingCountry>
[
<collectingRegion id="247A29A3855849652B696832FBD50B72" box="[1027,1114,1537,1559]" country="Portugal" name="Madeira" pageId="4" pageNumber="574">Madeira</collectingRegion>
], southern Atlantic Islands,
<collectingRegion id="247A29A3855849652A096832FEA90B50" country="Portugal" name="Leiria" pageId="4" pageNumber="574">Leiria district</collectingRegion>
)....................................................................
<emphasis id="D4CA3B53855849652B03682DFB330B51" bold="true" box="[1129,1212,1566,1588]" italics="true" pageId="4" pageNumber="574">silvestrii</emphasis>
(non-native[outdoors])
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5D4AFFE4855849652FFD680FFA130A59" pageId="4" pageNumber="574">
<paragraph id="E601E741855849652FFD680FFA120BA2" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD680FFA120BA2" pageId="4" pageNumber="574">
<emphasis id="D4CA3B53855849652FFD680FFF2C0B34" bold="true" box="[151,163,1596,1617]" pageId="4" pageNumber="574">6</emphasis>
Humerus without a projecting hair; pronotum dorsolaterally with sharp raised margination (
<emphasis id="D4CA3B53855849652B26680FFB390B34" box="[1100,1206,1596,1617]" italics="true" pageId="4" pageNumber="574">figure 10 A</emphasis>
). Dorsal mesosoma and first gastral tergite without standing hairs (
<emphasis id="D4CA3B53855849652D15686AFD640B0A" box="[639,747,1625,1647]" italics="true" pageId="4" pageNumber="574">figure 10 A</emphasis>
). Leading edge of scape with 1–2 hairs near the subbasal bend that are distinctly curved toward the scape base (
<emphasis id="D4CA3B53855849652D006844FD590BE9" box="[618,726,1655,1676]" italics="true" pageId="4" pageNumber="574">figure 10 B</emphasis>
). Dorsal surface of mandible basally with a very distinct transverse sharp edge or rim that extends across its width, parallel to and in front of the anterior clypeal margin (figure 10 B). CI 84–90, SI 51–57. (
<collectingCountry id="9EA9A7D1855849652E5A6881FEFA0BA2" box="[304,373,1714,1735]" name="France" pageId="4" pageNumber="574">France</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652E156882FE370BA2" box="[383,440,1713,1735]" name="Spain" pageId="4" pageNumber="574">Spain</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652EA86882FE600BA2" box="[450,495,1713,1735]" name="Italy" pageId="4" pageNumber="574">Italy</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652E926881FDB10BA2" box="[504,574,1714,1735]" name="Greece" pageId="4" pageNumber="574">Greece</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652D226882FD010BA2" box="[584,654,1713,1735]" name="Turkey" pageId="4" pageNumber="574">Turkey</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652DFD6881FD690BA2" box="[663,742,1714,1735]" name="Norway" pageId="4" pageNumber="574">Norway</collectingCountry>
)..................
<emphasis id="D4CA3B53855849652CA26882FBDA0BA2" bold="true" box="[968,1109,1713,1735]" italics="true" pageId="4" pageNumber="574">membranifera</emphasis>
(non-native[outdoors &amp; indoors])
</keyLead>
</paragraph>
<paragraph id="E601E741855849652FFD68FCFA130A59" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD68FCFA130A59" pageId="4" pageNumber="574">
- Humerus with a projecting simple hair; pronotum dorsolaterally without sharp raised margination (
<emphasis id="D4CA3B53855849652BA468FCFACB0B81" box="[1230,1348,1743,1764]" italics="true" pageId="4" pageNumber="574">figure 10 C</emphasis>
). Dorsal mesosoma and first gastral tergite with standing hairs (
<emphasis id="D4CA3B53855849652D9168DFFCE40A64" box="[763,875,1772,1793]" italics="true" pageId="4" pageNumber="574">figure 10 C</emphasis>
). Leading edge of scape with all hairs curved or inclined toward the scape apex (
<emphasis id="D4CA3B53855849652EAE693AFDBD0A7A" box="[452,562,1801,1823]" italics="true" pageId="4" pageNumber="574">figure 10 D</emphasis>
). Dorsal surface of mandible basally without a transverse sharp edge or rim running across its width (
<emphasis id="D4CA3B53855849652E296914FE3E0A59" box="[323,433,1831,1852]" italics="true" pageId="4" pageNumber="574">figure 10 D</emphasis>
). CI 66–72, SI 69–76.................................................................
<emphasis id="D4CA3B53855849652AFA6914FA130A59" bold="true" box="[1424,1436,1831,1852]" pageId="4" pageNumber="574">7</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5D4AFFE4855849642FFD6977FA130D82" lastPageId="5" lastPageNumber="575" pageId="4" pageNumber="574">
<paragraph id="E601E741855849652FFD6977FA120AAA" blockId="4.[151,1437,364,1999]" pageId="4" pageNumber="574">
<keyLead id="5D4F4474855849652FFD6977FA120AAA" pageId="4" pageNumber="574">
<emphasis id="D4CA3B53855849652FFD6977FF2C0A3C" bold="true" box="[151,163,1860,1881]" pageId="4" pageNumber="574">7</emphasis>
Dorsum of clypeus in full-face view densely clothed with conspicuous broadly spatulate to spoon-shaped hairs; in profile these hairs parallel with the surface from which they arise and closely applied (
<emphasis id="D4CA3B53855849652C8E6952FBC10A12" box="[996,1102,1889,1911]" italics="true" pageId="4" pageNumber="574">figure 11 A</emphasis>
). Medially curved ground-pilosity bordering the upper scrobe margins distinctly spatulate. (
<collectingCountry id="9EA9A7D1855849652C7A694CFCD30AF0" box="[784,860,1919,1941]" name="Algeria" pageId="4" pageNumber="574">Algeria</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652C0C694CFC320AF0" box="[870,957,1919,1941]" name="Armenia" pageId="4" pageNumber="574">Armenia</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652CA2694CFB920AF0" box="[968,1053,1919,1941]" name="Bulgaria" pageId="4" pageNumber="574">Bulgaria</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652B42694CFBFE0AF0" box="[1064,1137,1919,1941]" name="Croatia" pageId="4" pageNumber="574">Croatia</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652B17694CFB310AF1" box="[1149,1214,1919,1940]" name="France" pageId="4" pageNumber="574">France</collectingCountry>
(southern mainland &amp;
<collectingCountry id="9EA9A7D1855849652F8869AFFEA00AD7" box="[226,303,1948,1970]" name="France" pageId="4" pageNumber="574">Corsica</collectingCountry>
),
<collectingCountry id="9EA9A7D1855849652E2A69AFFE090AD4" box="[320,390,1948,1969]" name="Greece" pageId="4" pageNumber="574">Greece</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652EFB69AEFE690AD7" box="[401,486,1949,1970]" name="Hungary" pageId="4" pageNumber="574">Hungary</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652E9B69AFFD920AD7" box="[497,541,1948,1970]" name="Italy" pageId="4" pageNumber="574">Italy</collectingCountry>
(mainland,
<collectingRegion id="247A29A3855849652DF869AFFD6D0AD7" box="[658,738,1948,1970]" country="Italy" name="Sardegna" pageId="4" pageNumber="574">Sardinia</collectingRegion>
&amp;
<collectingRegion id="247A29A3855849652C6969AFFCB10AD7" box="[771,830,1948,1970]" country="Italy" name="Sicilia" pageId="4" pageNumber="574">Sicily</collectingRegion>
), Macedonia,
<collectingCountry id="9EA9A7D1855849652CAD69AFFB8E0AD7" box="[967,1025,1948,1970]" name="Malta" pageId="4" pageNumber="574">Malta</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652B6669AEFB0B0AD7" box="[1036,1156,1949,1970]" name="Serbia" pageId="4" pageNumber="574">Montenegro</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652BE569AEFB670AD7" box="[1167,1256,1949,1970]" name="Morocco" pageId="4" pageNumber="574">Morocco</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652B9869AFFAC40AD7" box="[1266,1355,1948,1970]" name="Romania" pageId="4" pageNumber="574">Romania</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652A3C69AFFA170AD7" box="[1366,1432,1948,1970]" name="Russia" pageId="4" pageNumber="574">Russia</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652F88698AFEAD0AAA" box="[226,290,1977,1999]" name="Serbia" pageId="4" pageNumber="574">Serbia</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652E41698AFEEB0AAA" box="[299,356,1977,1999]" name="Spain" pageId="4" pageNumber="574">Spain</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652E04698AFE6C0AAA" box="[366,483,1977,1999]" name="Switzerland" pageId="4" pageNumber="574">Switzerland</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652E86698AFDB90AAA" box="[492,566,1977,1999]" name="Tunisia" pageId="4" pageNumber="574">Tunisia</collectingCountry>
,
<collectingCountry id="9EA9A7D1855849652D55698AFD0A0AAA" box="[575,645,1977,1999]" name="Turkey" pageId="4" pageNumber="574">Turkey</collectingCountry>
)...................................................
<emphasis id="D4CA3B53855849652B98698AFAC40AAA" bold="true" box="[1266,1355,1977,1999]" italics="true" pageId="4" pageNumber="574">baudueri</emphasis>
(native)
</keyLead>
</paragraph>
<paragraph id="E601E741855949642FFD6EABFA130D82" blockId="5.[151,1437,151,349]" pageId="5" pageNumber="575">
<keyLead id="5D4F4474855949642FFD6EABFA130D82" pageId="5" pageNumber="575">
- Dorsum of clypeus in full-face view clothed with slender hairs that are very narrowly spatulate or simple and cylindrical; in profile these hairs markedly elevated and either arched or inclined anteriorly (
<emphasis id="D4CA3B53855949642CB56E86FBC30DAF" box="[991,1100,181,202]" italics="true" pageId="5" pageNumber="575">figure 11 B</emphasis>
). Medially curved ground-pilosity bordering the upper scrobe margins fine and simple..........................................................
<emphasis id="D4CA3B53855949642AFA6EE1FA130D82" bold="true" box="[1424,1436,210,231]" pageId="5" pageNumber="575">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<keyStep id="5D4AFFE4855949642FFD6EDCFA120C38" pageId="5" pageNumber="575">
<paragraph id="E601E741855949642FFD6EDCFA120C46" blockId="5.[151,1437,151,349]" pageId="5" pageNumber="575">
<keyLead id="5D4F4474855949642FFD6EDCFA120C46" pageId="5" pageNumber="575">
<emphasis id="D4CA3B53855949642FFD6EDCFF2C0C61" bold="true" box="[151,163,239,260]" pageId="5" pageNumber="575">8</emphasis>
Hairs on clypeal dorsum narrowly spatulate; in profile the hairs distinctly anteriorly curved or anteriorly arched (
<emphasis id="D4CA3B53855949642A496EC3FA000C60" box="[1315,1423,240,261]" italics="true" pageId="5" pageNumber="575">figure 12 A</emphasis>
). (
<collectingCountry id="9EA9A7D1855949642F826F3EFEB10C46" box="[232,318,269,291]" name="Bulgaria" pageId="5" pageNumber="575">Bulgaria</collectingCountry>
,
<collectingCountry id="9EA9A7D1855949642E226F3EFE060C47" box="[328,393,269,290]" name="France" pageId="5" pageNumber="575">France</collectingCountry>
(mainland &amp;
<collectingCountry id="9EA9A7D1855949642D7A6F3EFDD20C46" box="[528,605,269,291]" name="France" pageId="5" pageNumber="575">Corsica</collectingCountry>
),
<collectingCountry id="9EA9A7D1855949642D076F3EFD3C0C47" box="[621,691,269,290]" name="Greece" pageId="5" pageNumber="575">Greece</collectingCountry>
,
<collectingCountry id="9EA9A7D1855949642DD76F3EFD650C46" box="[701,746,269,291]" name="Italy" pageId="5" pageNumber="575">Italy</collectingCountry>
,
<collectingCountry id="9EA9A7D1855949642D996F3EFCA30C46" box="[755,812,269,291]" name="Spain" pageId="5" pageNumber="575">Spain</collectingCountry>
,
<collectingCountry id="9EA9A7D1855949642C5C6F3EFCF30C46" box="[822,892,269,291]" name="Turkey" pageId="5" pageNumber="575">Turkey</collectingCountry>
)..............................
<emphasis id="D4CA3B53855949642B9A6F3FFAC30C47" bold="true" box="[1264,1356,268,290]" italics="true" pageId="5" pageNumber="575">tenuipilis</emphasis>
(native)
</keyLead>
</paragraph>
<paragraph id="E601E741855949642FFD6F18FA120C38" blockId="5.[151,1437,151,349]" pageId="5" pageNumber="575">
<keyLead id="5D4F4474855949642FFD6F18FA120C38" pageId="5" pageNumber="575">
- Hairs on clypeal dorsum simple, fine and cylindrical throughout their length and tapered or truncated apically; in profile the hairs mostly straight, inclined anteriorly (
<emphasis id="D4CA3B53855949642D1A6F7BFD530C38" box="[624,732,328,349]" italics="true" pageId="5" pageNumber="575">figure 12 B</emphasis>
). (
<collectingCountry id="9EA9A7D1855949642D9C6F7BFCB60C38" box="[758,825,328,349]" name="France" pageId="5" pageNumber="575">France</collectingCountry>
(
<collectingCountry id="9EA9A7D1855949642C2F6F74FC1C0C38" box="[837,915,327,349]" name="France" pageId="5" pageNumber="575">Corsica</collectingCountry>
),
<collectingCountry id="9EA9A7D1855949642CCE6F7BFC650C38" box="[932,1002,328,349]" name="Greece" pageId="5" pageNumber="575">Greece</collectingCountry>
)...................
<emphasis id="D4CA3B53855949642B8B6F74FAC30C39" bold="true" box="[1249,1356,327,348]" italics="true" pageId="5" pageNumber="575">tenuissima</emphasis>
(native)
</keyLead>
</paragraph>
</keyStep>
</key>
</treatment>
</document>
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