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<document id="05A33A54C551E09CC8F83341848A2D49" ID-CLB-Dataset="297340" ID-DOI="10.5281/zenodo.12761936" ID-GBIF-Dataset="5e6c8fd8-74a8-4de2-8687-29b4ea71fac1" ID-ISSN="1525-9153" ID-Zenodo-Dep="12761936" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1714488167562" checkinUser="felipe" docAuthor="Conradie, Werner, Keates, Chad, Verburgt, Luke, Baptista, Ninda L. &amp; Harvey, James" docDate="2023" docId="43578788002C3F4088A1606AFC1A0283" docLanguage="en" docName="AmphibReptileCons.17.1-2.e325.pdf" docOrigin="Amphibian &amp; Reptile Conservation (e 325) 17 (1)" docStyle="DocumentStyle:D97478CDCB68366E5D4AECD282BFFB6F.2:AmphibReptileConse.2015-.journal_article.0cover" docStyleId="D97478CDCB68366E5D4AECD282BFFB6F" docStyleName="AmphibReptileConse.2015-.journal_article.0cover" docStyleVersion="2" docTitle="Hyperolius bocagei Steindachner 1867" docType="treatment" docVersion="6" lastPageNumber="36" masterDocId="BF6EFFF0003D3F52882C642AFFC80606" masterDocTitle="Contributions to the herpetofauna of the Angolan Okavango- Cuando-Zambezi river drainages. Part 3: Amphibians" masterLastPageNumber="56" masterPageNumber="19" pageNumber="36" updateTime="1721238984318" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0F7BBE95666D051EF7225C8E9007CEED">Contributions to the herpetofauna of the Angolan Okavango- Cuando-Zambezi river drainages. Part 3: Amphibians</mods:title>
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<mods:name id="574AB48082DEA8483F5C843060FC3E36" type="personal">
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<mods:namePart id="21B734C4B5F8D89689889E3F487B16CD">Conradie, Werner</mods:namePart>
<mods:affiliation id="8EA1D16374869478E2FF050C5B8041D3">National Geographic Okavango Wilderness Project, Wild Bird Trust, SOUTH AFRICA &amp; Department of Nature Conservation Management, Natural Resource Science and Management Cluster, Faculty of Science, George Campus, Nelson Mandela University, George, SOUTH AFRICA &amp; Port Elizabeth Museum, Beach Road, Humewood, Port Elizabeth 6013, SOUTH AFRICA</mods:affiliation>
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<mods:namePart id="03BC1CF130E2EACE3A94C4AC758F73A4">Keates, Chad</mods:namePart>
<mods:affiliation id="C429A38FF538B7DE3C4D7FA8B808500F">South African Institute for Aquatic Biodiversity (SAIAB), Makhanda, SOUTH AFRICA &amp; National Geographic Okavango Wilderness Project, Wild Bird Trust, SOUTH AFRICA &amp; Department of Nature Conservation Management, Natural Resource Science and Management Cluster, Faculty of Science, George Campus, Nelson Mandela University, George, SOUTH AFRICA</mods:affiliation>
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<mods:namePart id="830AB4EDDF50A01F4356EC1FC35C6427">Verburgt, Luke</mods:namePart>
<mods:affiliation id="34C9776DAEF2C0086FAA0168B2545BB7">Department of Zoology and Entomology, University of Pretoria, Pretoria, 0001, SOUTH AFRICA &amp; Enviro-Insight CC, Unit 8 Oppidraai Office Park, Pretoria 0050, SOUTH AFRICA &amp; National Geographic Okavango Wilderness Project, Wild Bird Trust, SOUTH AFRICA</mods:affiliation>
</mods:name>
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<mods:namePart id="523B0D4079442D6F52ED7398B12CF027">Baptista, Ninda L.</mods:namePart>
<mods:affiliation id="10F07843EB138C623AC69B15279E22B8">Museum für Naturkunde-Leibniz Institute for Evolution and Biodiversity Science, Invalidenstr. 43, 10115 Berlin, GERMANY &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, PORTUGAL &amp; Departamento de Biologia, Faculdade de Ciências, Universidade do Porto, 4169 - 007, Porto, PORTUGAL &amp; CIBIO / InBio, Centro de Investigação em Biodiversidade e Recursos Genéticos, Laboratório Associado, Universidade do Porto, Campus Agrário de Vairão, Rua Padre Armando Quintas, 4485 - 661 Vairão, PORTUGAL &amp; Instituto Superior de Ciências da Educação da Huíla (ISCED), Rua Sarmento Rodrigues, Lubango, ANGOLA &amp; National Geographic Okavango Wilderness Project, Wild Bird Trust, SOUTH AFRICA</mods:affiliation>
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<mods:namePart id="B2B22704C857AC113D2FD4747EA508D5">Harvey, James</mods:namePart>
<mods:affiliation id="66541D9F59C51AA3F2EDAAE9D593625D">Harvey Ecological, 41 Devonshire Avenue, Howick, 3290, SOUTH AFRICA &amp; National Geographic Okavango Wilderness Project, Wild Bird Trust, SOUTH AFRICA</mods:affiliation>
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<mods:title id="963841419A5A9129DAF4F7BFED4FE05F">Amphibian &amp; Reptile Conservation</mods:title>
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<mods:date id="BCE8A892CF8D0AD3CC785DC2B13B062E">2023</mods:date>
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<mods:title id="1177A071665481FA11B195EAB9EB72D9">e 325</mods:title>
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<mods:number id="2FBBDFCD19070DE77D72A9DEFBA30852">2023-08-04</mods:number>
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<mods:number id="B3C0EB4ED7805DDF7F1B6910610839E8">17</mods:number>
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<mods:start id="4385D6346BE5FED10CF3C4351C944F40">19</mods:start>
<mods:end id="2FF3CC58B400666CBEFB49B4909E1D17">56</mods:end>
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<mods:identifier id="56844994F086C4DCCE0C76E4DFDE8F37" type="ISSN">1525-9153</mods:identifier>
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<treatment id="43578788002C3F4088A1606AFC1A0283" ID-DOI="http://doi.org/10.5281/zenodo.11370902" ID-GBIF-Taxon="231569757" ID-Zenodo-Dep="11370902" LSID="urn:lsid:plazi:treatment:43578788002C3F4088A1606AFC1A0283" httpUri="http://treatment.plazi.org/id/43578788002C3F4088A1606AFC1A0283" lastPageId="18" lastPageNumber="36" pageId="17" pageNumber="36">
<subSubSection id="83E46515002C3F4388A1606AFD94027E" pageId="17" pageNumber="36" type="nomenclature">
<paragraph id="CB41369E002C3F4388A1606AFD94025E" blockId="17.[141,746,1088,1432]" box="[141,604,1088,1112]" pageId="17" pageNumber="36">
<heading id="900981F2002C3F4388A1606AFD94025E" box="[141,604,1088,1112]" fontSize="10" level="2" pageId="17" pageNumber="36" reason="4">
<taxonomicName id="0CFE4D1D002C3F4388A1606AFD94025E" ID-CoL="3NQRM" authority="Steindachner, 1867" authorityName="Steindachner" authorityYear="1867" box="[141,604,1088,1112]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" isUncertain="true" kingdom="Animalia" order="Anura" pageId="17" pageNumber="36" phylum="Chordata" rank="species" species="bocagei">
<emphasis id="F98AEA8C002C3F4388A1606AFECB025E" box="[141,259,1088,1112]" italics="true" pageId="17" pageNumber="36">Hyperolius</emphasis>
aff.
<emphasis id="F98AEA8C002C3F438918606AFE40025E" box="[308,392,1088,1112]" italics="true" pageId="17" pageNumber="36">bocagei</emphasis>
Steindachner, 1867
</taxonomicName>
</heading>
</paragraph>
<paragraph id="CB41369E002C3F4388A1604AFD94027E" blockId="17.[141,746,1088,1432]" box="[141,604,1120,1145]" pageId="17" pageNumber="36">
Bocage’s Reed Frog (
<figureCitation id="53C52A1B002C3F438958604AFE3E027E" box="[372,502,1120,1144]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="17.[791,826,1018,1040]" captionStartId-1="17.[791,826,1493,1515]" captionStartId-2="17.[789,824,1900,1922]" captionTargetBox-0="[791,1395,607,1010]" captionTargetBox-1="[791,1395,1083,1483]" captionTargetBox-2="[791,1395,1559,1890]" captionTargetId-0="figure-251@17.[791,1395,604,1012]" captionTargetId-1="figure-253@17.[791,1395,1081,1486]" captionTargetId-2="figure-255@17.[791,1395,1556,1893]" captionTargetPageId-0="17" captionTargetPageId-1="17" captionTargetPageId-2="17" captionText-0="Fig. 19. Adult male Hyperolius aff. bocagei (green form) from Cuito River source. Photo by Werner Conradie." captionText-1="Fig. 20. Adult male Hyperolius aff. bocagei (red form) from Cuanavale River source. Photo by Werner Conradie." captionText-2="Fig. 21. Adult male Hyperolius aff. bocagei (brown form) from Cuito River source. Photo by Werner Conradie." figureDoi-0="http://doi.org/10.5281/zenodo.12761982" figureDoi-1="http://doi.org/10.5281/zenodo.12761984" figureDoi-2="http://doi.org/10.5281/zenodo.12761986" httpUri-0="https://zenodo.org/record/12761982/files/figure.png" httpUri-1="https://zenodo.org/record/12761984/files/figure.png" httpUri-2="https://zenodo.org/record/12761986/files/figure.png" pageId="17" pageNumber="36">Figs. 19–21</figureCitation>
;
<figureCitation id="53C52A1B002C3F438A2D604BFD9C027E" box="[513,596,1120,1144]" captionStart="Map 18" captionStartId="17.[140,188,1919,1940]" captionTargetBox="[206,709,1467,1894]" captionTargetId="figure-240@17.[163,728,1439,1921]" captionTargetPageId="17" captionText="Map 18. Distribution of Hyperolius aff. bocagei in Angola." figureDoi="http://doi.org/10.5281/zenodo.12761988" httpUri="https://zenodo.org/record/12761988/files/figure.png" pageId="17" pageNumber="36">Map 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="83E46515002C3F4088A160AAFC1A0283" lastPageId="18" lastPageNumber="37" pageId="17" pageNumber="36" type="materials_examined">
<paragraph id="CB41369E002C3F4088A160AAFC1A0283" blockId="17.[141,746,1088,1432]" lastBlockId="18.[791,1396,141,1158]" lastPageId="18" lastPageNumber="37" pageId="17" pageNumber="36">
<emphasis id="F98AEA8C002C3F4388A160AAFD86029E" bold="true" box="[141,590,1152,1176]" pageId="17" pageNumber="36">
Material (
<specimenCount id="DDF8FD17002C3F4388D160AAFE5A029E" box="[253,402,1152,1176]" count="74" pageId="17" pageNumber="36" type="generic">74 specimens</specimenCount>
, 10 tadpole lots):
</emphasis>
<collectionCode id="ADEFAE5B002C3F438A7F60ABFD42029E" box="[595,650,1153,1176]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="17" pageNumber="36" type="Museum">PEM</collectionCode>
A12423, Cunde waterfall, -13.77390° 18.75520°,
<quantity id="0C069B7B002C3F438A75608AFD7002BE" box="[601,696,1184,1208]" metricMagnitude="3" metricUnit="m" metricValue="1.287" pageId="17" pageNumber="36" unit="m" value="1287.0">1,287 m</quantity>
asl;
<collectionCode id="ADEFAE5B002C3F4388A160EBFF0C02DE" box="[141,196,1217,1240]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="17" pageNumber="36" type="Museum">PEM</collectionCode>
A12455–60,
<collectionCode id="ADEFAE5B002C3F43897860EBFE4302DE" box="[340,395,1217,1240]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="17" pageNumber="36" type="Museum">PEM</collectionCode>
A14085–6 (tadpoles) Cuanavale River source lake, -13.08537° 18.89098°,
<quantity id="0C069B7B002C3F438AAB60CAFD2202FE" box="[647,746,1248,1272]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="17" pageNumber="36" unit="m" value="1340.0">1,340 m</quantity>
asl;
<collectionCode id="ADEFAE5B002C3F4388ED612BFF30031E" box="[193,248,1281,1304]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="17" pageNumber="36" type="Museum">PEM</collectionCode>
A12489,
<collectionCode id="ADEFAE5B002C3F438943612BFE6E031E" box="[367,422,1281,1304]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="17" pageNumber="36" type="Museum">PEM</collectionCode>
A14098 (tadpoles),
<collectionCode id="ADEFAE5B002C3F438ABA612AFD21031E" box="[662,745,1280,1304]" pageId="17" pageNumber="36">INBAC</collectionCode>
(no number), Dala River, near Samanunga village, -12.93169° 18.81458°,
<quantity id="0C069B7B002C3F4389A3616AFE23035E" box="[399,491,1344,1368]" metricMagnitude="3" metricUnit="m" metricValue="1.363" pageId="17" pageNumber="36" unit="m" value="1363.0">1,363 m</quantity>
asl;
<collectionCode id="ADEFAE5B002C3F438A09616BFD94035E" box="[549,604,1345,1368]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="17" pageNumber="36" type="Museum">PEM</collectionCode>
A12508–12, river crossing before Samboano village, -12.30700° 18.62350°,
<quantity id="0C069B7B002C3F43893F61AAFEBD039E" box="[275,373,1408,1432]" metricMagnitude="3" metricUnit="m" metricValue="1.3969999999999998" pageId="17" pageNumber="36" unit="m" value="1397.0">1,397 m</quantity>
asl;
<collectionCode id="ADEFAE5B002C3F43899761ABFE3A039E" box="[443,498,1409,1432]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="17" pageNumber="36" type="Museum">PEM</collectionCode>
A12543–52,
<collectionCode id="ADEFAE5B002C3F438ABA61AAFD21039E" box="[662,745,1408,1432]" pageId="17" pageNumber="36">INBAC</collectionCode>
(no number x2), Cuito River source lake, -12.68935° 18.36012°,
<quantity id="0C069B7B002F3F4089216484FEA106C3" box="[269,361,174,198]" metricMagnitude="3" metricUnit="m" metricValue="1.4309999999999998" pageId="18" pageNumber="37" unit="m" value="1431.0">1,431 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F40898F6484FE1206C3" box="[419,474,174,197]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A12592–8, Calua River source lake,
<quantity id="0C069B7B002F3F40893964E4FE8406E0" box="[277,332,206,230]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="37" unit="km" value="6.0">6 km</quantity>
SE of Cuito River source, -12.73675° 18.39310°,
<quantity id="0C069B7B002F3F40892064C4FEAF0703" box="[268,359,238,262]" metricMagnitude="3" metricUnit="m" metricValue="1.446" pageId="18" pageNumber="37" unit="m" value="1446.0">1,446 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F4089B264C4FE1D0703" box="[414,469,238,261]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A12619, Stop 1: road to Cuito River source, -12.25050° 18.63730°,
<quantity id="0C069B7B002F3F408A486527FD750723" box="[612,701,269,293]" metricMagnitude="3" metricUnit="m" metricValue="1.5559999999999998" pageId="18" pageNumber="37" unit="m" value="1556.0">1,556 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F4088A16504FF0C0743" box="[141,196,302,325]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A12674, Camp 3, Malova Village, Mipanha River, -14.09140° 16.41476°,
<quantity id="0C069B7B002F3F4089B86567FE3A0763" box="[404,498,333,357]" metricMagnitude="3" metricUnit="m" metricValue="1.553" pageId="18" pageNumber="37" unit="m" value="1553.0">1,553 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A1C6564FDAF0763" box="[560,615,334,357]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A12734–7,
<collectionCode id="ADEFAE5B002F3F4088A16544FF2C0780" box="[141,228,366,390]" pageId="18" pageNumber="37">INBAC</collectionCode>
: WC-4833,
<collectionCode id="ADEFAE5B002F3F40895D6544FE0C0780" box="[369,452,366,390]" pageId="18" pageNumber="37">INBAC</collectionCode>
(no number x2), Cuando River source, -13.00346° 19.12751°,
<quantity id="0C069B7B002F3F408A0F65A7FDB507A3" box="[547,637,397,421]" metricMagnitude="3" metricUnit="m" metricValue="1.553" pageId="18" pageNumber="37" unit="m" value="1553.0">1,553 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A9F65A4FD2207A3" box="[691,746,398,421]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A12788–9,
<collectionCode id="ADEFAE5B002F3F40893C6584FEAF07C0" box="[272,359,430,454]" pageId="18" pageNumber="37">INBAC</collectionCode>
: WC-4614, Quembo River, source trap 2, -13.13544° 19.04397°,
<quantity id="0C069B7B002F3F4089FC65E4FDE007E3" box="[464,552,462,486]" metricMagnitude="3" metricUnit="m" metricValue="1.374" pageId="18" pageNumber="37" unit="m" value="1374.0">1,374 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A7565E4FD5807E3" box="[601,656,462,485]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14112 (tadpoles), Cuiva River, -11.98346° 17.72841°,
<quantity id="0C069B7B002F3F408A8165C4FF6A0423" metricMagnitude="3" metricUnit="m" metricValue="1.264" pageId="18" pageNumber="37" unit="m" value="1264.0">1,264 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F4088F36624FEDE0423" box="[223,278,526,549]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A12815,
<collectionCode id="ADEFAE5B002F3F4089A56624FE080423" box="[393,448,526,549]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14125 (tadpoles), Cuiva River source, -12.66856° 18.35307°,
<quantity id="0C069B7B002F3F408A0F6604FDB50443" box="[547,637,558,582]" metricMagnitude="3" metricUnit="m" metricValue="1.4069999999999998" pageId="18" pageNumber="37" unit="m" value="1407.0">1,407 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A9F6604FD220443" box="[691,746,558,581]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A12857,
<collectionCode id="ADEFAE5B002F3F4088DA6664FE850460" box="[246,333,590,614]" pageId="18" pageNumber="37">INBAC</collectionCode>
: WC-4616;
<collectionCode id="ADEFAE5B002F3F4089F56664FDE40460" box="[473,556,590,614]" pageId="18" pageNumber="37">INBAC</collectionCode>
(no number x5) Quembo River source lake, -13.13624° 19.04591°,
<quantity id="0C069B7B002F3F4088A166A4FF2704A3" box="[141,239,654,678]" metricMagnitude="3" metricUnit="m" metricValue="1.411" pageId="18" pageNumber="37" unit="m" value="1411.0">1,411 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F40891B66A4FEA604A3" box="[311,366,654,677]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A13767–72,
<collectionCode id="ADEFAE5B002F3F408A3966A4FDA404A0" box="[533,620,654,678]" pageId="18" pageNumber="37">INBAC</collectionCode>
: WC-6979 Lungwebungu River, old oxbows, -12.58129° 18.67162°,
<quantity id="0C069B7B002F3F4088A166E4FF2F04E3" box="[141,231,718,742]" metricMagnitude="3" metricUnit="m" metricValue="1.304" pageId="18" pageNumber="37" unit="m" value="1304.0">1,304 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F40893166E4FE9C04E3" box="[285,340,718,741]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A13800–2,
<collectionCode id="ADEFAE5B002F3F4089F766E4FDDA04E3" box="[475,530,718,741]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A13803 (tadpoles), Lake Tchanssengwe, -12.41402° 18.64417°,
<quantity id="0C069B7B002F3F408A4466C7FD770503" box="[616,703,749,773]" metricMagnitude="3" metricUnit="m" metricValue="1.415" pageId="18" pageNumber="37" unit="m" value="1415.0">1,415 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F4088A16724FF0C0523" box="[141,196,782,805]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14090 (tadpoles),
<quantity id="0C069B7B002F3F4089B06724FE190520" box="[412,465,782,806]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="37" unit="km" value="4.0">4 km</quantity>
upstream from Cuanavale River source, -13.05084° 18.89726°,
<quantity id="0C069B7B002F3F408A0F6704FDB50543" box="[547,637,814,838]" metricMagnitude="3" metricUnit="m" metricValue="1.394" pageId="18" pageNumber="37" unit="m" value="1394.0">1,394 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A9F6704FD220543" box="[691,746,814,837]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14103 (tadpoles),
<quantity id="0C069B7B002F3F4089456764FE6B0560" box="[361,419,846,870]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="37" unit="km" value="4.0">4 km</quantity>
downstream from Cuanavale River source camp, -13.11585° 18.90246°,
<quantity id="0C069B7B002F3F408AA56747FD220583" box="[649,746,877,901]" metricMagnitude="3" metricUnit="m" metricValue="1.354" pageId="18" pageNumber="37" unit="m" value="1354.0">1,354 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F40889267A4FF3D05A3" box="[190,245,910,933]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14106 (tadpoles), confluence of Cuito and Calua rivers, -13.12458° 18.89989°,
<quantity id="0C069B7B002F3F408A0D6787FDB305C3" box="[545,635,941,965]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="18" pageNumber="37" unit="m" value="1345.0">1,345 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A9F6784FD2205C3" box="[691,746,942,965]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14686, detour route just across bridge over Cuiva River, -12.13942° 18.39393°,
<quantity id="0C069B7B002F3F4089E267C7FDEE0203" box="[462,550,1005,1029]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="18" pageNumber="37" unit="m" value="1385.0">1,385 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A7467C4FD470203" box="[600,655,1006,1029]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14693 (tadpoles), Lungwebungu River camp, first oxbow on right side, -12.58117° 18.67106°,
<quantity id="0C069B7B002F3F408A386004FDBA0243" box="[532,626,1070,1094]" metricMagnitude="3" metricUnit="m" metricValue="1.294" pageId="18" pageNumber="37" unit="m" value="1294.0">1,294 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A9F6004FD220243" box="[691,746,1070,1093]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14717, en route from Samanunga village to Cuanavale River source, -12.94331° 18.81118°,
<quantity id="0C069B7B002F3F408A0F6044FDB40283" box="[547,636,1134,1158]" metricMagnitude="3" metricUnit="m" metricValue="1.4069999999999998" pageId="18" pageNumber="37" unit="m" value="1407.0">1,407 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A9F6044FD220283" box="[691,746,1134,1157]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14727–8, Quembo River, oxbow near small waterfall, -13.54257° 19.29551°,
<quantity id="0C069B7B002F3F4089B86084FE3A02C3" box="[404,498,1198,1222]" metricMagnitude="3" metricUnit="m" metricValue="1.233" pageId="18" pageNumber="37" unit="m" value="1233.0">1,233 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A1C6084FDAF02C3" box="[560,615,1198,1221]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14748–9,
<collectionCode id="ADEFAE5B002F3F4088A160E4FF0C02E3" box="[141,196,1230,1253]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14763–65,
<collectionCode id="ADEFAE5B002F3F40897360E4FE5E02E3" box="[351,406,1230,1253]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14778, Quembo River right side tributary (Micongo River) past village, -13.51877° 19.28487°,
<quantity id="0C069B7B002F3F4089256124FEA90323" box="[265,353,1294,1318]" metricMagnitude="3" metricUnit="m" metricValue="1.248" pageId="18" pageNumber="37" unit="m" value="1248.0">1,248 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F4089BF6124FE020323" box="[403,458,1294,1317]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A14757, left side tributary (Condinde River) at Cuando River bridge, -13.60076° 19.52675°
<quantity id="0C069B7B002F3F40893D6164FEBE0363" box="[273,374,1358,1382]" metricMagnitude="3" metricUnit="m" metricValue="1.219" pageId="18" pageNumber="37" unit="m" value="1219.0">1,219 m</quantity>
asl.
<emphasis id="F98AEA8C002F3F4089EF6167FEEE0383" bold="true" pageId="18" pageNumber="37">Additional material (16 tadpole lots):</emphasis>
<collectionCode id="ADEFAE5B002F3F40891C6144FE480380" box="[304,384,1390,1414]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
204553 (7 tadpoles), Cuanavale River source lake, -13.08997° 18.89389°,
<quantity id="0C069B7B002F3F408A4C61A7FD7303A3" box="[608,699,1421,1445]" metricMagnitude="3" metricUnit="m" metricValue="1.358" pageId="18" pageNumber="37" unit="m" value="1358.0">1,358 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F4088A16184FF1503C0" box="[141,221,1454,1478]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
209059 (15 tadpoles), Cuanavale River near confluence, -13.12478° 18.90017°,
<quantity id="0C069B7B002F3F408A2061E4FDAD03E3" box="[524,613,1486,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.347" pageId="18" pageNumber="37" unit="m" value="1347.0">1,347 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408AB561E4FD2103E0" box="[665,745,1486,1510]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
204509 (29 tadpoles), outlet from Kuembo River source lake, -13.13689° 19.03144°,
<quantity id="0C069B7B002F3F4089926227FDDD0023" box="[446,533,1549,1573]" metricMagnitude="3" metricUnit="m" metricValue="1.375" pageId="18" pageNumber="37" unit="m" value="1375.0">1,375 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A686224FD5C0020" box="[580,660,1550,1574]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
204564 (25 tadpoles),
<collectionCode id="ADEFAE5B002F3F4089156204FE410040" box="[313,393,1582,1606]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
209074 (1 tadpole), Quembo River source lake, -13.13611° 19.04500°,
<quantity id="0C069B7B002F3F408A736264FD720063" box="[607,698,1614,1638]" metricMagnitude="3" metricUnit="m" metricValue="1.366" pageId="18" pageNumber="37" unit="m" value="1366.0">1,366 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F4088A16244FF150080" box="[141,221,1646,1670]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
209066 (9 tadpoles), Quembo River source lake, -13.14025° 19.04822°,
<quantity id="0C069B7B002F3F4089A162A7FE2000A3" box="[397,488,1677,1701]" metricMagnitude="3" metricUnit="m" metricValue="1.365" pageId="18" pageNumber="37" unit="m" value="1365.0">1,365 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A0C62A4FDB800A0" box="[544,624,1678,1702]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
209062 (2 tadpoles), swamp near Cuanavale River source lake camp, -13.10750° 18.86089°,
<quantity id="0C069B7B002F3F4089B462E4FE3000E3" box="[408,504,1742,1766]" metricMagnitude="3" metricUnit="m" metricValue="1.386" pageId="18" pageNumber="37" unit="m" value="1386.0">1,386 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A1662E4FD4200E0" box="[570,650,1742,1766]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
204465 (19 tadpoles), Cuando River source pool, -13.00383° 19.12719°,
<quantity id="0C069B7B002F3F4089226327FEA30123" box="[270,363,1805,1829]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="18" pageNumber="37" unit="m" value="1350.0">1,350 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F40898B6324FE3F0120" box="[423,503,1806,1830]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
209060 (2 tadpoles), stream outflow about
<quantity id="0C069B7B002F3F4089586304FE760140" box="[372,446,1838,1862]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="18" pageNumber="37" unit="km" value="2.6">2.6 km</quantity>
downstream of source lake, -13.00317° 19.15153°,
<quantity id="0C069B7B002F3F4089A66364FE2A0163" box="[394,482,1870,1894]" metricMagnitude="3" metricUnit="m" metricValue="1.331" pageId="18" pageNumber="37" unit="m" value="1331.0">1,331 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408A3A6364FDAE0160" box="[534,614,1870,1894]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
204472 (13 tadpoles), Cuando River, -13.09142° 19.35850°,
<quantity id="0C069B7B002F3F408A816344FF6A01A3" metricMagnitude="3" metricUnit="m" metricValue="1.283" pageId="18" pageNumber="37" unit="m" value="1283.0">1,283 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F4088FA63A4FEEE01A0" box="[214,294,1934,1958]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
209067 (8 tadpoles), on Cuanavale River road, -13.14190° 19.44503°,
<quantity id="0C069B7B002F3F408C6564A4FB5706A3" box="[1097,1183,142,166]" metricMagnitude="3" metricUnit="m" metricValue="1.301" pageId="18" pageNumber="37" unit="m" value="1301.0">1,301 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408CE264A4FAD606A0" box="[1230,1310,142,166]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
209070 (10 tadpoles), pan near Cuanavale River camp, seepage, -13.10750° 18.86089°,
<quantity id="0C069B7B002F3F408C3F64E4FBA406E3" box="[1043,1132,206,230]" metricMagnitude="3" metricUnit="m" metricValue="1.387" pageId="18" pageNumber="37" unit="m" value="1387.0">1,387 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408C8C64E4FB3806E0" box="[1184,1264,206,230]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
204569 (13 tadpoles), Samununga village, -12.93228° 18.81672°,
<quantity id="0C069B7B002F3F408B3B6527FCB90723" box="[791,881,269,293]" metricMagnitude="3" metricUnit="m" metricValue="1.365" pageId="18" pageNumber="37" unit="m" value="1365.0">1,365 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408B8B6524FC3F0720" box="[935,1015,270,294]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
209063 (6 tadpoles), small stream
<quantity id="0C069B7B002F3F408B3B6504FC870740" box="[791,847,302,326]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="37" unit="km" value="3.0">3 km</quantity>
below Cuanavale River source camp, -13.12539° 18.89914°,
<quantity id="0C069B7B002F3F408BBA6564FC390763" box="[918,1009,334,358]" metricMagnitude="3" metricUnit="m" metricValue="1.344" pageId="18" pageNumber="37" unit="m" value="1344.0">1,344 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408C066564FBB20760" box="[1066,1146,334,358]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
209069 (11 tadpoles), pool in wetland on road edge, west of Munhango, -12.17806° 18.24306°,
<quantity id="0C069B7B002F3F408C0D65A4FB4907A3" box="[1057,1153,398,422]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="18" pageNumber="37" unit="m" value="1370.0">1,370 m</quantity>
asl;
<collectionCode id="ADEFAE5B002F3F408CE865A4FADC07A0" box="[1220,1300,398,422]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="37" type="Other type of research institution/biorepository">SAIAB</collectionCode>
209075 (18 tadpoles), Cuando River source bog, -13.00383° 19.12719°,
<quantity id="0C069B7B002F3F408BBD65E7FC2007E3" box="[913,1000,461,485]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="18" pageNumber="37" unit="m" value="1350.0">1,350 m</quantity>
asl.
<emphasis id="F98AEA8C002F3F408C3B65E7FB6C07E3" bold="true" box="[1047,1188,461,485]" pageId="18" pageNumber="37">Description:</emphasis>
Medium sized reed frog; rounded snout; reduced webbing; ventrum granular. Three different color forms were observed, i.e., plain light to dark green, brown, and bright red; thin dorsolateral white stripe present; small white specks on dorsal side of legs; toes and webbing red. Tadpoles with ventrum spotted and ventral blotches remain present in juveniles but fade in adults. Adult females (n = 11) varied from 16.7–21.9 (19.6) mm (largest female:
<collectionCode id="ADEFAE5B002F3F408CE666E4FAC904E3" box="[1226,1281,718,741]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A12549); adult males (n = 52) varied from 14.8–21.1 (18.7) mm (largest male:
<collectionCode id="ADEFAE5B002F3F408BED6724FC300523" box="[961,1016,782,805]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="18" pageNumber="37" type="Museum">PEM</collectionCode>
A12592).
<emphasis id="F98AEA8C002F3F408C506727FC7B0540" bold="true" pageId="18" pageNumber="37">Habitat and natural history notes:</emphasis>
Found at the margins of large rivers and source lakes within dense emergent wetland vegetation. Sympatric with other species of
<emphasis id="F98AEA8C002F3F408CAC6744FB340580" box="[1152,1276,878,902]" italics="true" pageId="18" pageNumber="37">
<taxonomicName id="0CFE4D1D002F3F408CAC6744FB300580" box="[1152,1272,878,902]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="18" pageNumber="37" phylum="Chordata" rank="genus">Hyperolius</taxonomicName>
,
</emphasis>
mostly
<taxonomicName id="0CFE4D1D002F3F408D756744FCA005A3" authorityName="Gunther" authorityYear="1865" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="18" pageNumber="37" phylum="Chordata" rank="species" species="nasutus">
<emphasis id="F98AEA8C002F3F408D756744FCA005A3" italics="true" pageId="18" pageNumber="37">H. nasutus</emphasis>
</taxonomicName>
and
<emphasis id="F98AEA8C002F3F408B8367A4FB8405A0" box="[943,1100,910,934]" italics="true" pageId="18" pageNumber="37">
<taxonomicName id="0CFE4D1D002F3F408B8367A4FB8105A0" authorityName="Gunther" authorityYear="1858" box="[943,1097,910,934]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="18" pageNumber="37" phylum="Chordata" rank="species" species="parallelus">H. parallelus</taxonomicName>
.
</emphasis>
<emphasis id="F98AEA8C002F3F408C7767A7FB1705A3" bold="true" box="[1115,1247,909,933]" pageId="18" pageNumber="37">Comments:</emphasis>
Initially we assigned these specimens to
<taxonomicName id="0CFE4D1D002F3F408C686784FAA705C0" authority="(NGOWP 2017)" baseAuthorityName="NGOWP" baseAuthorityYear="2017" box="[1092,1391,942,966]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="18" pageNumber="37" phylum="Chordata" rank="species" species="bocagei">
<emphasis id="F98AEA8C002F3F408C686784FB7005C0" box="[1092,1208,942,966]" italics="true" pageId="18" pageNumber="37">H. bocagei</emphasis>
(NGOWP 2017)
</taxonomicName>
, however it is generally regarded as a larger species that is associated with the
<taxonomicName id="0CFE4D1D002F3F408C3667C4FB5A0200" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1841" box="[1050,1170,1006,1030]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="18" pageNumber="37" phylum="Chordata" rank="species" species="viridiflavus">
<emphasis id="F98AEA8C002F3F408C3667C4FB5A0200" box="[1050,1170,1006,1030]" italics="true" pageId="18" pageNumber="37">viridiflavus</emphasis>
</taxonomicName>
group (see
<bibRefCitation id="AF6F4B6F002F3F408D0F67C4FC980220" author="Schiotz A." pageId="18" pageNumber="37" refId="ref31477" refString="Schiotz A. 1999. Treefrogs of Africa. Editions Chimaira, Frankfurt am Main, Germany. 350 p." type="book" year="1999">Schiøtz 1999</bibRefCitation>
). This new material differs in size (smaller) and webbing (less webbing) from
<taxonomicName id="0CFE4D1D002F3F408C786004FAA70240" authority="(Schiotz 1999)" baseAuthorityName="Schiotz" baseAuthorityYear="1999" box="[1108,1391,1070,1094]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="18" pageNumber="37" phylum="Chordata" rank="species" species="bocagei">
<emphasis id="F98AEA8C002F3F408C786004FB000240" box="[1108,1224,1070,1094]" italics="true" pageId="18" pageNumber="37">H. bocagei</emphasis>
(
<bibRefCitation id="AF6F4B6F002F3F408CFA6004FAAF0240" author="Schiotz A." box="[1238,1383,1070,1094]" pageId="18" pageNumber="37" refId="ref31477" refString="Schiotz A. 1999. Treefrogs of Africa. Editions Chimaira, Frankfurt am Main, Germany. 350 p." type="book" year="1999">Schiøtz 1999</bibRefCitation>
)
</taxonomicName>
. Phylogenetic work is currently underway to determine its taxonomic status.
</paragraph>
</subSubSection>
</treatment>
</document>
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