161 lines
34 KiB
XML
161 lines
34 KiB
XML
<document id="EECA4219E5357DB6B6A4CC0FC58942DA" ID-CLB-Dataset="5966" ID-DOI="10.5852/ejt.2021.755.1405" ID-GBIF-Dataset="b932c9e5-ddad-4c11-8eee-ff17188d77cf" ID-ISSN="2118-9773" ID-Zenodo-Dep="5032970" ID-ZooBank="F9105E33-3E8B-4B3C-88B3-0316207B70F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1624643559605" checkinUser="felipe" docAuthor="Gale, Andy S. & Jagt, John W. M." docDate="2021" docId="F160A367716DFFA9FD87760EFEAE263A" docLanguage="en" docName="ejt-755_149-190.pdf" docOrigin="European Journal of Taxonomy 755" docSource="http://dx.doi.org/10.5852/ejt.2021.755.1405" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chrispaulia spinosa Gale & Jagt 2021, sp. nov." docType="treatment" docUuid="4C003A9F-1F91-44BC-91AB-2910B8EA756A" docUuidSource="ZooBank" docVersion="7" lastPageNumber="176" masterDocId="0D59DB1F7176FFB2FFF77706FFD02038" masterDocTitle="The fossil record of the family Benthopectinidae (Echinodermata, Asteroidea), a reappraisal" masterLastPageNumber="190" masterPageNumber="149" pageNumber="176" updateTime="1698976416854" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8CBA44F0F46AD2677F5C9B323C284AC6">The fossil record of the family Benthopectinidae (Echinodermata, Asteroidea), a reappraisal</mods:title>
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<mods:roleTerm id="110329806F17BF674FCB8AF14FC65996">Author</mods:roleTerm>
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<mods:namePart id="75069322D19B0DE9E4A52E7D0A6B2951">Gale, Andy S.</mods:namePart>
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<mods:affiliation id="611B7EFEE378543632B0A4530C9EECF2">Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom. School of the Environment, Geography and Geological Sciences, University of Portsmouth, Burnaby Building, Burnaby Road, Portsmouth PO 13 QL, United Kingdom. Natuurhistorisch Museum Maastricht, De Bosquetplein 6 - 7, 6211 KJ Maastricht, the Netherlands.</mods:affiliation>
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<mods:nameIdentifier id="6CD9FC5CEEA64CFE6B84755065346182" type="email">andy.gale@port.ac.uk</mods:nameIdentifier>
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<mods:namePart id="875AC4C87F19E7DD82E0AE2371D87739">Jagt, John W. M.</mods:namePart>
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<mods:nameIdentifier id="2DD1CB221786727524D229EDB5D0CDB7" type="email">john.jagt@maastricht.nl</mods:nameIdentifier>
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<mods:title id="4416F2D5A1C1E5DFA0FA27198CC66224">European Journal of Taxonomy</mods:title>
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<mods:date id="E91A76EFB6D80B3F2E01837D91F9D41C">2021</mods:date>
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<treatment id="F160A367716DFFA9FD87760EFEAE263A" ID-DOI="http://doi.org/10.5281/zenodo.5033000" ID-GBIF-Taxon="183195103" ID-Zenodo-Dep="5033000" LSID="urn:lsid:zoobank.org:act:4C003A9F-1F91-44BC-91AB-2910B8EA756A" httpUri="http://treatment.plazi.org/id/F160A367716DFFA9FD87760EFEAE263A" lastPageNumber="176" pageId="27" pageNumber="176">
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<subSubSection id="31D341FA716DFFA9FD87760EFC14211B" box="[624,964,264,291]" pageId="27" pageNumber="176" type="nomenclature">
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<paragraph id="79761271716DFFA9FD87760EFC14211B" blockId="27.[365,1224,264,371]" box="[624,964,264,291]" pageId="27" pageNumber="176">
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<heading id="223EA51D716DFFA9FD87760EFC14211B" box="[624,964,264,291]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="176" reason="2">
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<taxonomicName id="BEC969F2716DFFA9FD87760EFCB2211B" authority="Gale & Jagt, 2021" authorityName="Gale & Jagt" authorityYear="2021" box="[624,866,264,291]" class="Asteroidea" family="Goniopectinidae" genus="Chrispaulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paxillosida" pageId="27" pageNumber="176" phylum="Echinodermata" rank="species" species="spinosa" status="sp. nov.">
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<emphasis id="4BBDCE63716DFFA9FD87760EFCB2211B" bold="true" box="[624,866,264,291]" italics="true" pageId="27" pageNumber="176">Chrispaulia spinosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="508E7318716DFFA9FC9E760FFC14211B" box="[873,964,265,291]" pageId="27" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="31D341FA716DFFA9FE9A7637FC52214B" pageId="27" pageNumber="176" type="description">
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<paragraph id="79761271716DFFA9FE9A7637FB182174" blockId="27.[365,1224,264,371]" box="[365,1224,305,332]" pageId="27" pageNumber="176">
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urn:lsid:zoobank.org:act:
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<uuid id="0D6F28A4716DFFA9FD797637FB182174" box="[654,1224,305,332]" pageId="27" pageNumber="176">4C003A9F-1F91-44BC-91AB-2910B8EA756A</uuid>
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</uri>
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</paragraph>
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<paragraph id="79761271716DFFA9FD46765FFC52214B" blockId="27.[365,1224,264,371]" box="[689,898,345,371]" pageId="27" pageNumber="176">
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<figureCitation id="E1F20EF4716DFFA9FD46765FFC52214B" box="[689,898,345,371]" captionStart="Fig" captionStartId="15.[189,232,1328,1354]" captionTargetId="figure-15@16.[189,1398,283,1870]" captionTargetPageId="16" captionText="Fig. 7 (next page). A–J. Jurapecten infrajurensis sp. nov., various ossicle types (paratypes). A, H. Ambulacrals, in lateral and actinal aspects, respectively (SNSB-BGSP2020XLV 21,SNSB- BGSP2020XLV 28). B, E–F. Adambulacrals, in actinal and abactinal views (SNSB-BGSP2020XLV 22, SNSB-BGSP2020XLV 25–26). C, G, J. Superomarginals, in lateral views (SNSB-BGSP2020XLV 23, SNSB-BGSP2020XLV 27, SNSB-BGSP2020XLV 30). D, I. Inferomarginals, in lateral views (SNSB-BGSP2020XLV 24, SNSB-BGSP2020XLV 29). — K–Y. Jurapecten hessi Gale, 2011. K, P, U. Adambulacral ossicles. K. Abactinal view (NHMUK PI EE 17996). P. NHMUK EE 13598 (the original of Gale 2011a: pl. 19 fig. 11). U. NHMUK EE 13599, the original of Gale (2011a: pl. 20 fig. 12). L. Circumoral ossicle (NHMUK EE 13605, the original of Gale 2011a: pl. 20 fig. 13). M–N. Ambulacral ossicles, in actinal and abactinal views, respectively (NHMUK EE 13595–13596, the originals of Gale 2011a: pl. 19 figs 8–9). O. Ambulacral, in proximal/distal view (NHMUK 13597). Q–S. Superomarginal ossicles (NHMUK EE 13601–13602; the originals of Gale 2011a: pl. 20 figs 6–7). T, V–W. Inferomarginals (NHMUK EE 13600, EE 13603–13604, EE 13608, the originals of Gale 2011a: pl. 20 figs 5, 8, 16). X–Y. Oral ossicles (NHMUK EE 13606– 13607, the originals of Gale 2011a: pl. 20 figs 10, 14). Provenance: A–J. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; see Hess 2014). K–Y. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). Abbreviations: d = distal; p = proximal. See Fig. 3 for detailed morphological terminology. Scale bars: A–J = 0.2 mm; K–Y = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5036870" httpUri="https://zenodo.org/record/5036870/files/figure.png" pageId="27" pageNumber="176">Fig. 13D–E, H–O</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="31D341FA716DFFA9FF4A7699FDB5222C" pageId="27" pageNumber="176" type="diagnosis">
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<paragraph id="79761271716DFFA9FF4A7699FEE62181" blockId="27.[189,310,415,441]" box="[189,310,415,441]" pageId="27" pageNumber="176">
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||
<heading id="223EA51D716DFFA9FF4A7699FEE62181" bold="true" box="[189,310,415,441]" fontSize="11" level="3" pageId="27" pageNumber="176" reason="3">
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||
<emphasis id="4BBDCE63716DFFA9FF4A7699FEE62181" bold="true" box="[189,310,415,441]" pageId="27" pageNumber="176">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="79761271716DFFA9FF4A76D4FDB5222C" blockId="27.[189,1399,466,532]" pageId="27" pageNumber="176">
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<taxonomicName id="BEC969F2716DFFA9FF4A76D4FE9921D4" authorityName="Gale" authorityYear="2005" box="[189,329,466,492]" class="Asteroidea" family="Goniopectinidae" genus="Chrispaulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paxillosida" pageId="27" pageNumber="176" phylum="Echinodermata" rank="genus">
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<emphasis id="4BBDCE63716DFFA9FF4A76D4FE9921D4" box="[189,329,466,492]" italics="true" pageId="27" pageNumber="176">Chrispaulia</emphasis>
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</taxonomicName>
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in which distal inferomarginals bear an oblique row of large, bifid spine attachment sites, which carried long, flattened spines.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="31D341FA716DFFA9FF4A7547FA8D22B6" box="[189,1373,577,654]" pageId="27" pageNumber="176" type="etymology">
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||
<paragraph id="79761271716DFFA9FF4A7547FE942263" blockId="27.[189,324,577,603]" box="[189,324,577,603]" pageId="27" pageNumber="176">
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||
<heading id="223EA51D716DFFA9FF4A7547FE942263" bold="true" box="[189,324,577,603]" fontSize="11" level="3" pageId="27" pageNumber="176" reason="3">
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||
<emphasis id="4BBDCE63716DFFA9FF4A7547FE942263" bold="true" box="[189,324,577,603]" pageId="27" pageNumber="176">Etymology</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="79761271716DFFA9FF4A7572FA8D22B6" blockId="27.[189,1373,628,654]" box="[189,1373,628,654]" pageId="27" pageNumber="176">Latin for ‘bearing spines’, in reference to the row of spine attachment sites on distal inferomarginals.</paragraph>
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</subSubSection>
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<subSubSection id="31D341FA716DFFA9FF4A75BCFD032390" pageId="27" pageNumber="176" type="materials_examined">
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<paragraph id="79761271716DFFA9FF4A75BCFE7A22EC" blockId="27.[189,426,698,724]" box="[189,426,698,724]" pageId="27" pageNumber="176">
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<heading id="223EA51D716DFFA9FF4A75BCFE7A22EC" bold="true" box="[189,426,698,724]" fontSize="11" level="3" pageId="27" pageNumber="176" reason="3">
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<emphasis id="4BBDCE63716DFFA9FF4A75BCFE7A22EC" bold="true" box="[189,426,698,724]" pageId="27" pageNumber="176">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="79761271716DFFA9FF4A75E8FD032390" blockId="27.[188,1399,749,936]" pageId="27" pageNumber="176">
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<materialsCitation id="C9A1182C716DFFA9FF4A75E8FD032390" ID-GBIF-Occurrence="3311523305" collectionCode="NHMUK" pageId="27" pageNumber="176" specimenCode="Nds LH 105.107, NHMUK PI EE 17998-18004" specimenCount="1">
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A distal arm fragment (
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<specimenCode id="296FBA0A716DFFA9FE1575EBFD672330" box="[482,695,749,776]" pageId="27" pageNumber="176">Nds LH 105.107</specimenCode>
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), comprising five marginal pairs, with articulating spines preserved, is the holotype; it is from the lower Hauterivian (
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<taxonomicName id="BEC969F2716DFFA9FC5D7413FB262317" box="[938,1270,789,815]" class="Cephalopoda" genus="Endemoceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="27" pageNumber="176" phylum="Mollusca" rank="species" species="amblygonium">
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<emphasis id="4BBDCE63716DFFA9FC5D7413FB262317" box="[938,1270,789,815]" italics="true" pageId="27" pageNumber="176">Endemoceras amblygonium</emphasis>
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</taxonomicName>
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ammonite Zone) at Engelbostel near Hannover (northern Germany). Paratypes (
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<specimenCode id="296FBA0A716DFFA9FC17743BFA94236F" box="[992,1348,829,856]" pageId="27" pageNumber="176">NHMUK PI EE 17998–18004</specimenCode>
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) are 35 marginal ossicles and a single oral ossicle from the upper 3 metres of the Tealby Clay (Hauterivian) at Nettleton, Lincolnshire (United Kingdom).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="31D341FA716DFFA9FF4A74D5FACF2558" pageId="27" pageNumber="176" type="description">
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||
<paragraph id="79761271716DFFA9FF4A74D5FE9E23D5" blockId="27.[189,334,979,1005]" box="[189,334,979,1005]" pageId="27" pageNumber="176">
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<heading id="223EA51D716DFFA9FF4A74D5FE9E23D5" bold="true" box="[189,334,979,1005]" fontSize="11" level="3" pageId="27" pageNumber="176" reason="3">
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<emphasis id="4BBDCE63716DFFA9FF4A74D5FE9E23D5" bold="true" box="[189,334,979,1005]" pageId="27" pageNumber="176">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="79761271716DFFA9FF4A7301FACF2558" blockId="27.[187,1399,1031,1377]" pageId="27" pageNumber="176">
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Arms elongated, narrow, tapering slowly (
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<figureCitation id="E1F20EF4716DFFA9FD3E7301FC8E2419" box="[713,862,1031,1057]" captionStart="Fig" captionStartId="15.[189,232,1328,1354]" captionTargetId="figure-15@16.[189,1398,283,1870]" captionTargetPageId="16" captionText="Fig. 7 (next page). A–J. Jurapecten infrajurensis sp. nov., various ossicle types (paratypes). A, H. Ambulacrals, in lateral and actinal aspects, respectively (SNSB-BGSP2020XLV 21,SNSB- BGSP2020XLV 28). B, E–F. Adambulacrals, in actinal and abactinal views (SNSB-BGSP2020XLV 22, SNSB-BGSP2020XLV 25–26). C, G, J. Superomarginals, in lateral views (SNSB-BGSP2020XLV 23, SNSB-BGSP2020XLV 27, SNSB-BGSP2020XLV 30). D, I. Inferomarginals, in lateral views (SNSB-BGSP2020XLV 24, SNSB-BGSP2020XLV 29). — K–Y. Jurapecten hessi Gale, 2011. K, P, U. Adambulacral ossicles. K. Abactinal view (NHMUK PI EE 17996). P. NHMUK EE 13598 (the original of Gale 2011a: pl. 19 fig. 11). U. NHMUK EE 13599, the original of Gale (2011a: pl. 20 fig. 12). L. Circumoral ossicle (NHMUK EE 13605, the original of Gale 2011a: pl. 20 fig. 13). M–N. Ambulacral ossicles, in actinal and abactinal views, respectively (NHMUK EE 13595–13596, the originals of Gale 2011a: pl. 19 figs 8–9). O. Ambulacral, in proximal/distal view (NHMUK 13597). Q–S. Superomarginal ossicles (NHMUK EE 13601–13602; the originals of Gale 2011a: pl. 20 figs 6–7). T, V–W. Inferomarginals (NHMUK EE 13600, EE 13603–13604, EE 13608, the originals of Gale 2011a: pl. 20 figs 5, 8, 16). X–Y. Oral ossicles (NHMUK EE 13606– 13607, the originals of Gale 2011a: pl. 20 figs 10, 14). Provenance: A–J. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; see Hess 2014). K–Y. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). Abbreviations: d = distal; p = proximal. See Fig. 3 for detailed morphological terminology. Scale bars: A–J = 0.2 mm; K–Y = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5036870" httpUri="https://zenodo.org/record/5036870/files/figure.png" pageId="27" pageNumber="176">Fig. 13D–E</figureCitation>
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); proximal superomarginals tall, block-like, bearing a single, abactinally directed, crater-like spine base. Central region of external face narrow, poorly defined, lateral surfaces broad with small rugosities for attachment of tiny cribriform spines (
|
||
<figureCitation id="E1F20EF4716DFFA9FF317379FEBA24A1" box="[198,362,1151,1177]" captionStart="Fig" captionStartId="15.[189,232,1328,1354]" captionTargetId="figure-15@16.[189,1398,283,1870]" captionTargetPageId="16" captionText="Fig. 7 (next page). A–J. Jurapecten infrajurensis sp. nov., various ossicle types (paratypes). A, H. Ambulacrals, in lateral and actinal aspects, respectively (SNSB-BGSP2020XLV 21,SNSB- BGSP2020XLV 28). B, E–F. Adambulacrals, in actinal and abactinal views (SNSB-BGSP2020XLV 22, SNSB-BGSP2020XLV 25–26). C, G, J. Superomarginals, in lateral views (SNSB-BGSP2020XLV 23, SNSB-BGSP2020XLV 27, SNSB-BGSP2020XLV 30). D, I. Inferomarginals, in lateral views (SNSB-BGSP2020XLV 24, SNSB-BGSP2020XLV 29). — K–Y. Jurapecten hessi Gale, 2011. K, P, U. Adambulacral ossicles. K. Abactinal view (NHMUK PI EE 17996). P. NHMUK EE 13598 (the original of Gale 2011a: pl. 19 fig. 11). U. NHMUK EE 13599, the original of Gale (2011a: pl. 20 fig. 12). L. Circumoral ossicle (NHMUK EE 13605, the original of Gale 2011a: pl. 20 fig. 13). M–N. Ambulacral ossicles, in actinal and abactinal views, respectively (NHMUK EE 13595–13596, the originals of Gale 2011a: pl. 19 figs 8–9). O. Ambulacral, in proximal/distal view (NHMUK 13597). Q–S. Superomarginal ossicles (NHMUK EE 13601–13602; the originals of Gale 2011a: pl. 20 figs 6–7). T, V–W. Inferomarginals (NHMUK EE 13600, EE 13603–13604, EE 13608, the originals of Gale 2011a: pl. 20 figs 5, 8, 16). X–Y. Oral ossicles (NHMUK EE 13606– 13607, the originals of Gale 2011a: pl. 20 figs 10, 14). Provenance: A–J. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; see Hess 2014). K–Y. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). Abbreviations: d = distal; p = proximal. See Fig. 3 for detailed morphological terminology. Scale bars: A–J = 0.2 mm; K–Y = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5036870" httpUri="https://zenodo.org/record/5036870/files/figure.png" pageId="27" pageNumber="176">Fig. 13H–I, N</figureCitation>
|
||
; spines still articulated in
|
||
<figureCitation id="E1F20EF4716DFFA9FD667379FD2A24A1" box="[657,762,1151,1177]" captionStart="Fig" captionStartId="15.[189,232,1328,1354]" captionTargetId="figure-15@16.[189,1398,283,1870]" captionTargetPageId="16" captionText="Fig. 7 (next page). A–J. Jurapecten infrajurensis sp. nov., various ossicle types (paratypes). A, H. Ambulacrals, in lateral and actinal aspects, respectively (SNSB-BGSP2020XLV 21,SNSB- BGSP2020XLV 28). B, E–F. Adambulacrals, in actinal and abactinal views (SNSB-BGSP2020XLV 22, SNSB-BGSP2020XLV 25–26). C, G, J. Superomarginals, in lateral views (SNSB-BGSP2020XLV 23, SNSB-BGSP2020XLV 27, SNSB-BGSP2020XLV 30). D, I. Inferomarginals, in lateral views (SNSB-BGSP2020XLV 24, SNSB-BGSP2020XLV 29). — K–Y. Jurapecten hessi Gale, 2011. K, P, U. Adambulacral ossicles. K. Abactinal view (NHMUK PI EE 17996). P. NHMUK EE 13598 (the original of Gale 2011a: pl. 19 fig. 11). U. NHMUK EE 13599, the original of Gale (2011a: pl. 20 fig. 12). L. Circumoral ossicle (NHMUK EE 13605, the original of Gale 2011a: pl. 20 fig. 13). M–N. Ambulacral ossicles, in actinal and abactinal views, respectively (NHMUK EE 13595–13596, the originals of Gale 2011a: pl. 19 figs 8–9). O. Ambulacral, in proximal/distal view (NHMUK 13597). Q–S. Superomarginal ossicles (NHMUK EE 13601–13602; the originals of Gale 2011a: pl. 20 figs 6–7). T, V–W. Inferomarginals (NHMUK EE 13600, EE 13603–13604, EE 13608, the originals of Gale 2011a: pl. 20 figs 5, 8, 16). X–Y. Oral ossicles (NHMUK EE 13606– 13607, the originals of Gale 2011a: pl. 20 figs 10, 14). Provenance: A–J. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; see Hess 2014). K–Y. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). Abbreviations: d = distal; p = proximal. See Fig. 3 for detailed morphological terminology. Scale bars: A–J = 0.2 mm; K–Y = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5036870" httpUri="https://zenodo.org/record/5036870/files/figure.png" pageId="27" pageNumber="176">Fig. 13D</figureCitation>
|
||
). Distal superomarginals with or without an abactinally situated, distally directed large spine base (
|
||
<figureCitation id="E1F20EF4716DFFA9FD4673A1FCFC24F9" box="[689,812,1191,1217]" captionStart="Fig" captionStartId="15.[189,232,1328,1354]" captionTargetId="figure-15@16.[189,1398,283,1870]" captionTargetPageId="16" captionText="Fig. 7 (next page). A–J. Jurapecten infrajurensis sp. nov., various ossicle types (paratypes). A, H. Ambulacrals, in lateral and actinal aspects, respectively (SNSB-BGSP2020XLV 21,SNSB- BGSP2020XLV 28). B, E–F. Adambulacrals, in actinal and abactinal views (SNSB-BGSP2020XLV 22, SNSB-BGSP2020XLV 25–26). C, G, J. Superomarginals, in lateral views (SNSB-BGSP2020XLV 23, SNSB-BGSP2020XLV 27, SNSB-BGSP2020XLV 30). D, I. Inferomarginals, in lateral views (SNSB-BGSP2020XLV 24, SNSB-BGSP2020XLV 29). — K–Y. Jurapecten hessi Gale, 2011. K, P, U. Adambulacral ossicles. K. Abactinal view (NHMUK PI EE 17996). P. NHMUK EE 13598 (the original of Gale 2011a: pl. 19 fig. 11). U. NHMUK EE 13599, the original of Gale (2011a: pl. 20 fig. 12). L. Circumoral ossicle (NHMUK EE 13605, the original of Gale 2011a: pl. 20 fig. 13). M–N. Ambulacral ossicles, in actinal and abactinal views, respectively (NHMUK EE 13595–13596, the originals of Gale 2011a: pl. 19 figs 8–9). O. Ambulacral, in proximal/distal view (NHMUK 13597). Q–S. Superomarginal ossicles (NHMUK EE 13601–13602; the originals of Gale 2011a: pl. 20 figs 6–7). T, V–W. Inferomarginals (NHMUK EE 13600, EE 13603–13604, EE 13608, the originals of Gale 2011a: pl. 20 figs 5, 8, 16). X–Y. Oral ossicles (NHMUK EE 13606– 13607, the originals of Gale 2011a: pl. 20 figs 10, 14). Provenance: A–J. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; see Hess 2014). K–Y. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). Abbreviations: d = distal; p = proximal. See Fig. 3 for detailed morphological terminology. Scale bars: A–J = 0.2 mm; K–Y = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5036870" httpUri="https://zenodo.org/record/5036870/files/figure.png" pageId="27" pageNumber="176">Fig. 13I–J</figureCitation>
|
||
), rugose (
|
||
<figureCitation id="E1F20EF4716DFFA9FC6873A1FBD924F9" box="[927,1033,1191,1217]" captionStart="Fig" captionStartId="15.[189,232,1328,1354]" captionTargetId="figure-15@16.[189,1398,283,1870]" captionTargetPageId="16" captionText="Fig. 7 (next page). A–J. Jurapecten infrajurensis sp. nov., various ossicle types (paratypes). A, H. Ambulacrals, in lateral and actinal aspects, respectively (SNSB-BGSP2020XLV 21,SNSB- BGSP2020XLV 28). B, E–F. Adambulacrals, in actinal and abactinal views (SNSB-BGSP2020XLV 22, SNSB-BGSP2020XLV 25–26). C, G, J. Superomarginals, in lateral views (SNSB-BGSP2020XLV 23, SNSB-BGSP2020XLV 27, SNSB-BGSP2020XLV 30). D, I. Inferomarginals, in lateral views (SNSB-BGSP2020XLV 24, SNSB-BGSP2020XLV 29). — K–Y. Jurapecten hessi Gale, 2011. K, P, U. Adambulacral ossicles. K. Abactinal view (NHMUK PI EE 17996). P. NHMUK EE 13598 (the original of Gale 2011a: pl. 19 fig. 11). U. NHMUK EE 13599, the original of Gale (2011a: pl. 20 fig. 12). L. Circumoral ossicle (NHMUK EE 13605, the original of Gale 2011a: pl. 20 fig. 13). M–N. Ambulacral ossicles, in actinal and abactinal views, respectively (NHMUK EE 13595–13596, the originals of Gale 2011a: pl. 19 figs 8–9). O. Ambulacral, in proximal/distal view (NHMUK 13597). Q–S. Superomarginal ossicles (NHMUK EE 13601–13602; the originals of Gale 2011a: pl. 20 figs 6–7). T, V–W. Inferomarginals (NHMUK EE 13600, EE 13603–13604, EE 13608, the originals of Gale 2011a: pl. 20 figs 5, 8, 16). X–Y. Oral ossicles (NHMUK EE 13606– 13607, the originals of Gale 2011a: pl. 20 figs 10, 14). Provenance: A–J. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; see Hess 2014). K–Y. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). Abbreviations: d = distal; p = proximal. See Fig. 3 for detailed morphological terminology. Scale bars: A–J = 0.2 mm; K–Y = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5036870" httpUri="https://zenodo.org/record/5036870/files/figure.png" pageId="27" pageNumber="176">Fig. 13D</figureCitation>
|
||
) or smooth (
|
||
<figureCitation id="E1F20EF4716DFFA9FB6973A1FACB24F9" box="[1182,1307,1191,1217]" captionStart="Fig" captionStartId="15.[189,232,1328,1354]" captionTargetId="figure-15@16.[189,1398,283,1870]" captionTargetPageId="16" captionText="Fig. 7 (next page). A–J. Jurapecten infrajurensis sp. nov., various ossicle types (paratypes). A, H. Ambulacrals, in lateral and actinal aspects, respectively (SNSB-BGSP2020XLV 21,SNSB- BGSP2020XLV 28). B, E–F. Adambulacrals, in actinal and abactinal views (SNSB-BGSP2020XLV 22, SNSB-BGSP2020XLV 25–26). C, G, J. Superomarginals, in lateral views (SNSB-BGSP2020XLV 23, SNSB-BGSP2020XLV 27, SNSB-BGSP2020XLV 30). D, I. Inferomarginals, in lateral views (SNSB-BGSP2020XLV 24, SNSB-BGSP2020XLV 29). — K–Y. Jurapecten hessi Gale, 2011. K, P, U. Adambulacral ossicles. K. Abactinal view (NHMUK PI EE 17996). P. NHMUK EE 13598 (the original of Gale 2011a: pl. 19 fig. 11). U. NHMUK EE 13599, the original of Gale (2011a: pl. 20 fig. 12). L. Circumoral ossicle (NHMUK EE 13605, the original of Gale 2011a: pl. 20 fig. 13). M–N. Ambulacral ossicles, in actinal and abactinal views, respectively (NHMUK EE 13595–13596, the originals of Gale 2011a: pl. 19 figs 8–9). O. Ambulacral, in proximal/distal view (NHMUK 13597). Q–S. Superomarginal ossicles (NHMUK EE 13601–13602; the originals of Gale 2011a: pl. 20 figs 6–7). T, V–W. Inferomarginals (NHMUK EE 13600, EE 13603–13604, EE 13608, the originals of Gale 2011a: pl. 20 figs 5, 8, 16). X–Y. Oral ossicles (NHMUK EE 13606– 13607, the originals of Gale 2011a: pl. 20 figs 10, 14). Provenance: A–J. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; see Hess 2014). K–Y. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). Abbreviations: d = distal; p = proximal. See Fig. 3 for detailed morphological terminology. Scale bars: A–J = 0.2 mm; K–Y = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5036870" httpUri="https://zenodo.org/record/5036870/files/figure.png" pageId="27" pageNumber="176">Fig. 13I–J</figureCitation>
|
||
). Distal infero- and superomarginals thin, imbricating proximally; distal inferomarginals with oblique row of large, bifid spine bases, which bore flattened, lanceolate spines (
|
||
<figureCitation id="E1F20EF4716DFFA9FC5A73F0FBCA2529" box="[941,1050,1270,1297]" captionStart="Fig" captionStartId="15.[189,232,1328,1354]" captionTargetId="figure-15@16.[189,1398,283,1870]" captionTargetPageId="16" captionText="Fig. 7 (next page). A–J. Jurapecten infrajurensis sp. nov., various ossicle types (paratypes). A, H. Ambulacrals, in lateral and actinal aspects, respectively (SNSB-BGSP2020XLV 21,SNSB- BGSP2020XLV 28). B, E–F. Adambulacrals, in actinal and abactinal views (SNSB-BGSP2020XLV 22, SNSB-BGSP2020XLV 25–26). C, G, J. Superomarginals, in lateral views (SNSB-BGSP2020XLV 23, SNSB-BGSP2020XLV 27, SNSB-BGSP2020XLV 30). D, I. Inferomarginals, in lateral views (SNSB-BGSP2020XLV 24, SNSB-BGSP2020XLV 29). — K–Y. Jurapecten hessi Gale, 2011. K, P, U. Adambulacral ossicles. K. Abactinal view (NHMUK PI EE 17996). P. NHMUK EE 13598 (the original of Gale 2011a: pl. 19 fig. 11). U. NHMUK EE 13599, the original of Gale (2011a: pl. 20 fig. 12). L. Circumoral ossicle (NHMUK EE 13605, the original of Gale 2011a: pl. 20 fig. 13). M–N. Ambulacral ossicles, in actinal and abactinal views, respectively (NHMUK EE 13595–13596, the originals of Gale 2011a: pl. 19 figs 8–9). O. Ambulacral, in proximal/distal view (NHMUK 13597). Q–S. Superomarginal ossicles (NHMUK EE 13601–13602; the originals of Gale 2011a: pl. 20 figs 6–7). T, V–W. Inferomarginals (NHMUK EE 13600, EE 13603–13604, EE 13608, the originals of Gale 2011a: pl. 20 figs 5, 8, 16). X–Y. Oral ossicles (NHMUK EE 13606– 13607, the originals of Gale 2011a: pl. 20 figs 10, 14). Provenance: A–J. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; see Hess 2014). K–Y. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). Abbreviations: d = distal; p = proximal. See Fig. 3 for detailed morphological terminology. Scale bars: A–J = 0.2 mm; K–Y = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5036870" httpUri="https://zenodo.org/record/5036870/files/figure.png" pageId="27" pageNumber="176">Fig. 13D</figureCitation>
|
||
). Sharply defined grooves for cribriform organs between each infero-/superomarginal pair. Oral ossicle (
|
||
<figureCitation id="E1F20EF4716DFFA9FBD07218FB412501" box="[1063,1169,1310,1337]" captionStart="Fig" captionStartId="15.[189,232,1328,1354]" captionTargetId="figure-15@16.[189,1398,283,1870]" captionTargetPageId="16" captionText="Fig. 7 (next page). A–J. Jurapecten infrajurensis sp. nov., various ossicle types (paratypes). A, H. Ambulacrals, in lateral and actinal aspects, respectively (SNSB-BGSP2020XLV 21,SNSB- BGSP2020XLV 28). B, E–F. Adambulacrals, in actinal and abactinal views (SNSB-BGSP2020XLV 22, SNSB-BGSP2020XLV 25–26). C, G, J. Superomarginals, in lateral views (SNSB-BGSP2020XLV 23, SNSB-BGSP2020XLV 27, SNSB-BGSP2020XLV 30). D, I. Inferomarginals, in lateral views (SNSB-BGSP2020XLV 24, SNSB-BGSP2020XLV 29). — K–Y. Jurapecten hessi Gale, 2011. K, P, U. Adambulacral ossicles. K. Abactinal view (NHMUK PI EE 17996). P. NHMUK EE 13598 (the original of Gale 2011a: pl. 19 fig. 11). U. NHMUK EE 13599, the original of Gale (2011a: pl. 20 fig. 12). L. Circumoral ossicle (NHMUK EE 13605, the original of Gale 2011a: pl. 20 fig. 13). M–N. Ambulacral ossicles, in actinal and abactinal views, respectively (NHMUK EE 13595–13596, the originals of Gale 2011a: pl. 19 figs 8–9). O. Ambulacral, in proximal/distal view (NHMUK 13597). Q–S. Superomarginal ossicles (NHMUK EE 13601–13602; the originals of Gale 2011a: pl. 20 figs 6–7). T, V–W. Inferomarginals (NHMUK EE 13600, EE 13603–13604, EE 13608, the originals of Gale 2011a: pl. 20 figs 5, 8, 16). X–Y. Oral ossicles (NHMUK EE 13606– 13607, the originals of Gale 2011a: pl. 20 figs 10, 14). Provenance: A–J. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; see Hess 2014). K–Y. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). Abbreviations: d = distal; p = proximal. See Fig. 3 for detailed morphological terminology. Scale bars: A–J = 0.2 mm; K–Y = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5036870" httpUri="https://zenodo.org/record/5036870/files/figure.png" pageId="27" pageNumber="176">Fig. 13O</figureCitation>
|
||
) with broad actinal face, bearing large rugosities for attachment of
|
||
<emphasis id="4BBDCE63716DFFA9FD1F7241FCDD2559" bold="true" box="[744,781,1351,1377]" italics="true" pageId="27" pageNumber="176">sos</emphasis>
|
||
; large elongate
|
||
<emphasis id="4BBDCE63716DFFA9FC3E7240FBC72559" bold="true" box="[969,1047,1350,1377]" italics="true" pageId="27" pageNumber="176">oradm</emphasis>
|
||
, low, broad apophyse.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31D341FA716DFFA9FF4A728BFEAE263A" pageId="27" pageNumber="176" type="discussion">
|
||
<paragraph id="79761271716DFFA9FF4A728BFEFF259F" blockId="27.[189,303,1421,1447]" box="[189,303,1421,1447]" pageId="27" pageNumber="176">
|
||
<heading id="223EA51D716DFFA9FF4A728BFEFF259F" bold="true" box="[189,303,1421,1447]" fontSize="11" level="3" pageId="27" pageNumber="176" reason="3">
|
||
<emphasis id="4BBDCE63716DFFA9FF4A728BFEFF259F" bold="true" box="[189,303,1421,1447]" pageId="27" pageNumber="176">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="79761271716DFFA9FF4A72C6FEAE263A" blockId="27.[189,1399,1472,1538]" pageId="27" pageNumber="176">
|
||
<taxonomicName id="BEC969F2716DFFA9FF4A72C6FE7A25E2" authority="Gale & Jagt, 2021" authorityName="Gale & Jagt" authorityYear="2021" box="[189,426,1472,1498]" class="Asteroidea" family="Goniopectinidae" genus="Chrispaulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paxillosida" pageId="27" pageNumber="176" phylum="Echinodermata" rank="species" species="spinosa" status="sp. nov.">
|
||
<emphasis id="4BBDCE63716DFFA9FF4A72C6FE7A25E2" box="[189,426,1472,1498]" italics="true" pageId="27" pageNumber="176">Chrispaulia spinosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="508E7318716DFFA9FE4672C7FDDC25E3" box="[433,524,1473,1499]" pageId="27" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from its congeners in its possession of 3–4 bifid spine pits on distal inferomarginals.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |