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<document id="5075668A935928255E5FA78C3841FEE4" ID-DOI="10.5281/zenodo.199681" ID-GBIF-Dataset="62e1623e-d85e-46ea-8edf-6cb01d7c88eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="199681" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460343126729" checkinUser="plazi" docAuthor="Blaimer, Bonnie B." docDate="2010" docId="6516CC460D0CFFEAFF7A8AB661EE623A" docLanguage="en" docName="zt02714p039.pdf" docOrigin="Zootaxa 2714" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Crematogaster (Decacrema)" docType="key" docVersion="8" lastPageNumber="14" masterDocId="992FB43E0D01FFE7FFED8A216475640D" masterDocTitle="Taxonomy and Natural History of the Crematogaster (Decacrema) - group (Hymenoptera: Formicidae) in Madagascar" masterLastPageNumber="39" masterPageNumber="1" pageNumber="14" updateTime="1698261518362" updateUser="plazi">
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<mods:title id="0375D26EDE054BDE4DDB7FD2536164D4">Taxonomy and Natural History of the Crematogaster (Decacrema) - group (Hymenoptera: Formicidae) in Madagascar</mods:title>
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<mods:namePart id="F4CD9905FDBD69625D64E486F8E050C4">Blaimer, Bonnie B.</mods:namePart>
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<mods:date id="B1B90947F21E7E4D6E31EDBC6A3EA707">2010</mods:date>
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<mods:number id="CF693396CD8447BA0511482CD2BD8071">2714</mods:number>
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<mods:identifier id="9F9BF9A180CD193BB959DA3E033885F6" type="DOI">10.5281/zenodo.199681</mods:identifier>
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<mods:identifier id="4F3926E0BD268467B2F0436EC91FAE5E" type="ISSN">1175-5326</mods:identifier>
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<treatment id="6516CC460D0CFFEAFF7A8AB661EE623A" ID-DOI="http://doi.org/10.5281/zenodo.6199368" ID-GBIF-Taxon="119406765" ID-Zenodo-Dep="6199368" LSID="urn:lsid:plazi:treatment:6516CC460D0CFFEAFF7A8AB661EE623A" httpUri="http://treatment.plazi.org/id/6516CC460D0CFFEAFF7A8AB661EE623A" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="A5A52EDB0D0CFFEAFF7A8AB6601464BC" box="[151,1121,151,177]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="ED007D500D0CFFEAFF7A8AB6601464BC" blockId="13.[151,1121,151,177]" box="[151,1121,151,177]" pageId="13" pageNumber="14">
<heading id="B648CA3C0D0CFFEAFF7A8AB6601464BC" bold="true" box="[151,1121,151,177]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="DFCBA1420D0CFFEAFF7A8AB6601464BC" bold="true" box="[151,1121,151,177]" pageId="13" pageNumber="14">
Key to the workers of the
<taxonomicName id="2ABF06D30D0CFFEAFE368AB6675B64BC" box="[475,814,151,177]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="10" phylum="Arthropoda" rank="subGenus" subGenus="Decacrema">
<emphasis id="DFCBA1420D0CFFEAFE368AB666F864BC" bold="true" box="[475,653,151,177]" italics="true" pageId="13" pageNumber="14">Crematogaster</emphasis>
(
<emphasis id="DFCBA1420D0CFFEAFD708AB6675364BC" bold="true" box="[669,806,151,177]" italics="true" pageId="13" pageNumber="14">Decacrema</emphasis>
)
</taxonomicName>
-group,
<collectingCountry id="95A83DC00D0CFFEAFC7F8AB6607E64BC" box="[914,1035,151,177]" name="Madagascar" pageId="13" pageNumber="14">Malagasy</collectingCountry>
region
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D0CFFEAFF7A8AC161EE623A" pageId="13" pageNumber="14" type="key">
<keyStep id="564B65F50D0CFFEAFF7A8AC161EE659A" pageId="13" pageNumber="14">
<paragraph id="ED007D500D0CFFEAFF7A8AC161EE653A" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A8AC161EE653A" pageId="13" pageNumber="14">
1. Petiole dorsally with postero-lateral tubercles, each bearing a single, stiff and long erect seta (
<figureCitation id="758461D50D0CFFEAFB5B8AC1617064FA" box="[1206,1285,224,247]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 21</figureCitation>
a); postpetiole distinctly bilobed and broadly medially impressed, with a pair of dorso-posterior stiff, long erect setae (
<figureCitation id="758461D50D0CFFEAFADD8B2161F5651A" box="[1328,1408,256,279]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 21</figureCitation>
b); abdominal tergites 4–7 usually with scattered erect pilosity; mandibles with 4 teeth .................................................. 2
</keyLead>
</paragraph>
<paragraph id="ED007D500D0CFFEAFF7A8B6161EE659A" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A8B6161EE659A" pageId="13" pageNumber="14">- Petiole dorsally lacking postero-lateral tubercle, setae commonly absent, if present (rare) then short and feeble; postpetiole variable, usually without erect setae; erect pilosity absent from abdominal tergites 4–7; mandibles with 4–5 teeth............................................................................................................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="564B65F50D0CFFEAFF7A8B8161EE665A" pageId="13" pageNumber="14">
<paragraph id="ED007D500D0CFFEAFF7A8B8161E965FA" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A8B8161E965FA" pageId="13" pageNumber="14">
2(1) Lateral portions of promesonotum raised with respect to median portion, and with postero-lateral carinate denticles and lateral carinae bordering metanotal groove, in lateral view promesonotum appearing dorsally flat (
<figureCitation id="758461D50D0CFFEAFAE38BE1612865DA" box="[1294,1373,448,471]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 22</figureCitation>
, or as in
<figureCitation id="758461D50D0CFFEAFF0F8BC1654565FA" box="[226,304,480,503]" captionStart="FIGURES 32 – 37" captionStartId="29.[151,270,1974,1998]" captionTargetBox="[190,1397,196,1939]" captionTargetId="figure@29.[179,1407,166,1953]" captionTargetPageId="29" captionText="FIGURES 32 – 37. Malagasy Crematogaster (Decacrema) species; a: full-face view, b: lateral view, c: dorsal view. 32: C. nosibeensis; 33: C. sisa; 34: C. sabatra; 35: C. mahery; 36: C. malala; 37: C. grevei." httpUri="https://zenodo.org/record/199685/files/figure.png" pageId="13" pageNumber="14">Fig. 37</figureCitation>
b);
<emphasis id="DFCBA1420D0CFFEAFEBF8BC167CD65FA" box="[338,952,480,503]" italics="true" pageId="13" pageNumber="14">widespread in western and southern dry and spiny forests</emphasis>
..........................................................
<taxonomicName id="2ABF06D30D0CFFEAFAD68BC161E965FA" ID-CoL="6BDC5" box="[1339,1436,480,503]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D0CFFEAFAD68BC161E965FA" box="[1339,1436,480,503]" italics="true" pageId="13" pageNumber="14">C. grevei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="ED007D500D0CFFEAFF7A882161EE665A" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A882161EE665A" pageId="13" pageNumber="14">
- Lateral portions of promesonotum not raised with respect to median portion, postero-laterally rounded or angular and metanotal groove without lateral carinae, in lateral view promesonotum forming a more or less convex outline (
<figureCitation id="758461D50D0CFFEAFF228861656B665A" box="[207,286,576,599]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 23</figureCitation>
, or as in
<figureCitation id="758461D50D0CFFEAFE93886165B9665A" box="[382,460,576,599]" captionStart="FIGURES 32 – 37" captionStartId="29.[151,270,1974,1998]" captionTargetBox="[190,1397,196,1939]" captionTargetId="figure@29.[179,1407,166,1953]" captionTargetPageId="29" captionText="FIGURES 32 – 37. Malagasy Crematogaster (Decacrema) species; a: full-face view, b: lateral view, c: dorsal view. 32: C. nosibeensis; 33: C. sisa; 34: C. sabatra; 35: C. mahery; 36: C. malala; 37: C. grevei." httpUri="https://zenodo.org/record/199685/files/figure.png" pageId="13" pageNumber="14">Fig. 36</figureCitation>
b);
<emphasis id="DFCBA1420D0CFFEAFE008861666F665A" box="[493,538,576,599]" italics="true" pageId="13" pageNumber="14">rare</emphasis>
,
<emphasis id="DFCBA1420D0CFFEAFDC5886161EE665A" box="[552,1435,576,599]" italics="true" pageId="13" pageNumber="14">
montane forests of southern central
<collectingCountry id="95A83DC00D0CFFEAFC4588616058665A" box="[936,1069,576,599]" name="Madagascar" pageId="13" pageNumber="14">Madagascar</collectingCountry>
......................................
<taxonomicName id="2ABF06D30D0CFFEAFADF886161EE665A" box="[1330,1435,576,599]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="malala">C. malala</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="564B65F50D0CFFEAFF7A884161EE66DA" pageId="13" pageNumber="14">
<paragraph id="ED007D500D0CFFEAFF7A884161E9669A" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A884161E9669A" pageId="13" pageNumber="14">
3(1) Head and promesonotal sculpture well pronounced, deeply costulate (
<figureCitation id="758461D50D0CFFEAFC428841678E667A" box="[943,1019,608,631]" captionStart="FIGURES 1 – 11" captionStartId="5.[151,271,1905,1929]" captionTargetBox="[223,1345,227,1856]" captionTargetId="figure@5.[223,1360,197,1878]" captionTargetPageId="5" captionText="FIGURES 1 – 11. Measurements and sculpture patterns. 1: head width, head length and eye length; 2: scape length; 3: length of hind tibia; 4: Weber’s length and spine length; 5: petiole length and petiole height; 6: petiole width, postpetiole width and postpetiole height; 7: mesonotal width and mesonotal length; 8: aciculate, 9: areolate, 10: reticulate, 11: costulate sculpture." httpUri="https://zenodo.org/record/199682/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
and
<figureCitation id="758461D50D0CFFEAFBC28841603E667A" box="[1071,1099,608,631]" captionStart="FIGURES 44 – 48" captionStartId="31.[151,269,1707,1731]" captionTargetBox="[189,1398,208,1677]" captionTargetId="figure@31.[181,1406,194,1685]" captionTargetPageId="31" captionText="FIGURES 44 – 48. The Crematogaster (Decacrema) hova - complex; a: full-face view, b: lateral view, c: dorsal view. 44: morphotype 1; 45: morphotype 2; 46: morphotype 3; 47: morphotype 4; 48: morphotype 5." httpUri="https://zenodo.org/record/199687/files/figure.png" pageId="13" pageNumber="14">47</figureCitation>
a) or areolate (
<figureCitation id="758461D50D0CFFEAFB0988416156667A" box="[1252,1315,608,631]" captionStart="FIGURES 1 – 11" captionStartId="5.[151,271,1905,1929]" captionTargetBox="[223,1345,227,1856]" captionTargetId="figure@5.[223,1360,197,1878]" captionTargetPageId="5" captionText="FIGURES 1 – 11. Measurements and sculpture patterns. 1: head width, head length and eye length; 2: scape length; 3: length of hind tibia; 4: Weber’s length and spine length; 5: petiole length and petiole height; 6: petiole width, postpetiole width and postpetiole height; 7: mesonotal width and mesonotal length; 8: aciculate, 9: areolate, 10: reticulate, 11: costulate sculpture." httpUri="https://zenodo.org/record/199682/files/figure.png" pageId="13" pageNumber="14">Fig. 9</figureCitation>
and
<figureCitation id="758461D50D0CFFEAFABA88416106667A" box="[1367,1395,608,631]" captionStart="FIGURES 44 – 48" captionStartId="31.[151,269,1707,1731]" captionTargetBox="[189,1398,208,1677]" captionTargetId="figure@31.[181,1406,194,1685]" captionTargetPageId="31" captionText="FIGURES 44 – 48. The Crematogaster (Decacrema) hova - complex; a: full-face view, b: lateral view, c: dorsal view. 44: morphotype 1; 45: morphotype 2; 46: morphotype 3; 47: morphotype 4; 48: morphotype 5." httpUri="https://zenodo.org/record/199687/files/figure.png" pageId="13" pageNumber="14">48</figureCitation>
a) ..
<emphasis id="DFCBA1420D0CFFEAFF2388A167AD669A" box="[206,984,640,663]" italics="true" pageId="13" pageNumber="14">.......................................................................................................C. hova-</emphasis>
complex (morphotypes 4, 5, and part of 3)
</keyLead>
</paragraph>
<paragraph id="ED007D500D0CFFEAFF7A888161EE66DA" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A888161EE66DA" pageId="13" pageNumber="14">
- Head and promesonotal sculpture less developed, from reduced-aciculate (
<figureCitation id="758461D50D0CFFEAFC088881605366BA" box="[997,1062,672,695]" captionStart="FIGURES 1 – 11" captionStartId="5.[151,271,1905,1929]" captionTargetBox="[223,1345,227,1856]" captionTargetId="figure@5.[223,1360,197,1878]" captionTargetPageId="5" captionText="FIGURES 1 – 11. Measurements and sculpture patterns. 1: head width, head length and eye length; 2: scape length; 3: length of hind tibia; 4: Weber’s length and spine length; 5: petiole length and petiole height; 6: petiole width, postpetiole width and postpetiole height; 7: mesonotal width and mesonotal length; 8: aciculate, 9: areolate, 10: reticulate, 11: costulate sculpture." httpUri="https://zenodo.org/record/199682/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
, or e.g. 44a) to reticulate (
<figureCitation id="758461D50D0CFFEAFAAD888161FA66BA" box="[1344,1423,672,695]" captionStart="FIGURES 1 – 11" captionStartId="5.[151,271,1905,1929]" captionTargetBox="[223,1345,227,1856]" captionTargetId="figure@5.[223,1360,197,1878]" captionTargetPageId="5" captionText="FIGURES 1 – 11. Measurements and sculpture patterns. 1: head width, head length and eye length; 2: scape length; 3: length of hind tibia; 4: Weber’s length and spine length; 5: petiole length and petiole height; 6: petiole width, postpetiole width and postpetiole height; 7: mesonotal width and mesonotal length; 8: aciculate, 9: areolate, 10: reticulate, 11: costulate sculpture." httpUri="https://zenodo.org/record/199682/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
), reticulate-areolate (e.g.
<figureCitation id="758461D50D0CFFEAFE2988E1666466DA" box="[452,529,704,727]" captionStart="FIGURES 44 – 48" captionStartId="31.[151,269,1707,1731]" captionTargetBox="[189,1398,208,1677]" captionTargetId="figure@31.[181,1406,194,1685]" captionTargetPageId="31" captionText="FIGURES 44 – 48. The Crematogaster (Decacrema) hova - complex; a: full-face view, b: lateral view, c: dorsal view. 44: morphotype 1; 45: morphotype 2; 46: morphotype 3; 47: morphotype 4; 48: morphotype 5." httpUri="https://zenodo.org/record/199687/files/figure.png" pageId="13" pageNumber="14">Fig. 46</figureCitation>
a) to weakly costulate .................................................................................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="564B65F50D0CFFEAFF7A88C161EE67BA" pageId="13" pageNumber="14">
<paragraph id="ED007D500D0CFFEAFF7A88C161E9675A" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A88C161E9675A" pageId="13" pageNumber="14">
4(3) Face lacking erect setae except a single pair near frontal carinae; subpetiolar process developed as broad protuberance (
<figureCitation id="758461D50D0CFFEAFEE589216523671A" box="[264,342,768,791]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 25</figureCitation>
and
<figureCitation id="758461D50D0CFFEAFE60892165DF671A" box="[397,426,768,791]" captionStart="FIGURES 38 – 43" captionStartId="30.[151,270,1894,1918]" captionTargetBox="[221,1364,193,1874]" captionTargetId="figure@30.[200,1385,164,1882]" captionTargetPageId="30" captionText="FIGURES 38 – 43. Malagasy Crematogaster (Decacrema) species; a: petiole &amp; postpetiole in dorsal view, b: petiole &amp; postpetiole in lateral view. 38: C. nosibeensis; 39: C. sisa, 40: C. sabatra; 41: C. mahery; 42: C. malala; 43: C. grevei." httpUri="https://zenodo.org/record/199686/files/figure.png" pageId="13" pageNumber="14">38</figureCitation>
b); small to medium size (HW 0.86–1.04, WL 0.80–1.05); propodeal spines small to medium size (SPI 0.15–0.20); sculpture on head and promesonotum always reduced-aciculate;
<emphasis id="DFCBA1420D0CFFEAFB8989016642675A" italics="true" pageId="13" pageNumber="14">
northern, northwestern
<collectingCountry id="95A83DC00D0CFFEAFA8F89016568675A" name="Madagascar" pageId="13" pageNumber="14">Madagascar</collectingCountry>
and Ambohitantely forest.
</emphasis>
..........................................................................................................
<taxonomicName id="2ABF06D30D0CFFEAFAEF896161E9675A" box="[1282,1436,832,855]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="nosibeensis">
<emphasis id="DFCBA1420D0CFFEAFAEF896161E9675A" box="[1282,1436,832,855]" italics="true" pageId="13" pageNumber="14">C. nosibeensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="ED007D500D0CFFEAFF7A894161EE67BA" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A894161EE67BA" pageId="13" pageNumber="14">- Face with additional erect setae (careful, could be abraded) other than those close to frontal carinae; subpetiolar process variable; size highly variable, small to very large (HW 0.74–1.31, WL 0.78–1.34); propodeal spines variable (SPI 0.17–0.40), but commonly SPI&gt; 0.20; sculpture variable. .................................................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="564B65F50D0CFFEAFF7A89E161EE60BA" pageId="13" pageNumber="14">
<paragraph id="ED007D500D0CFFEAFF7A89E161E9603A" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A89E161E9603A" pageId="13" pageNumber="14">
5(4) Subpetiolar process developed as acute, long tooth (
<figureCitation id="758461D50D0CFFEAFCF789E1671967DA" box="[794,876,960,983]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 24</figureCitation>
and
<figureCitation id="758461D50D0CFFEAFC4789E167B267DA" box="[938,967,960,983]" captionStart="FIGURES 38 – 43" captionStartId="30.[151,270,1894,1918]" captionTargetBox="[221,1364,193,1874]" captionTargetId="figure@30.[200,1385,164,1882]" captionTargetPageId="30" captionText="FIGURES 38 – 43. Malagasy Crematogaster (Decacrema) species; a: petiole &amp; postpetiole in dorsal view, b: petiole &amp; postpetiole in lateral view. 38: C. nosibeensis; 39: C. sisa, 40: C. sabatra; 41: C. mahery; 42: C. malala; 43: C. grevei." httpUri="https://zenodo.org/record/199686/files/figure.png" pageId="13" pageNumber="14">39</figureCitation>
b); promesonotal suture usually complete; medium to large size (HW 0.89–1.14, WL 0.91–1.12); propodeal spines small to medium size (SPI 0.17–0.21); head and promesonotal sculpture always reduced-aciculate;
<emphasis id="DFCBA1420D0CFFEAFCE28E2161E9603A" italics="true" pageId="13" pageNumber="14">
rare, known only from montane forests in R. S. Manongarivo .............................................................................................................................................................................
<taxonomicName id="2ABF06D30D0CFFEAFABE8E0161E9603A" box="[1363,1436,1056,1079]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sisa">C. sisa</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="ED007D500D0CFFEAFF7A8E6161EE60BA" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A8E6161EE60BA" pageId="13" pageNumber="14">
- Subpetiolar process variable, moderately articulated as angular dent (
<figureCitation id="758461D50D0CFFEAFC5D8E61678A605A" box="[944,1023,1088,1111]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 26</figureCitation>
,
<figureCitation id="758461D50D0CFFEAFBE68E616052605A" box="[1035,1063,1088,1111]" captionStart="FIGURES 38 – 43" captionStartId="30.[151,270,1894,1918]" captionTargetBox="[221,1364,193,1874]" captionTargetId="figure@30.[200,1385,164,1882]" captionTargetPageId="30" captionText="FIGURES 38 – 43. Malagasy Crematogaster (Decacrema) species; a: petiole &amp; postpetiole in dorsal view, b: petiole &amp; postpetiole in lateral view. 38: C. nosibeensis; 39: C. sisa, 40: C. sabatra; 41: C. mahery; 42: C. malala; 43: C. grevei." httpUri="https://zenodo.org/record/199686/files/figure.png" pageId="13" pageNumber="14">41</figureCitation>
b, 49b, 50b, 52b and 53b), close to absent (
<figureCitation id="758461D50D0CFFEAFEF48E416510607A" box="[281,357,1120,1143]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 27</figureCitation>
and
<figureCitation id="758461D50D0CFFEAFE758E4165C1607A" box="[408,436,1120,1143]" captionStart="FIGURES 49 – 53" captionStartId="32.[151,269,1818,1842]" captionTargetBox="[190,1397,201,1792]" captionTargetId="figure@32.[182,1404,194,1797]" captionTargetPageId="32" captionText="FIGURES 49 – 53. The Crematogaster (Decacrema) hova - complex; a: petiole &amp; postpetiole in dorsal view, b: petiole &amp; postpetiole in lateral view. 49: morphotype 1; 50: morphotype 2; 51: morphotype 3; 52: morphotype 4; 53: morphotype 5." httpUri="https://zenodo.org/record/199688/files/figure.png" pageId="13" pageNumber="14">51</figureCitation>
b), or rarely as a broad protuberance (
<figureCitation id="758461D50D0CFFEAFCAC8E4167E5607A" box="[833,912,1120,1143]" captionStart="FIGURES 38 – 43" captionStartId="30.[151,270,1894,1918]" captionTargetBox="[221,1364,193,1874]" captionTargetId="figure@30.[200,1385,164,1882]" captionTargetPageId="30" captionText="FIGURES 38 – 43. Malagasy Crematogaster (Decacrema) species; a: petiole &amp; postpetiole in dorsal view, b: petiole &amp; postpetiole in lateral view. 38: C. nosibeensis; 39: C. sisa, 40: C. sabatra; 41: C. mahery; 42: C. malala; 43: C. grevei." httpUri="https://zenodo.org/record/199686/files/figure.png" pageId="13" pageNumber="14">Fig. 40</figureCitation>
b); promesonotal suture often incomplete, if complete, then large to very large species (HW 1.09–1.21, WL 1.18–1.32); otherwise size highly variable; propodeal spines usually with SPI&gt; 0.20; sculpture variable. ...................................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="564B65F50D0CFFEAFF7A8EE161E9619A" pageId="13" pageNumber="14">
<paragraph id="ED007D500D0CFFEAFF7A8EE161EE611A" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A8EE161EE611A" pageId="13" pageNumber="14">
6(5) Mesonotum postero-laterally distinctly angular to tuberculate (
<figureCitation id="758461D50D0CFFEAFC9F8EE167B460DA" box="[882,961,1216,1239]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 28</figureCitation>
b and 30b, or as in 32b–35b); large to very large species (HW 1.09–1.31, WL 1.18–1.34); head usually reduced-aciculate, sometimes reticulate to feebly costulate;
<emphasis id="DFCBA1420D0CFFEAFF168F2167B5611A" box="[251,960,1280,1303]" italics="true" pageId="13" pageNumber="14">
uncommon, central, eastern central and southeastern
<collectingCountry id="95A83DC00D0CFFEAFCD68F2167B5611A" box="[827,960,1280,1303]" name="Madagascar" pageId="13" pageNumber="14">Madagascar</collectingCountry>
</emphasis>
.
<emphasis id="DFCBA1420D0CFFEAFC218F2161FA611A" box="[972,1423,1280,1303]" italics="true" pageId="13" pageNumber="14">...................................................................</emphasis>
7
</keyLead>
</paragraph>
<paragraph id="ED007D500D0CFFEAFF7A8F0161E9619A" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A8F0161E9619A" pageId="13" pageNumber="14">
- Mesonotum postero-laterally rounded or slightly tuberculate (
<figureCitation id="758461D50D0CFFEAFCB18F0167DE613A" box="[860,939,1312,1335]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 29</figureCitation>
, and as in 44b–48b),
<emphasis id="DFCBA1420D0CFFEAFB7C8F0160A7613A" box="[1169,1234,1312,1335]" italics="true" pageId="13" pageNumber="14">rarely</emphasis>
angular; body size small to large (HW 0.74–1.19, WL 0.78–1.22); head sculpture variable, aciculate, reticulate or reticulate-areolate, if head sculpture reduced and aciculate, then usually at least promesonotal sculpture more developed;
<emphasis id="DFCBA1420D0CFFEAFACF8F4167AD619A" italics="true" pageId="13" pageNumber="14">
widespread throughout humid forests in
<collectingCountry id="95A83DC00D0CFFEAFE198FA1660E619A" box="[500,635,1408,1431]" name="Madagascar" pageId="13" pageNumber="14">Madagascar</collectingCountry>
......................................C. hova-
</emphasis>
complex (morphotypes 1, 2, and part of 3)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="564B65F50D0CFFEAFF7A8F8161EE623A" pageId="13" pageNumber="14">
<paragraph id="ED007D500D0CFFEAFF7A8F8161E961DA" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A8F8161E961DA" pageId="13" pageNumber="14">
7(6) Mesonotum raised with respect to pronotum and medially at least slightly tuberculate (
<figureCitation id="758461D50D0CFFEAFB798F81609261BA" box="[1172,1255,1440,1463]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 30</figureCitation>
a); promesonotal suture usually complete (
<figureCitation id="758461D50D0CFFEAFE3E8FE1665461DA" box="[467,545,1472,1495]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 31</figureCitation>
a); head and promesonotal sculpture
<emphasis id="DFCBA1420D0CFFEAFC458FE1678761DA" box="[936,1010,1472,1495]" italics="true" pageId="13" pageNumber="14">always</emphasis>
aciculate.
<emphasis id="DFCBA1420D0CFFEAFBB28FE161E961DA" box="[1119,1436,1472,1495]" italics="true" pageId="13" pageNumber="14">
..............................
<taxonomicName id="2ABF06D30D0CFFEAFAC08FE161E961DA" box="[1325,1436,1472,1495]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mahery">C. mahery</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="ED007D500D0CFFEAFF7A8FC161EE623A" blockId="13.[151,1436,224,1591]" pageId="13" pageNumber="14">
<keyLead id="564EDE650D0CFFEAFF7A8FC161EE623A" pageId="13" pageNumber="14">
- Mesonotum usually not raised with respect to pronotum nor medially tuberculate (
<figureCitation id="758461D50D0CFFEAFBA38FC160EA61FA" box="[1102,1183,1504,1527]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 28</figureCitation>
a), promesonotal suture usually incomplete (
<figureCitation id="758461D50D0CFFEAFE4D8C216599621A" box="[416,492,1536,1559]" captionStart="FIGURES 21 – 31" captionStartId="12.[151,270,1851,1875]" captionTargetBox="[260,1337,246,1806]" captionTargetId="figure@12.[244,1353,194,1856]" captionTargetPageId="12" captionText="FIGURES 21 – 31. Key to Malagasy Decacrema. 21 a: petiole with erect setae; 21 b: postpetiole distinctly bilobed and with erect setae; 22: promesonotum in lateral view flat; 23: promesonotum in lateral view convex; 24: subpetiolar process as acute tooth; 25: subpetiolar process as broad protuberance; 26: subpetiolar process as angular dent; 27: subpetiolar process absent; 28: mesonotum a: not raised in respect to pronotum b: postero-laterally angular, 29: mesonotum rounded postero-laterally; 30: mesonotum a: raised in respect to pronotum b: postero-laterally angular; 31: promesonotal suture a: complete b: incomplete." httpUri="https://zenodo.org/record/199684/files/figure.png" pageId="13" pageNumber="14">Fig. 31</figureCitation>
b) and only laterally impressed; head sculpture and promesonotal sculpture either aciculate or reticulate to weakly costulate............................................................................................................
<taxonomicName id="2ABF06D30D0CFFEAFAF38C0161FB623A" box="[1310,1422,1568,1591]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sabatra">
<emphasis id="DFCBA1420D0CFFEAFAF38C0161FB623A" box="[1310,1422,1568,1591]" italics="true" pageId="13" pageNumber="14">C. sabatra</emphasis>
</taxonomicName>
*
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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