254 lines
36 KiB
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254 lines
36 KiB
XML
<document id="7CDCD29B77B0469F1DCF83348A095FCD" ID-CLB-Dataset="32271" ID-DOI="10.11646/zootaxa.4312.2.5" ID-GBIF-Dataset="2a43bafe-bc08-4970-a7b5-6d986ddcd28a" ID-ISSN="1175-5326" ID-Zenodo-Dep="852866" ID-ZooBank="A167Fbbf-Ceef-4Bec-88B4-Cbb4238195E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503986631928" checkinUser="plazi" docAuthor="Zhou, Hong-Zhang" docDate="2017" docId="6E160623FFD4B9640FAEFC0B923CF851" docLanguage="en" docName="zootaxa.4312.2.5.pdf" docOrigin="Zootaxa 4312 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ocypus Leach 1819" docType="treatment" docVersion="8" lastPageNumber="294" masterDocId="922F7E5BFFD5B9650F39FFFE9757FF8C" masterDocTitle="Taxonomy of the subgenus Ocypus Leach (Coleoptera: Staphylinidae: Staphylinini) and three new species from China" masterLastPageNumber="310" masterPageNumber="293" pageNumber="294" updateTime="1698467505264" updateUser="ExternalLinkService">
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<mods:title id="AEFF9195E75793CAAE57E2FD93AC647F">Taxonomy of the subgenus Ocypus Leach (Coleoptera: Staphylinidae: Staphylinini) and three new species from China</mods:title>
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<mods:namePart id="7B300A80FD9D80B6706A49CCC328D88C">Zhou, Hong-Zhang</mods:namePart>
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<mods:title id="E0B6DF4A8675404C30AB6211235002E1">Zootaxa</mods:title>
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<mods:part id="2EAA134192DC123E36AFC8EBBFA253C9">
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<mods:date id="4FE5724077C5F7F607045CA6461E090E">2017</mods:date>
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<mods:number id="4A922A2416DC68B87C660650E394A1E0">2017-08-29</mods:number>
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<mods:number id="80E091E645EAECD180887725498B7CCC">4312</mods:number>
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<treatment id="6E160623FFD4B9640FAEFC0B923CF851" ID-DOI="http://doi.org/10.5281/zenodo.6028536" ID-GBIF-Taxon="133010619" ID-Zenodo-Dep="6028536" LSID="urn:lsid:plazi:treatment:6E160623FFD4B9640FAEFC0B923CF851" httpUri="http://treatment.plazi.org/id/6E160623FFD4B9640FAEFC0B923CF851" lastPageNumber="294" pageId="1" pageNumber="294">
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<subSubSection id="AEA5E4BEFFD4B9640FAEFC0B9488FA84" pageId="1" pageNumber="294" type="nomenclature">
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<paragraph id="E600B735FFD4B9640FAEFC0B9559FB83" blockId="1.[151,526,1012,1039]" box="[151,526,1012,1039]" pageId="1" pageNumber="294">
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<heading id="BD480059FFD4B9640FAEFC0B9559FB83" bold="true" box="[151,526,1012,1039]" fontSize="11" level="1" pageId="1" pageNumber="294" reason="1">
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<emphasis id="D4CB6B27FFD4B9640FAEFC0B9559FB83" bold="true" box="[151,526,1012,1039]" pageId="1" pageNumber="294">
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Subgenus
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<taxonomicName id="21BFCCB6FFD4B9640E2EFC0A9559FB83" ID-CoL="8VRWY" authority="Leach, 1819" authorityName="Leach" authorityYear="1819" box="[279,526,1012,1039]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
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<emphasis id="D4CB6B27FFD4B9640E2EFC0A9627FB82" bold="true" box="[279,368,1012,1038]" italics="true" pageId="1" pageNumber="294">Ocypus</emphasis>
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<bibRefCitation id="822ECAC4FFD4B9640E4EFC0B9559FB83" author="Leach" box="[375,526,1013,1039]" pageId="1" pageNumber="294" refString="Leach, W. E. (1819) In: Samouelle, G., The Entomologist's useful compendium; or an introduction to the knowledge of British insects, comprising the best means of obtaining and preserving them, and a description of the apparatus generally used; together with the genera of Linne, and the modern method of arranging the classes Crustacea, Myriapoda, spiders, mites, and insects from their affinities and structure, according to the views of Dr. Leach. Also an explanation of the terms used in entomology; a calendar of the times of appearance, and usual situations of near 3000 species of British insects; with instructions for collecting and fitting up objects for the microscope. Thomas Boys, London, 496 pp." type="book" year="1819">Leach, 1819</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E600B735FFD4B9640FAEFBC49488FA84" blockId="1.[151,1436,1082,1288]" pageId="1" pageNumber="294">
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<bibRefCitation id="822ECAC4FFD4B9640FAEFBC4964DFBDC" author="Leach" box="[151,282,1082,1104]" pageId="1" pageNumber="294" refString="Leach, W. E. (1819) In: Samouelle, G., The Entomologist's useful compendium; or an introduction to the knowledge of British insects, comprising the best means of obtaining and preserving them, and a description of the apparatus generally used; together with the genera of Linne, and the modern method of arranging the classes Crustacea, Myriapoda, spiders, mites, and insects from their affinities and structure, according to the views of Dr. Leach. Also an explanation of the terms used in entomology; a calendar of the times of appearance, and usual situations of near 3000 species of British insects; with instructions for collecting and fitting up objects for the microscope. Thomas Boys, London, 496 pp." type="book" year="1819">Leach, 1819</bibRefCitation>
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: 172 (species included:
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<taxonomicName id="21BFCCB6FFD4B9640D20FBC5953DFBDC" authorityName="Paykull" authorityYear="1789" box="[537,618,1083,1104]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="cyaneus">
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<emphasis id="D4CB6B27FFD4B9640D20FBC5953DFBDC" box="[537,618,1083,1104]" italics="true" pageId="1" pageNumber="294">cyaneus</emphasis>
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</taxonomicName>
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);
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<bibRefCitation id="822ECAC4FFD4B9640DBAFBC49494FBDC" author="Bernhauer" box="[643,963,1082,1104]" pageId="1" pageNumber="294" refString="Bernhauer, M. & Schubert, K. (1914) Staphylinidae IV. Pars 57. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Fol. F. Staphylinidae. W. Junk, Berlin, pp. 289 - 408." type="book chapter" year="1914">Bernhauer and Schubert, 1914</bibRefCitation>
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: 374 (subgenus of
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<taxonomicName id="21BFCCB6FFD4B9640BB4FBC49255FBDC" box="[1165,1282,1082,1104]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
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<emphasis id="D4CB6B27FFD4B9640BB4FBC49255FBDC" box="[1165,1282,1082,1104]" italics="true" pageId="1" pageNumber="294">Staphylinus</emphasis>
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</taxonomicName>
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; authorship as Stephens, 1832; world catalog);
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<bibRefCitation id="822ECAC4FFD4B9640D23FBA795EBFBE3" author="Cameron" box="[538,700,1113,1135]" pageId="1" pageNumber="294" refString="Cameron, M. (1932) The fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Fol. 3. Taylor and Francis, London, xiii + 443 pp." type="book" year="1932">Cameron, 1932</bibRefCitation>
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: 180 (subgenus of
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<taxonomicName id="21BFCCB6FFD4B9640CBCFBA794ADFBE3" box="[901,1018,1113,1135]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
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<emphasis id="D4CB6B27FFD4B9640CBCFBA794ADFBE3" box="[901,1018,1113,1135]" italics="true" pageId="1" pageNumber="294">Staphylinus</emphasis>
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</taxonomicName>
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);
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<bibRefCitation id="822ECAC4FFD4B9640B2AFBA79398FBE3" author="Scheerpeltz" box="[1043,1231,1113,1135]" pageId="1" pageNumber="294" refString="Scheerpeltz, O. (1933): Staphylinidae VII (Pars 129). In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Fol. FI. Staphylinidae. Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">Scheerpeltz, 1933</bibRefCitation>
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: 1381 (subgenus of
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<taxonomicName id="21BFCCB6FFD4B9640FFEFB86966BFB02" box="[199,316,1144,1166]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
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<emphasis id="D4CB6B27FFD4B9640FFEFB86966BFB02" box="[199,316,1144,1166]" italics="true" pageId="1" pageNumber="294">Staphylinus</emphasis>
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</taxonomicName>
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, catalog supplement);
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<bibRefCitation id="822ECAC4FFD4B9640D18FB8695F8FB02" author="Shibata" box="[545,687,1144,1166]" pageId="1" pageNumber="294" refString="Shibata, Y. (1973) Preliminary check list of the family Staphylinidae of Taiwan (Insecta: Coleoptera). Annual Bulletin of the Nichidai Sanko, 16, 21 - 88." type="journal article" year="1973">Shibata, 1973</bibRefCitation>
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: 62 (checklist of species of Taiwan);
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<bibRefCitation id="822ECAC4FFD4B9640B1CFB8693E1FB02" author="Coiffait" box="[1061,1206,1144,1166]" pageId="1" pageNumber="294" refString="Coiffait, H. (1974) Coleopteres staphylinides de la region palearctique occidentale II. Sous famille Staphilininae, Tribus Philonthini et Staphylinini. Nouvelle Revue d'Entomologie, Supplement 4 (4), 1 - 593." type="journal article" year="1974">Coiffait, 1974</bibRefCitation>
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: 420 (key to species of the western Palearctic Region);
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<bibRefCitation id="822ECAC4FFD4B9640D29FB6895F7FB20" author="Shibata" box="[528,672,1174,1196]" pageId="1" pageNumber="294" refString="Shibata, Y. (1984) Provisional check list of the family Staphylinidae of Japan. IV. (Insecta: Coleoptera). Annual Bulletin of the Nichidai Sanko, 22, 79 - 141." type="journal article" year="1984">Shibata, 1984</bibRefCitation>
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: 87 (checklist of species of Japan);
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<bibRefCitation id="822ECAC4FFD4B9640B2BFB68927EFB20" author="Smetana" box="[1042,1321,1174,1196]" pageId="1" pageNumber="294" refString="Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1 - 88." type="journal article" year="2000">Smetana and Davies, 2000</bibRefCitation>
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: 21, 31, 42 (characters; checklist of species of north temperate region);
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<bibRefCitation id="822ECAC4FFD4B9640C07FB489482FB47" author="Herman" box="[830,981,1205,1227]" pageId="1" pageNumber="294" refString="Herman, L. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini. Staphylinidae Incertae sedis, fossils, Protactinae. Bulletin of the American Museum of Natural History, 265, i - v + 3021 - 3839." type="journal article" year="2001">Herman, 2001</bibRefCitation>
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: 3355 (world catalog);
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<bibRefCitation id="822ECAC4FFD4B9640BF5FB4B923DFB47" author="Smetana" box="[1228,1386,1205,1227]" pageId="1" pageNumber="294" refString="Smetana, A. (2004) Staphylininae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Fol. 2. Apollo Books, Stenstrup, pp. 624 - 698." type="journal article" year="2004">Smetana, 2004</bibRefCitation>
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: 674 (Catalog of Palaearctic Region);
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<bibRefCitation id="822ECAC4FFD4B9640D2FFB2A95E7FB66" author="Smetana" box="[534,688,1236,1258]" pageId="1" pageNumber="294" refString="Smetana, A. (2007) Contributions to the knowledge of the " Staphylinus - complex " (Coleoptera, Staphylinidae, Staphylinini) of China. Part 16. The genus Ocypus Leach, 1819, subgenera Ocypus s. str. and Matidus Motschulsky, 1860. Elytra, 35, 441 - 459." type="journal article" year="2007">Smetana, 2007</bibRefCitation>
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: 443 (key to species of China);
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<bibRefCitation id="822ECAC4FFD4B9640CCDFB2A9256FB66" author="Schulke" box="[1012,1281,1236,1258]" pageId="1" pageNumber="294" refString="Schulke, M. & Smetana, A. (2015) Staphylinidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Fol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 901 - 1134." type="journal article" year="2015">Schülke & Smetana, 2015</bibRefCitation>
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: 1089 (Catalog of Palaearctic Region, including the synonyms
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<taxonomicName id="21BFCCB6FFD4B9640D9CFB0C944AFA84" authorityName="Coiffait" authorityYear="1982" box="[677,797,1266,1288]" class="Insecta" family="Staphylinidae" genus="Nudabemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
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<emphasis id="D4CB6B27FFD4B9640D9CFB0C944AFA84" box="[677,797,1266,1288]" italics="true" pageId="1" pageNumber="294">Nudabemus</emphasis>
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</taxonomicName>
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and
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<emphasis id="D4CB6B27FFD4B9640C77FB0C9486FA84" box="[846,977,1266,1288]" italics="true" pageId="1" pageNumber="294">Xanthocypus</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="AEA5E4BEFFD4B9640FAEFACB969BFA2B" pageId="1" pageNumber="294" type="materials_examined">
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<paragraph id="E600B735FFD4B9640FAEFACB92CBFA04" blockId="1.[151,1436,1333,1509]" pageId="1" pageNumber="294">
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<emphasis id="D4CB6B27FFD4B9640FAEFACB9671FAC7" bold="true" box="[151,294,1333,1355]" pageId="1" pageNumber="294">Type species:</emphasis>
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<taxonomicName id="21BFCCB6FFD4B9640E12FACB95D9FAC7" authority="Paykull, 1789" authorityName="Paykull" authorityYear="1789" box="[299,654,1333,1355]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="cyaneus">
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<emphasis id="D4CB6B27FFD4B9640E12FACB96AFFAC7" box="[299,504,1333,1355]" italics="true" pageId="1" pageNumber="294">Staphylinus cyaneus</emphasis>
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Paykull, 1789
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</taxonomicName>
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, fixed by original designation and monotypy.
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<emphasis id="D4CB6B27FFD4B9640FAEFAAA9799FAE6" bold="true" box="[151,206,1364,1386]" pageId="1" pageNumber="294">Syn.:</emphasis>
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<taxonomicName id="21BFCCB6FFD4B9640FEDFAAA9557FAE6" authority="Westwood, 1827: 58" authorityName="Westwood" authorityPageNumber="58" authorityYear="1827" box="[212,512,1364,1386]" class="Insecta" family="Staphylinidae" genus="Goerius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
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<emphasis id="D4CB6B27FFD4B9640FEDFAAA9672FAE5" box="[212,293,1364,1385]" italics="true" pageId="1" pageNumber="294">Goerius</emphasis>
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<bibRefCitation id="822ECAC4FFD4B9640E15FAAA968DFAE6" author="Westwood" box="[300,474,1364,1386]" pageId="1" pageNumber="294" refString="Westwood, J. O. (1827) Observations upon Siagonium quadricorne of Kirby, and on other portions of the Brachelytra (Staphylinus Lin.). The Zoological Journal, 3, 56 - 66." type="journal article" year="1827">Westwood, 1827</bibRefCitation>
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: 58
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</taxonomicName>
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. Type species:
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<taxonomicName id="21BFCCB6FFD4B9640DA1FAAA94A5FAE6" authority="O. Muller, 1764" authorityName="O. Muller" authorityYear="1764" box="[664,1010,1364,1386]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="olens">
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<emphasis id="D4CB6B27FFD4B9640DA1FAAA941DFAE6" box="[664,842,1364,1386]" italics="true" pageId="1" pageNumber="294">Staphylinus olens</emphasis>
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O. Müller, 1764
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</taxonomicName>
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by monotypy.
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<taxonomicName id="21BFCCB6FFD4B9640FAEFA8D9687FA04" authority="Gistel, 1856: 388" authorityName="Gistel" authorityPageNumber="388" authorityYear="1856" box="[151,464,1394,1416]" class="Insecta" family="Staphylinidae" genus="Isopterum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
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<emphasis id="D4CB6B27FFD4B9640FAEFA8D97A9FA04" box="[151,254,1395,1416]" italics="true" pageId="1" pageNumber="294">Isopterum</emphasis>
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<bibRefCitation id="822ECAC4FFD4B9640E35FA8C96C2FA04" author="Gistel" box="[268,405,1394,1416]" pageId="1" pageNumber="294" refString="Gistel, J. N. F. X. (1856) Die Mysterien der europaischen Insectenwelt. Ein geheimer Schlussel fur Sammler aller Insecten- Ordnungen und Stande, behufs des Fangs, des Aufenhalts-Orts, der Wohnung, Tag- und Jahreszeit u. s. w., oder autoptische Darstellung des Insectenstaats in seinem Zusammenhange zum Bestehen des Naturhaushaltes uberhaupt und insbesondere in seinem Einflusse auf die phanerogamische und cryptogamische Pflanzenbevolkerung Europa's. Zum ersten Male nach funfundzwanzigjahrigen eigenen Erfahrungen zusammengestellt und herausgegeben. T. Dannheimer, Kempten, xii + 530 pp." type="book" year="1856">Gistel, 1856</bibRefCitation>
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: 388
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</taxonomicName>
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. Type species:
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<taxonomicName id="21BFCCB6FFD4B9640DBDFA8C9357FA04" authority="Paykull, 1789" authorityName="Paykull" authorityYear="1789" box="[644,1024,1394,1416]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="cyaneus">
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<emphasis id="D4CB6B27FFD4B9640DBDFA8C940DFA04" box="[644,858,1394,1416]" italics="true" pageId="1" pageNumber="294">Staphylinus cyaneus</emphasis>
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Paykull, 1789
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</taxonomicName>
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, fixed by subsequent designation by
|
|
</paragraph>
|
|
<paragraph id="E600B735FFD4B9640FFEFA6F969BFA2B" blockId="1.[151,1436,1333,1509]" box="[199,460,1425,1447]" pageId="1" pageNumber="294">Blackwelder, 1952: 2015.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="AEA5E4BEFFD4B9640FAEFA4E9614F952" pageId="1" pageNumber="294" type="reference_group">
|
|
<paragraph id="E600B735FFD4B9640FAEFA4E94F1FA68" blockId="1.[151,1436,1333,1509]" pageId="1" pageNumber="294">
|
|
<treatmentCitationGroup id="C6AF901BFFD4B9640FAEFA4E94F1FA68" pageId="1" pageNumber="294">
|
|
<taxonomicName id="21BFCCB6FFD4B9640FAEFA4E968FFA4A" ID-CoL="9LFH5" authority="Coiffait, 1982 b: 74" authorityName="Coiffait" authorityPageNumber="74" authorityYear="1982" box="[151,472,1456,1478]" class="Insecta" family="Staphylinidae" genus="Nudabemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="D4CB6B27FFD4B9640FAEFA4E9658FA4A" box="[151,271,1456,1478]" italics="true" pageId="1" pageNumber="294">Nudabemus</emphasis>
|
|
<treatmentCitation id="671E9124FFD4B9640E2DFA4E968FFA4A" author="Coiffait" box="[276,472,1456,1478]" page="74" pageId="1" pageNumber="294" year="1982">
|
|
<bibRefCitation id="822ECAC4FFD4B9640E2DFA4E96E5FA4A" author="Coiffait" box="[276,434,1456,1478]" pageId="1" pageNumber="294" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982" yearSuffix="b">Coiffait, 1982b</bibRefCitation>
|
|
: 74
|
|
</treatmentCitation>
|
|
</taxonomicName>
|
|
. Type species
|
|
<taxonomicName id="21BFCCB6FFD4B9640D53FA4E9356FA4A" ID-CoL="483J3" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[618,1025,1456,1478]" class="Insecta" family="Staphylinidae" genus="Nudabemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="caerulescens">
|
|
<emphasis id="D4CB6B27FFD4B9640D53FA4E943CFA4A" box="[618,875,1456,1478]" italics="true" pageId="1" pageNumber="294">Nudabemus caerulescens</emphasis>
|
|
<bibRefCitation id="822ECAC4FFD4B9640C48FA4E9356FA4A" author="Coiffait" box="[881,1025,1456,1478]" pageId="1" pageNumber="294" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
|
|
</taxonomicName>
|
|
, fixed by original designation.
|
|
<emphasis id="D4CB6B27FFD4B9640FAEFA30964DFA68" box="[151,282,1486,1508]" italics="true" pageId="1" pageNumber="294">Xanthocypus</emphasis>
|
|
<treatmentCitation id="671E9124FFD4B9640E19FA3196B8FA68" author="Muller" box="[288,495,1486,1508]" page="40" pageId="1" pageNumber="294" year="1925">
|
|
<bibRefCitation id="822ECAC4FFD4B9640E19FA31969DFA68" author="Muller" box="[288,458,1486,1508]" pageId="1" pageNumber="294" refString="Muller, J. (1925) Terzo contribute alia conoscenza del genere Staphylinus L .. Bollettino della Societa Entomologica Italiana, 57, 40 - 48." type="journal article" year="1925" yearSuffix="d">J. Müller, 1925d</bibRefCitation>
|
|
: 40
|
|
</treatmentCitation>
|
|
. Type species
|
|
<taxonomicName id="21BFCCB6FFD4B9640DBBFA3194F5FA68" ID-CoL="48MLK" authority="Harold, 1877" authorityName="Harold" authorityYear="1877" box="[642,930,1486,1509]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="weisei">
|
|
<emphasis id="D4CB6B27FFD4B9640DBBFA319444FA69" box="[642,787,1487,1509]" italics="true" pageId="1" pageNumber="294">Ocypus weisei</emphasis>
|
|
<bibRefCitation id="822ECAC4FFD4B9640C20FA3094F5FA68" author="Harold" box="[793,930,1486,1508]" pageId="1" pageNumber="294" refString="Harold, E. (1877) Beitrage zur Kaferfauna von Japan. Deutsche Entomologische Zeitschrift, 21, 337 - 367. https: // doi. org / 10.1002 / mmnd. 4800210220" type="journal article" year="1877">Harold, 1877</bibRefCitation>
|
|
</taxonomicName>
|
|
.
|
|
</treatmentCitationGroup>
|
|
</paragraph>
|
|
<paragraph id="E600B735FFD4B9640FAEF9EF9614F952" blockId="1.[151,1436,1553,2014]" pageId="1" pageNumber="294">
|
|
The species of the subgenus
|
|
<taxonomicName id="21BFCCB6FFD4B9640EE4F9EC9566F9A5" ID-CoL="8VRWY" authority="Zhou, 2017" authorityName="Zhou" authorityYear="2017" box="[477,561,1554,1577]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|
<emphasis id="D4CB6B27FFD4B9640EE4F9EC9566F9A5" box="[477,561,1554,1577]" italics="true" pageId="1" pageNumber="294">Ocypus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="CFF8D65CFFD4B9640D00F9EC9522F9A5" box="[569,629,1554,1577]" pageId="1" pageNumber="294" sensu="stricto">s. str.</taxonomicNameLabel>
|
|
can be easily distinguished from those of other subgenera of the genus
|
|
<taxonomicName id="21BFCCB6FFD4B9640FAEF9C897BCF9C1" ID-CoL="8VRWY" authorityName="Zhou" authorityYear="2017" box="[151,235,1590,1613]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="D4CB6B27FFD4B9640FAEF9C897BCF9C1" box="[151,235,1590,1613]" italics="true" pageId="1" pageNumber="294">Ocypus</emphasis>
|
|
</taxonomicName>
|
|
by the following characteristics: last segment of maxillary palpi asetose; that of labial palpi setose, usually short, with truncate apex; body size relatively large (more than
|
|
<quantity id="21471AD0FFD4B9640C5EF9A794ECF9FE" box="[871,955,1625,1650]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="294" unit="mm" value="15.0">15 mm</quantity>
|
|
), body color usually totally black; palpifer usually with three setae on apical portion; eyes small and tempora relatively long; genal seta at least four puncture diameters away from posterior margin of eye; sternite 9 of male genital segment with tapered basal portion rather short and wide.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="AEA5E4BEFFD4B9640FFEF910923CF851" pageId="1" pageNumber="294" type="description">
|
|
<paragraph id="E600B735FFD4B9640FFEF910923CF851" blockId="1.[151,1436,1553,2014]" pageId="1" pageNumber="294">
|
|
In the current taxonomy (
|
|
<bibRefCitation id="822ECAC4FFD4B9640ECDF9139467F889" author="Smetana" box="[500,816,1773,1798]" pageId="1" pageNumber="294" refString="Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1 - 88." type="journal article" year="2000">Smetana and Davies, 2000</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="822ECAC4FFD4B9640C79F91094BFF889" author="Herman" box="[832,1000,1773,1797]" pageId="1" pageNumber="294" refString="Herman, L. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini. Staphylinidae Incertae sedis, fossils, Protactinae. Bulletin of the American Museum of Natural History, 265, i - v + 3021 - 3839." type="journal article" year="2001">Herman, 2001</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="822ECAC4FFD4B9640CC1F91393C9F889" author="Smetana" box="[1016,1182,1773,1797]" pageId="1" pageNumber="294" refString="Smetana, A. (2007) Contributions to the knowledge of the " Staphylinus - complex " (Coleoptera, Staphylinidae, Staphylinini) of China. Part 16. The genus Ocypus Leach, 1819, subgenera Ocypus s. str. and Matidus Motschulsky, 1860. Elytra, 35, 441 - 459." type="journal article" year="2007">Smetana 2007</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="822ECAC4FFD4B9640B97F9139784F8A6" author="Schulke" pageId="1" pageNumber="294" refString="Schulke, M. & Smetana, A. (2015) Staphylinidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Fol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 901 - 1134." type="journal article" year="2015">Schülke & Smetana, 2015</bibRefCitation>
|
|
), the subgenus
|
|
<taxonomicName id="21BFCCB6FFD4B9640EB7F8EF96B5F8A4" authority="Zhou, 2017" authorityName="Zhou" authorityYear="2017" box="[398,482,1809,1832]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|
<emphasis id="D4CB6B27FFD4B9640EB7F8EF96B5F8A4" box="[398,482,1809,1832]" italics="true" pageId="1" pageNumber="294">Ocypus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="CFF8D65CFFD4B9640ED7F8EC9579F8A5" box="[494,558,1810,1833]" pageId="1" pageNumber="294" sensu="stricto">s. str.</taxonomicNameLabel>
|
|
includes the synonyms
|
|
<taxonomicName id="21BFCCB6FFD4B9640C76F8EF932EF8A5" authority="Westwood, 1827" authorityName="Westwood" authorityYear="1827" box="[847,1145,1809,1834]" class="Insecta" family="Staphylinidae" genus="Goerius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="D4CB6B27FFD4B9640C76F8EF94FEF8A4" box="[847,937,1809,1832]" italics="true" pageId="1" pageNumber="294">Goerius</emphasis>
|
|
<bibRefCitation id="822ECAC4FFD4B9640C8CF8EF932EF8A5" author="Westwood" box="[949,1145,1809,1834]" pageId="1" pageNumber="294" refString="Westwood, J. O. (1827) Observations upon Siagonium quadricorne of Kirby, and on other portions of the Brachelytra (Staphylinus Lin.). The Zoological Journal, 3, 56 - 66." type="journal article" year="1827">Westwood, 1827</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="21BFCCB6FFD4B9640BB0F8EC92CFF8A6" authority="Gistel, 1856" authorityName="Gistel" authorityYear="1856" box="[1161,1432,1809,1834]" class="Insecta" family="Staphylinidae" genus="Isopterum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="D4CB6B27FFD4B9640BB0F8EC93ADF8A5" box="[1161,1274,1810,1833]" italics="true" pageId="1" pageNumber="294">Isopterum</emphasis>
|
|
<bibRefCitation id="822ECAC4FFD4B9640A3FF8EF92CFF8A6" author="Gistel" box="[1286,1432,1809,1834]" pageId="1" pageNumber="294" refString="Gistel, J. N. F. X. (1856) Die Mysterien der europaischen Insectenwelt. Ein geheimer Schlussel fur Sammler aller Insecten- Ordnungen und Stande, behufs des Fangs, des Aufenhalts-Orts, der Wohnung, Tag- und Jahreszeit u. s. w., oder autoptische Darstellung des Insectenstaats in seinem Zusammenhange zum Bestehen des Naturhaushaltes uberhaupt und insbesondere in seinem Einflusse auf die phanerogamische und cryptogamische Pflanzenbevolkerung Europa's. Zum ersten Male nach funfundzwanzigjahrigen eigenen Erfahrungen zusammengestellt und herausgegeben. T. Dannheimer, Kempten, xii + 530 pp." type="book" year="1856">Gistel, 1856</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="21BFCCB6FFD4B9640FAEF8CB9690F8C1" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[151,455,1845,1870]" class="Insecta" family="Staphylinidae" genus="Nudabemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="D4CB6B27FFD4B9640FAEF8CB964CF8C1" box="[151,283,1845,1869]" italics="true" pageId="1" pageNumber="294">Nudabemus</emphasis>
|
|
<bibRefCitation id="822ECAC4FFD4B9640E1DF8CB9690F8C1" author="Coiffait" box="[292,455,1845,1870]" pageId="1" pageNumber="294" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
|
|
</taxonomicName>
|
|
, and
|
|
<emphasis id="D4CB6B27FFD4B9640D3FF8CB95C0F8C1" box="[518,663,1845,1869]" italics="true" pageId="1" pageNumber="294">Xanthocypus</emphasis>
|
|
<bibRefCitation id="822ECAC4FFD4B9640D99F8C89405F8C2" author="Muller" box="[672,850,1845,1870]" pageId="1" pageNumber="294" refString="Muller, J. (1925) Terzo contribute alia conoscenza del genere Staphylinus L .. Bollettino della Societa Entomologica Italiana, 57, 40 - 48." type="journal article" year="1925">J. Müller, 1925</bibRefCitation>
|
|
. The taxonomic position of the name
|
|
<emphasis id="D4CB6B27FFD4B9640A32F8CB92CBF8C1" box="[1291,1436,1845,1869]" italics="true" pageId="1" pageNumber="294">Xanthocypus</emphasis>
|
|
was once challenged if it should be maintained within the subgenus
|
|
<taxonomicName id="21BFCCB6FFD4B9640CA3F8A794B9F8FC" authority="Zhou, 2017" authorityName="Zhou" authorityYear="2017" box="[922,1006,1881,1904]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|
<emphasis id="D4CB6B27FFD4B9640CA3F8A794B9F8FC" box="[922,1006,1881,1904]" italics="true" pageId="1" pageNumber="294">Ocypus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="CFF8D65CFFD4B9640CCCF8A49366F8FD" box="[1013,1073,1882,1905]" pageId="1" pageNumber="294" sensu="stricto">s. str.</taxonomicNameLabel>
|
|
(
|
|
<bibRefCitation id="822ECAC4FFD4B9640B79F8A793BFF8FD" author="Hayashi" box="[1088,1256,1881,1906]" pageId="1" pageNumber="294" refString="Hayashi, Y. (1993) Studies on the Asian Staphylinidae, I, (Coleoptera, Staphylinidae). Elytra, 21 (2), 281 - 301." type="journal article" year="1993">Hayashi, 1993</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="822ECAC4FFD4B9640BCDF8A792CFF8FD" author="Smetana" box="[1268,1432,1881,1905]" pageId="1" pageNumber="294" refString="Smetana, A. (2007) Contributions to the knowledge of the " Staphylinus - complex " (Coleoptera, Staphylinidae, Staphylinini) of China. Part 16. The genus Ocypus Leach, 1819, subgenera Ocypus s. str. and Matidus Motschulsky, 1860. Elytra, 35, 441 - 459." type="journal article" year="2007">Smetana 2007</bibRefCitation>
|
|
: “tentatively set aside as a possible junior synonym of
|
|
<taxonomicName id="21BFCCB6FFD4B9640DC1F88394A5F819" authority="Sharp, 1889" authorityName="Sharp" authorityYear="1889" box="[760,1010,1917,1942]" class="Insecta" family="Staphylinidae" genus="Agelosus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="D4CB6B27FFD4B9640DC1F883940AF819" box="[760,861,1917,1941]" italics="true" pageId="1" pageNumber="294">Agelosus</emphasis>
|
|
<bibRefCitation id="822ECAC4FFD4B9640C5CF88394A5F819" author="Sharp" box="[869,1010,1917,1942]" pageId="1" pageNumber="294" refString="Sharp, D. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3, 28 - 44, 108 - 121, 249 - 267, 319 - 334, 406 - 419, 463 - 476. https: // doi. org / 10.1080 / 00222938909460371" type="journal article" year="1889">Sharp, 1889</bibRefCitation>
|
|
</taxonomicName>
|
|
”), but this was not confirmed by later publications (Smetana, 2009;
|
|
<bibRefCitation id="822ECAC4FFD4B9640EDAF85F945CF836" author="Schulke" box="[483,779,1953,1978]" pageId="1" pageNumber="294" refString="Schulke, M. & Smetana, A. (2015) Staphylinidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Fol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 901 - 1134." type="journal article" year="2015">Schülke & Smetana, 2015</bibRefCitation>
|
|
). We therefore maintained it within the subgenus
|
|
<taxonomicName id="21BFCCB6FFD4B9640A09F85F92D3F834" authority="Zhou, 2017" authorityName="Zhou" authorityYear="2017" box="[1328,1412,1953,1976]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|
<emphasis id="D4CB6B27FFD4B9640A09F85F92D3F834" box="[1328,1412,1953,1976]" italics="true" pageId="1" pageNumber="294">Ocypus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="CFF8D65CFFD4B9640AB3F85C97EEF851" pageId="1" pageNumber="294" sensu="stricto">s. str.</taxonomicNameLabel>
|
|
and included here two Chinese species:
|
|
<taxonomicName id="21BFCCB6FFD4B9640D46F83B94A5F851" authority="(Bernhauer, 1943)" baseAuthorityName="Bernhauer" baseAuthorityYear="1943" box="[639,1010,1989,2014]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="miwai">
|
|
<emphasis id="D4CB6B27FFD4B9640D46F83B9449F851" box="[639,798,1989,2013]" italics="true" pageId="1" pageNumber="294">Ocypus miwai</emphasis>
|
|
(
|
|
<bibRefCitation id="822ECAC4FFD4B9640C14F83B94BEF851" author="Bernhauer" box="[813,1001,1989,2014]" pageId="1" pageNumber="294" refString="Bernhauer, M. (1943) Neuheiten der palaearktischen Staphylinidenfauna. Mitteilungen der Munchner Entomologischen Gesellschaft, 33, 169 - 188." type="journal article" year="1943">Bernhauer, 1943</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="21BFCCB6FFD4B9640B11F83B9230F851" authority="Harold, 1877" authorityName="Harold" authorityYear="1877" box="[1064,1383,1989,2014]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="weisei">
|
|
<emphasis id="D4CB6B27FFD4B9640B11F83B939EF851" box="[1064,1225,1989,2013]" italics="true" pageId="1" pageNumber="294">Ocypus weisei</emphasis>
|
|
<bibRefCitation id="822ECAC4FFD4B9640BE9F83B9230F851" author="Harold" box="[1232,1383,1989,2014]" pageId="1" pageNumber="294" refString="Harold, E. (1877) Beitrage zur Kaferfauna von Japan. Deutsche Entomologische Zeitschrift, 21, 337 - 367. https: // doi. org / 10.1002 / mmnd. 4800210220" type="journal article" year="1877">Harold, 1877</bibRefCitation>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |