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<document id="1ADB1E3106D0A7D3A00B7348AC53091A" ID-DOI="10.18590/euscorpius.2006.vol2006.iss34.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="4673157" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617911401397" checkinUser="carolina" docAuthor="Kovařík, František" docDate="2006" docId="9B7687CA0F5EFFEF6C94F9FE3E2A133F" docLanguage="en" docName="Euscorpius.34.1-16.pdf" docOrigin="Euscorpius 34 (34)" docSource="https://mds.marshall.edu/euscorpius/vol2006/iss34/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Buthus intumescens" docType="treatment" docVersion="3" lastPageNumber="11" masterDocId="674FFFB20F55FFE36C52FF283D43155A" masterDocTitle="Review of Tunisian species of the genus Buthus with descriptions of two new species and a discussion of Ehrenberg s types (Scorpiones: Buthidae)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="10" updateTime="1698902796642" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="DB1874174E8624F0B72A5985AAE0837C">Review of Tunisian species of the genus Buthus with descriptions of two new species and a discussion of Ehrenberg s types (Scorpiones: Buthidae)</mods:title>
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<mods:namePart id="5526FF42DD9219C38357327563DBF3F7">Kovařík, František</mods:namePart>
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<mods:title id="18543B5DF59BBF204282871830A1B6E3">Euscorpius</mods:title>
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<mods:part id="1AB384CDF4A026767D457FF67FB9A94B">
<mods:date id="6580BB0E7308A1BFACA14FA812C6ADC2">2006</mods:date>
<mods:detail id="367E6048268D95F5451462219F9B7A3B" type="pubDate">
<mods:number id="4820C58AD2F9815F6F46579CBA858FC1">2006-12-31</mods:number>
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<mods:number id="B382EF4D7BC5B164D1EC59B9085F635B">34</mods:number>
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<mods:number id="8D4819C9A4234F647B52DBE5C8C40893">34</mods:number>
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<mods:classification id="90253BCD7931348D5EC349C4192F9E4B">journal article</mods:classification>
<mods:identifier id="7BBB3262490A79BEE883CD28ED50526F" type="DOI">10.18590/euscorpius.2006.vol2006.iss34.1</mods:identifier>
<mods:identifier id="DE251989606C112B37A9FF75612D2E60" type="ISSN">1536-9307</mods:identifier>
<mods:identifier id="01D4AA3A9B4FB55098EF969F5992C652" type="Zenodo-Dep">4673157</mods:identifier>
<mods:identifier id="B5694DE423A910B8CC11C5D60A20C197" type="ZooBank">59572554-C3AE-41B5-B6BB-3F2384FF3619</mods:identifier>
</mods:mods>
<treatment id="9B7687CA0F5EFFEF6C94F9FE3E2A133F" LSID="urn:lsid:plazi:treatment:9B7687CA0F5EFFEF6C94F9FE3E2A133F" httpUri="http://treatment.plazi.org/id/9B7687CA0F5EFFEF6C94F9FE3E2A133F" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10">
<subSubSection id="5BC565570F5EFFE86C94F9FE3F261250" pageId="11" pageNumber="10" type="nomenclature">
<paragraph id="136036DC0F5EFFE86C94F9FE3E5F13B4" blockId="11.[198,796,1750,1829]" box="[198,796,1750,1774]" pageId="11" pageNumber="10">
<heading id="482881B00F5EFFE86C94F9FE3E5F13B4" box="[198,796,1750,1774]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="10" reason="4">
<taxonomicName id="D4DF4D5F0F5EFFE86C94F9FE3CE613B4" ID-CoL="68TTV" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[198,421,1750,1774]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="intumescens">
<emphasis id="21ABEACE0F5EFFE86C94F9FE3CE613B4" bold="true" box="[198,421,1750,1774]" italics="true" pageId="11" pageNumber="10">Buthus intumescens</emphasis>
</taxonomicName>
(Ehrenberg in Hemprich &amp; Ehren-
</heading>
</paragraph>
<paragraph id="136036DC0F5EFFE86D2EF9DB3F261250" blockId="11.[198,796,1750,1829]" box="[380,613,1779,1802]" pageId="11" pageNumber="10">
<heading id="482881B00F5EFFE86D2EF9DB3F261250" box="[380,613,1779,1802]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="10" reason="2">
berg, 1829),
<emphasis id="21ABEACE0F5EFFE86E57F9DB3F261250" bold="true" box="[517,613,1779,1802]" pageId="11" pageNumber="10">comb. n.</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5BC565570F5EFFE86D90F826384E10D4" pageId="11" pageNumber="10" type="description">
<paragraph id="136036DC0F5EFFE86D90F8263F63127F" blockId="11.[198,796,1750,1829]" box="[450,544,1806,1829]" pageId="11" pageNumber="10">
(
<figureCitation id="8BE42A590F5EFFE86D98F8263F54127F" box="[458,535,1806,1829]" captionStart="Figure 20" captionStartId="16.[192,268,930,953]" captionTargetBox="[191,1440,191,886]" captionTargetId="figure-11@16.[192,1439,192,885]" captionTargetPageId="16" captionText="Figure 20: Buthus intumescens (Ehrenberg in Hemprich &amp; Ehrenberg, 1829), comb. n., ♀, holotype, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4673225" httpUri="https://zenodo.org/record/4673225/files/figure.png" pageId="11" pageNumber="10">Fig. 20</figureCitation>
)
</paragraph>
<paragraph id="136036DC0F5EFFE86C92F86D3F2C1220" blockId="11.[192,802,1861,1914]" pageId="11" pageNumber="10">
<taxonomicName id="D4DF4D5F0F5EFFE86C92F86D3E611206" authority="Ehrenberg" authorityName=": Hemprich &amp; Ehrenberg" authorityYear="1831" box="[192,802,1861,1884]" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="tunetanus" subGenus="Leiurus" subSpecies="intumescens">
<emphasis id="21ABEACE0F5EFFE86C92F86D3C0B1206" box="[192,328,1861,1884]" italics="true" pageId="11" pageNumber="10">Androctonus</emphasis>
(
<emphasis id="21ABEACE0F5EFFE86D0AF86D3CE91206" box="[344,426,1861,1884]" italics="true" pageId="11" pageNumber="10">Leiurus</emphasis>
)
<emphasis id="21ABEACE0F5EFFE86DE8F86D3FE81206" box="[442,683,1861,1884]" italics="true" pageId="11" pageNumber="10">tunetanus intumescens</emphasis>
Ehrenberg
</taxonomicName>
in Hemprich &amp; Ehrenberg, 1829: 354.
</paragraph>
<paragraph id="136036DC0F5EFFE86F6CFBF6384E104E" blockId="11.[830,1440,1246,1545]" pageId="11" pageNumber="10">
<taxonomicName id="D4DF4D5F0F5EFFE86F6CFBF63967104E" authority=": Hemprich &amp; Ehrenberg, 1831" authorityName=": Hemprich &amp; Ehrenberg" authorityYear="1831" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="tunetanus" subGenus="Liurus" subSpecies="intumescens">
<emphasis id="21ABEACE0F5EFFE86F6CFBF63E8511AF" box="[830,966,1246,1269]" italics="true" pageId="11" pageNumber="10">Androctonus</emphasis>
(
<emphasis id="21ABEACE0F5EFFE86F8BFBF6395C11AF" box="[985,1055,1246,1269]" italics="true" pageId="11" pageNumber="10">Liurus</emphasis>
)
<emphasis id="21ABEACE0F5EFFE86863FBF6386711AF" box="[1073,1316,1246,1269]" italics="true" pageId="11" pageNumber="10">tunetanus intumescens</emphasis>
: Hemprich &amp; Ehrenberg, 1831
</taxonomicName>
(pages unnumbered).
</paragraph>
<paragraph id="136036DC0F5EFFE86F6CFA33387D100B" blockId="11.[830,1440,1246,1545]" pageId="11" pageNumber="10">
<taxonomicName id="D4DF4D5F0F5EFFE86F6CFA3338E31068" authority="Ehrenberg" authorityName=": Hemprich &amp; Ehrenberg" authorityYear="1831" box="[830,1440,1307,1330]" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="tunetanus" subGenus="Leiurus" subSpecies="intermedius">
<emphasis id="21ABEACE0F5EFFE86F6CFA333E841068" box="[830,967,1307,1330]" italics="true" pageId="11" pageNumber="10">Androctonus</emphasis>
(
<emphasis id="21ABEACE0F5EFFE86F8AFA3339691068" box="[984,1066,1307,1330]" italics="true" pageId="11" pageNumber="10">Leiurus</emphasis>
)
<emphasis id="21ABEACE0F5EFFE86869FA33386A1068" box="[1083,1321,1307,1330]" italics="true" pageId="11" pageNumber="10">tunetanus intermedius</emphasis>
Ehrenberg
</taxonomicName>
in Hemprich &amp; Ehrenberg, 1829: 355,
<emphasis id="21ABEACE0F5EFFE868A7FA12387D100B" bold="true" box="[1269,1342,1338,1361]" pageId="11" pageNumber="10">syn. n.</emphasis>
</paragraph>
<paragraph id="136036DC0F5EFFE86F6CFA70384E10D4" blockId="11.[830,1440,1246,1545]" pageId="11" pageNumber="10">
<taxonomicName id="D4DF4D5F0F5EFFE86F6CFA70396710D4" authority=": Hemprich &amp; Ehrenberg, 1831" authorityName=": Hemprich &amp; Ehrenberg" authorityYear="1831" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="tunetanus" subGenus="Liurus" subSpecies="intermedius">
<emphasis id="21ABEACE0F5EFFE86F6CFA703E851035" box="[830,966,1368,1391]" italics="true" pageId="11" pageNumber="10">Androctonus</emphasis>
(
<emphasis id="21ABEACE0F5EFFE86F88FA7039631035" box="[986,1056,1368,1391]" italics="true" pageId="11" pageNumber="10">Liurus</emphasis>
)
<emphasis id="21ABEACE0F5EFFE86861FA7038671035" box="[1075,1316,1368,1391]" italics="true" pageId="11" pageNumber="10">tunetanus intermedius</emphasis>
: Hemprich &amp; Ehrenberg, 1831
</taxonomicName>
(pages unnumbered).
</paragraph>
</subSubSection>
<subSubSection id="5BC565570F5EFFE86F6CFABE39921353" pageId="11" pageNumber="10" type="reference_group">
<paragraph id="136036DC0F5EFFE86F6CFABE39021091" blockId="11.[830,1440,1246,1545]" pageId="11" pageNumber="10">
<taxonomicName id="D4DF4D5F0F5EFFE86F6CFABE3EA81091" ID-CoL="NYZF" authority="Shulov &amp; Amitai, 1959: 219" authorityName="Shulov &amp; Amitai" authorityPageNumber="219" authorityYear="1959" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="variety" species="occitanus" subSpecies="mardochei" variety="israelis">
<emphasis id="21ABEACE0F5EFFE86F6CFABE392D10F7" box="[830,1134,1430,1453]" italics="true" pageId="11" pageNumber="10">Buthus occitanus mardochei</emphasis>
var.
<emphasis id="21ABEACE0F5EFFE868F7FABE39B710F7" box="[1189,1268,1430,1453]" italics="true" pageId="11" pageNumber="10">israelis</emphasis>
Shulov &amp; Amitai, 1959: 219
</taxonomicName>
,
<emphasis id="21ABEACE0F5EFFE86FAAFA9C39021091" bold="true" box="[1016,1089,1460,1483]" pageId="11" pageNumber="10">
<taxonomicNameLabel id="3A9857B50F5EFFE86FAAFA9C39021091" box="[1016,1089,1460,1483]" pageId="11" pageNumber="10">syn. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="136036DC0F5EFFE86F6CFAFB39921353" blockId="11.[830,1440,1246,1545]" pageId="11" pageNumber="10">
<taxonomicName id="D4DF4D5F0F5EFFE86F6CFAFB38D910B0" ID-CoL="NYZF" authority=": Levy &amp; Amitai, 1980: 16" authorityName=": Levy &amp; Amitai" authorityPageNumber="16" authorityYear="1980" box="[830,1434,1491,1514]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="occitanus" subSpecies="israelis">
<emphasis id="21ABEACE0F5EFFE86F6CFAFB391A10B0" box="[830,1113,1491,1514]" italics="true" pageId="11" pageNumber="10">Buthus occitanus israelis</emphasis>
: Levy &amp; Amitai, 1980: 16
</taxonomicName>
, figs. 25–29; Fet &amp; Lowe, 2000: 95.
</paragraph>
</subSubSection>
<subSubSection id="5BC565570F5EFFEF6F6CF9073E2A133F" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10" type="materials_examined">
<paragraph id="136036DC0F5EFFE86F6CF90739E31247" blockId="11.[830,1442,1583,1913]" pageId="11" pageNumber="10">
<emphasis id="21ABEACE0F5EFFE86F6CF9073948131C" box="[830,1035,1583,1606]" italics="true" pageId="11" pageNumber="10">Material examined</emphasis>
:
<materialsCitation id="A3B73C810F5EFFE86849F9073EF413D9" collectionCode="ZMHB" collectorName="F. W. Hemprich &amp; C. G. Ehrenberg" country="Egypt" location="Egypt" pageId="11" pageNumber="10" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="21ABEACE0F5EFFE86849F9073925131C" bold="true" box="[1051,1126,1583,1606]" pageId="11" pageNumber="10">
<collectingCountry id="6BC8764C0F5EFFE86849F9073922131C" box="[1051,1121,1583,1606]" name="Egypt" pageId="11" pageNumber="10">Egypt</collectingCountry>
,
</emphasis>
<specimenCount id="05D9FD550F5EFFE8683DF90739D3131C" box="[1135,1168,1583,1606]" pageId="11" pageNumber="10" type="female">1♀</specimenCount>
, leg.
<collectorName id="BE2A530A0F5EFFE86883F90738C0131C" box="[1233,1411,1583,1606]" pageId="11" pageNumber="10">F. W. Hemprich</collectorName>
&amp;
<collectorName id="BE2A530A0F5EFFE86F6CF9663EB0133F" box="[830,1011,1614,1637]" pageId="11" pageNumber="10">C. G. Ehrenberg</collectorName>
(
<collectionCode id="75CEAE190F5EFFE86851F9663911133F" box="[1027,1106,1614,1637]" pageId="11" pageNumber="10">ZMHB</collectionCode>
No. 145,
<typeStatus id="CC64887E0F5EFFE86896F9663862133F" box="[1220,1313,1614,1637]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
of
<emphasis id="21ABEACE0F5EFFE86919F9663EEF13D9" italics="true" pageId="11" pageNumber="10">
A. t.
<taxonomicName id="D4DF4D5F0F5EFFE869D1F9663EEF13D9" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="intumescens">intumescens</taxonomicName>
</emphasis>
)
</materialsCitation>
.
<emphasis id="21ABEACE0F5EFFE86F96F9443E8C13D9" bold="true" box="[964,975,1644,1667]" pageId="11" pageNumber="10">?</emphasis>
<materialsCitation id="A3B73C810F5EFFE86F9DF9443E351385" collectionCode="ZMHB" collectorName="F. W. Hemprich &amp; C. G. Ehrenberg" country="Yemen" location="Lohaie" municipality="Al Luhayyah" pageId="11" pageNumber="10" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="21ABEACE0F5EFFE86F9DF944396413D9" bold="true" box="[975,1063,1644,1667]" pageId="11" pageNumber="10">
<collectingCountry id="6BC8764C0F5EFFE86F9DF944396013D9" box="[975,1059,1644,1667]" name="Yemen" pageId="11" pageNumber="10">Yemen</collectingCountry>
,
</emphasis>
<location id="160060070F5EFFE86862F944393913D9" LSID="urn:lsid:plazi:treatment:9B7687CA0F5EFFEF6C94F9FE3E2A133F:160060070F5EFFE86862F944393913D9" box="[1072,1146,1644,1667]" country="Yemen" municipality="Al Luhayyah" name="Lohaie" pageId="11" pageNumber="10">Lohaie</location>
, now
<collectingMunicipality id="F304ACA60F5EFFE868EDF944380D13D9" box="[1215,1358,1644,1667]" pageId="11" pageNumber="10">Al Luhayyah</collectingMunicipality>
, probably incorrect locality,
<specimenCount id="05D9FD550F5EFFE8687DF9A3391313F8" box="[1071,1104,1675,1698]" pageId="11" pageNumber="10" type="female">1♀</specimenCount>
, leg.
<collectorName id="BE2A530A0F5EFFE868DDF9A3380313F8" box="[1167,1344,1675,1698]" pageId="11" pageNumber="10">F. W. Hemprich</collectorName>
&amp;
<collectorName id="BE2A530A0F5EFFE86937F9A33EED139B" pageId="11" pageNumber="10">C. G. Ehrenberg</collectorName>
(
<collectionCode id="75CEAE190F5EFFE86FEDF982394D139B" box="[959,1038,1706,1729]" pageId="11" pageNumber="10">ZMHB</collectionCode>
No. 146,
<typeStatus id="CC64887E0F5EFFE868D1F98239A3139B" box="[1155,1248,1706,1729]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
of
<emphasis id="21ABEACE0F5EFFE8695EF9823E291385" italics="true" pageId="11" pageNumber="10">A. t. intermedius</emphasis>
)
</materialsCitation>
.
<materialsCitation id="A3B73C810F5EFFE86FD0F9E039DF1247" collectingDate="2004-11" collectingDateMax="2004-12-31" collectingDateMin="2004-11" collectionCode="FKCP" collectorName="J. Kral" country="Israel" county="Sede Boqer" location="Negev" municipality="Vadi Hazaz" pageId="11" pageNumber="10" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="holotype">
<emphasis id="21ABEACE0F5EFFE86FD0F9E03E801385" bold="true" box="[898,963,1736,1759]" pageId="11" pageNumber="10">
<collectingCountry id="6BC8764C0F5EFFE86FD0F9E03E801385" box="[898,963,1736,1759]" name="Israel" pageId="11" pageNumber="10">Israel</collectingCountry>
</emphasis>
, south part of
<location id="160060070F5EFFE8680DF9E039E61385" LSID="urn:lsid:plazi:treatment:9B7687CA0F5EFFEF6C94F9FE3E2A133F:160060070F5EFFE8680DF9E039E61385" box="[1119,1189,1736,1759]" country="Israel" county="Sede Boqer" municipality="Vadi Hazaz" name="Negev" pageId="11" pageNumber="10">Negev</location>
,
<collectingMunicipality id="F304ACA60F5EFFE868E1F9E038731385" box="[1203,1328,1736,1759]" pageId="11" pageNumber="10">Vadi Hazaz</collectingMunicipality>
near
<collectingCounty id="FA014E500F5EFFE8693EF9E03EC313A4" pageId="11" pageNumber="10">Sede Boqer</collectingCounty>
(Haluqim Ridge),
<date id="6761101C0F5EFFE86818F9CF383713A4" box="[1098,1396,1767,1790]" pageId="11" pageNumber="10" value="2004-11" valueMax="2004-12-31" valueMin="2004-11-00">
<collectingDate id="7725E9F40F5EFFE86818F9CF383713A4" box="[1098,1396,1767,1790]" pageId="11" pageNumber="10" value="2004-11" valueMax="2004-12-31" valueMin="2004-11-00">November–December 2004</collectingDate>
</date>
,
<specimenCount id="05D9FD550F5EFFE8692DF9CF38E313A4" box="[1407,1440,1767,1790]" pageId="11" pageNumber="10" type="male">1♂</specimenCount>
<specimenCount id="05D9FD550F5EFFE86F6CF82E3E231247" box="[830,864,1798,1821]" pageId="11" pageNumber="10" type="female">3♀</specimenCount>
, 5 juvs., leg.
<collectorName id="BE2A530A0F5EFFE86FA4F82E397D1247" box="[1014,1086,1798,1821]" pageId="11" pageNumber="10">J. Král</collectorName>
(
<collectionCode id="75CEAE190F5EFFE8681EF82E39D61247" box="[1100,1173,1798,1821]" pageId="11" pageNumber="10">FKCP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="136036DC0F5EFFEF6F6CF80C3F41104E" blockId="11.[830,1442,1583,1913]" lastBlockId="12.[192,803,1124,1883]" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10">
<emphasis id="21ABEACE0F5EFFE86F6CF80C3EEC1261" box="[830,943,1828,1851]" italics="true" pageId="11" pageNumber="10">Comments</emphasis>
. It was important to examine the type of
<taxonomicName id="D4DF4D5F0F5EFFE8692EF80C38C01200" authority="Ehrenberg" authorityName=": Hemprich &amp; Ehrenberg" authorityYear="1831" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="tunetanus" subGenus="Leiurus" subSpecies="intumescens">
<emphasis id="21ABEACE0F5EFFE8692EF80C3EEA1200" italics="true" pageId="11" pageNumber="10">Androctonus</emphasis>
(
<emphasis id="21ABEACE0F5EFFE86FE8F86B39481200" box="[954,1035,1859,1882]" italics="true" pageId="11" pageNumber="10">Leiurus</emphasis>
)
<emphasis id="21ABEACE0F5EFFE8684EF86B384E1200" box="[1052,1293,1859,1882]" italics="true" pageId="11" pageNumber="10">tunetanus intumescens</emphasis>
Ehrenberg
</taxonomicName>
in Hemprich &amp; Ehrenberg, 1829, hitherto considered a syn- onym of
<taxonomicName id="D4DF4D5F0F59FFEF6D71FB4C3C9D1121" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[291,478,1124,1147]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="tunetanus">
<emphasis id="21ABEACE0F59FFEF6D71FB4C3C9D1121" box="[291,478,1124,1147]" italics="true" pageId="12" pageNumber="11">Buthus tunetanus</emphasis>
</taxonomicName>
(see Fet &amp; Lowe, 2000: 97), to verify that it is not one of the species proposed in this study as new. The
<typeStatus id="CC64887E0F59FFEF6DDCFB893CA911E2" box="[398,490,1185,1208]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
in question is a female (
<figureCitation id="8BE42A590F59FFEF6EA4FB893D9E118D" captionStart="Figure 20" captionStartId="16.[192,268,930,953]" captionTargetBox="[191,1440,191,886]" captionTargetId="figure-11@16.[192,1439,192,885]" captionTargetPageId="16" captionText="Figure 20: Buthus intumescens (Ehrenberg in Hemprich &amp; Ehrenberg, 1829), comb. n., ♀, holotype, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4673225" httpUri="https://zenodo.org/record/4673225/files/figure.png" pageId="12" pageNumber="11">Fig. 20</figureCitation>
) from
<collectingCountry id="6BC8764C0F59FFEF6D60FBE83C31118D" box="[306,370,1216,1239]" name="Egypt" pageId="12" pageNumber="11">Egypt</collectingCountry>
(
<collectionCode id="75CEAE190F59FFEF6DD5FBE83C95118D" box="[391,470,1216,1239]" pageId="12" pageNumber="11">ZMHB</collectionCode>
No. 145) and has the chela markedly narrower than
<taxonomicName id="D4DF4D5F0F59FFEF6D86FBF63F1D11AF" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[468,606,1246,1269]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="tunetanus">
<emphasis id="21ABEACE0F59FFEF6D86FBF63F1D11AF" box="[468,606,1246,1269]" italics="true" pageId="12" pageNumber="11">B. tunetanus</emphasis>
</taxonomicName>
; its movable fingers bear 11 rows of granules.
</paragraph>
<caption id="47A066540F59FFEF6C92FB263CE11164" ID-DOI="http://doi.org/10.5281/zenodo.4673200" ID-Zenodo-Dep="4673200" httpUri="https://zenodo.org/record/4673200/files/figure.png" pageId="12" pageNumber="11" startId="12.[192,268,1038,1061]" targetBox="[191,1440,191,1006]" targetPageId="12">
<paragraph id="136036DC0F59FFEF6C92FB263CE11164" blockId="12.[192,1440,1038,1086]" pageId="12" pageNumber="11">
<emphasis id="21ABEACE0F59FFEF6C92FB263C7B117F" bold="true" box="[192,312,1038,1061]" pageId="12" pageNumber="11">Figure 13:</emphasis>
Locality of
<taxonomicName id="D4DF4D5F0F59FFEF6DE4FB383F1D117F" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[438,606,1040,1061]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="tunetanus">
<emphasis id="21ABEACE0F59FFEF6DE4FB383F1D117F" box="[438,606,1040,1061]" italics="true" pageId="12" pageNumber="11">Buthus tunetanus</emphasis>
</taxonomicName>
(Tunisia, Beja Province, Jebel Sabbah Mts., Bou Salem, 10 km NW Balta, 27 May 2005 and 2 June 2005).
</paragraph>
</caption>
<paragraph id="136036DC0F59FFEF6CA2FA343E3B11C3" blockId="12.[192,803,1124,1883]" lastBlockId="12.[830,1441,1124,1637]" pageId="12" pageNumber="11">
I have not been able to find any differences between this type, the
<typeStatus id="CC64887E0F59FFEF6D00FA123CF3100B" box="[338,432,1338,1361]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
of
<taxonomicName id="D4DF4D5F0F59FFEF6D8EFA123C29102A" authorityName=": Hemprich &amp; Ehrenberg" authorityYear="1831" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="tunetanus" subGenus="Leiurus" subSpecies="intermedius">
<emphasis id="21ABEACE0F59FFEF6D8EFA123F27100B" box="[476,612,1338,1361]" italics="true" pageId="12" pageNumber="11">Androctonus</emphasis>
(
<emphasis id="21ABEACE0F59FFEF6E26FA123F86100B" box="[628,709,1338,1361]" italics="true" pageId="12" pageNumber="11">Leiurus</emphasis>
)
<emphasis id="21ABEACE0F59FFEF6E84FA123C29102A" italics="true" pageId="12" pageNumber="11">tunetanus intermedius</emphasis>
</taxonomicName>
, and
<taxonomicName id="D4DF4D5F0F59FFEF6DF4FA713CB3102A" authorityName="Leach" authorityYear="1815" box="[422,496,1369,1392]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="21ABEACE0F59FFEF6DF4FA713CB3102A" box="[422,496,1369,1392]" italics="true" pageId="12" pageNumber="11">Buthus</emphasis>
</taxonomicName>
specimens from
<collectingCountry id="6BC8764C0F59FFEF6EFAFA713FA0102A" box="[680,739,1369,1392]" name="Israel" pageId="12" pageNumber="11">Israel</collectingCountry>
identified as
<taxonomicName id="D4DF4D5F0F59FFEF6D76FA503F7B10D5" authorityName=": Levy &amp; Amitai" authorityYear="1980" box="[292,568,1400,1423]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="occitanus" subSpecies="israelis">
<emphasis id="21ABEACE0F59FFEF6D76FA503F7B10D5" box="[292,568,1400,1423]" italics="true" pageId="12" pageNumber="11">Buthus occitanus israelis</emphasis>
</taxonomicName>
. The inevitable conclusion is that
<taxonomicName id="D4DF4D5F0F59FFEF6D36FABE3F7810F7" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[356,571,1430,1453]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="intumescens">
<emphasis id="21ABEACE0F59FFEF6D36FABE3F7810F7" box="[356,571,1430,1453]" italics="true" pageId="12" pageNumber="11">Buthus intumescens</emphasis>
</taxonomicName>
(Ehrenberg in Hemprich &amp; Ehrenberg, 1829), comb. n. is a valid species occurring in
<collectingCountry id="6BC8764C0F59FFEF6D19FAFC3CC810B1" box="[331,395,1492,1515]" name="Egypt" pageId="12" pageNumber="11">Egypt</collectingCountry>
(
<collectingRegion id="D11BF83E0F59FFEF6DCEFAFC3C9710B1" box="[412,468,1492,1515]" country="Egypt" name="Shamal Sina'" pageId="12" pageNumber="11">Sinai</collectingRegion>
) and
<collectingCountry id="6BC8764C0F59FFEF6E45FAFC3F1110B1" box="[535,594,1492,1515]" name="Israel" pageId="12" pageNumber="11">Israel</collectingCountry>
, and its synonyms are
<taxonomicName id="D4DF4D5F0F59FFEF6CB9FADA3C0D1372" authority="Shulov &amp; Amitai, 1959" authorityName="Shulov &amp; Amitai" authorityYear="1959" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="variety" species="occitanus" subSpecies="mardochei" variety="israelis">
<emphasis id="21ABEACE0F59FFEF6CB9FADA3F621353" box="[235,545,1522,1545]" italics="true" pageId="12" pageNumber="11">Buthus occitanus mardochei</emphasis>
var.
<emphasis id="21ABEACE0F59FFEF6E0DFADA3FEE1353" box="[607,685,1522,1545]" italics="true" pageId="12" pageNumber="11">israelis</emphasis>
Shulov &amp; Amitai, 1959
</taxonomicName>
,
<taxonomicNameLabel id="3A9857B50F59FFEF6D0FF9393CE61372" box="[349,421,1553,1576]" pageId="12" pageNumber="11">syn. n.</taxonomicNameLabel>
and
<taxonomicName id="D4DF4D5F0F59FFEF6D8EF9393CAA131D" authority="Ehrenberg" authorityName=": Hemprich &amp; Ehrenberg" authorityYear="1831" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="tunetanus" subGenus="Leiurus" subSpecies="intermedius">
<emphasis id="21ABEACE0F59FFEF6D8EF9393F201372" box="[476,611,1553,1576]" italics="true" pageId="12" pageNumber="11">Androctonus</emphasis>
(
<emphasis id="21ABEACE0F59FFEF6E26F9393F851372" box="[628,710,1553,1576]" italics="true" pageId="12" pageNumber="11">Leiurus</emphasis>
)
<emphasis id="21ABEACE0F59FFEF6E84F9393C2C131D" italics="true" pageId="12" pageNumber="11">tunetanus intermedius</emphasis>
Ehrenberg
</taxonomicName>
in Hemprich &amp; Ehrenberg, 1829, syn. n. The
<typeStatus id="CC64887E0F59FFEF6DD1F9663CA3133F" box="[387,480,1614,1637]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
of the latter taxon is a female labeled as from Lohaie (
<collectionCode id="75CEAE190F59FFEF6D9DF9453F5D13DE" box="[463,542,1645,1668]" pageId="12" pageNumber="11">ZMHB</collectionCode>
No. 146). Lohaie, now Al Luhayyah,
<collectingCountry id="6BC8764C0F59FFEF6D0DF9A43CEF13F9" box="[351,428,1676,1699]" name="Yemen" pageId="12" pageNumber="11">Yemen</collectingCountry>
, probably is an incorrect locality, because no specimen of
<taxonomicName id="D4DF4D5F0F59FFEF6D98F9823F50139B" authorityName="Leach" authorityYear="1815" box="[458,531,1706,1729]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="21ABEACE0F59FFEF6D98F9823F50139B" box="[458,531,1706,1729]" italics="true" pageId="12" pageNumber="11">Buthus</emphasis>
</taxonomicName>
from the Arabian Peninsula has ever been confirmed (Braunwalder &amp; Fet, 1998: 34; Fet &amp; Lowe, 2000: 94). Although
<taxonomicName id="D4DF4D5F0F59FFEF6E0DF9C03CB91247" authorityName=": Hemprich &amp; Ehrenberg" authorityYear="1831" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="tunetanus" subGenus="Leiurus" subSpecies="intumescens">
<emphasis id="21ABEACE0F59FFEF6E0DF9C03FA513A5" box="[607,742,1768,1791]" italics="true" pageId="12" pageNumber="11">Androctonus</emphasis>
(
<emphasis id="21ABEACE0F59FFEF6EA5F9C03DB31247" italics="true" pageId="12" pageNumber="11">Leiurus</emphasis>
)
<emphasis id="21ABEACE0F59FFEF6D57F82E3CB91247" box="[261,506,1798,1821]" italics="true" pageId="12" pageNumber="11">tunetanus intumescens</emphasis>
</taxonomicName>
and
<taxonomicName id="D4DF4D5F0F59FFEF6E68F82E3C6A1266" authorityName=": Hemprich &amp; Ehrenberg" authorityYear="1831" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="tunetanus" subGenus="Leiurus" subSpecies="intermedius">
<emphasis id="21ABEACE0F59FFEF6E68F82E3F121247" box="[570,593,1798,1821]" italics="true" pageId="12" pageNumber="11">A.</emphasis>
(
<emphasis id="21ABEACE0F59FFEF6E34F82E3F3F1247" box="[614,636,1798,1821]" italics="true" pageId="12" pageNumber="11">L.</emphasis>
)
<emphasis id="21ABEACE0F59FFEF6EC2F82E3C6A1266" italics="true" pageId="12" pageNumber="11">tunetanus intermedius</emphasis>
</taxonomicName>
are described in the same work, the description of
<emphasis id="21ABEACE0F59FFEF6D46F86C3C8D1201" box="[276,462,1860,1883]" italics="true" pageId="12" pageNumber="11">
A. t.
<taxonomicName id="D4DF4D5F0F59FFEF6D1EF86C3C8D1201" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[332,462,1860,1883]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="intumescens">intumescens</taxonomicName>
</emphasis>
(p. 354) precedes that of
<emphasis id="21ABEACE0F59FFEF6EA1F86C3EFF1121" box="[755,956,1124,1883]" italics="true" pageId="12" pageNumber="11">A. t. intermedius</emphasis>
(p. 355), thus the first name must have priority.
</paragraph>
<paragraph id="136036DC0F59FFEF6F3CFB893E2A133F" blockId="12.[830,1441,1124,1637]" pageId="12" pageNumber="11">
In case of
<taxonomicName id="D4DF4D5F0F59FFEF6FB7FB89390B118D" authority="Shulov &amp; Amitai, 1959" authorityName="Shulov &amp; Amitai" authorityYear="1959" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="variety" species="occitanus" subSpecies="mardochei" variety="israelis">
<emphasis id="21ABEACE0F59FFEF6FB7FB89385411E2" box="[997,1303,1185,1208]" italics="true" pageId="12" pageNumber="11">Buthus occitanus mardochei</emphasis>
var.
<emphasis id="21ABEACE0F59FFEF6901FB8938DC11E2" box="[1363,1439,1185,1208]" italics="true" pageId="12" pageNumber="11">israelis</emphasis>
Shulov &amp; Amitai, 1959
</taxonomicName>
, I have not studied
<typeStatus id="CC64887E0F59FFEF6967FBE8382E118D" box="[1333,1389,1216,1239]" pageId="12" pageNumber="11">types</typeStatus>
and base my conclusions on nine studied specimens from Negev. As noted above, the primary criterion for assessing species of the
<taxonomicName id="D4DF4D5F0F59FFEF6850FA3439FB1069" baseAuthorityName="Amoreux" baseAuthorityYear="1789" box="[1026,1208,1308,1331]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="occitanus">
<emphasis id="21ABEACE0F59FFEF6850FA3439FB1069" box="[1026,1208,1308,1331]" italics="true" pageId="12" pageNumber="11">Buthus occitanus</emphasis>
</taxonomicName>
complex, i.e. all taxa formerly considered subspecies of
<taxonomicName id="D4DF4D5F0F59FFEF688CFA123EBA102A" authority="(Amoreux, 1789)" baseAuthorityName="Amoreux" baseAuthorityYear="1789" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="occitanus">
<emphasis id="21ABEACE0F59FFEF688CFA1238E3100B" box="[1246,1440,1338,1361]" italics="true" pageId="12" pageNumber="11">Buthus occitanus</emphasis>
(Amoreux, 1789)
</taxonomicName>
or of
<taxonomicName id="D4DF4D5F0F59FFEF686FFA7138D9102A" authority="(Herbst, 1800)" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[1085,1434,1369,1392]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="tunetanus">
<emphasis id="21ABEACE0F59FFEF686FFA7139B5102A" box="[1085,1270,1369,1392]" italics="true" pageId="12" pageNumber="11">Buthus tunetanus</emphasis>
(Herbst, 1800)
</taxonomicName>
, is the width of pedipalp chela and its relation to sexual dimorphism. In the case of
<taxonomicName id="D4DF4D5F0F59FFEF6822FABE380610F7" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[1136,1349,1430,1453]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="intumescens">
<emphasis id="21ABEACE0F59FFEF6822FABE380610F7" box="[1136,1349,1430,1453]" italics="true" pageId="12" pageNumber="11">Buthus intumescens</emphasis>
</taxonomicName>
(Ehrenberg in Hemprich &amp; Ehrenberg, 1829), comb. n. the different width of chela shows this taxon to have the status of a species rather than a subspecies (
<figureCitation id="8BE42A590F59FFEF68A3FADA38051353" box="[1265,1350,1522,1545]" captionStart="Figure 20" captionStartId="16.[192,268,930,953]" captionTargetBox="[191,1440,191,886]" captionTargetId="figure-11@16.[192,1439,192,885]" captionTargetPageId="16" captionText="Figure 20: Buthus intumescens (Ehrenberg in Hemprich &amp; Ehrenberg, 1829), comb. n., ♀, holotype, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4673225" httpUri="https://zenodo.org/record/4673225/files/figure.png" pageId="12" pageNumber="11">Fig. 20</figureCitation>
), as the chela is narrower than in
<taxonomicName id="D4DF4D5F0F59FFEF6837F9393E3E131D" authority="(Amoreux, 1789)" baseAuthorityName="Amoreux" baseAuthorityYear="1789" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="occitanus">
<emphasis id="21ABEACE0F59FFEF6837F93938631372" box="[1125,1312,1553,1576]" italics="true" pageId="12" pageNumber="11">Buthus occitanus</emphasis>
(Amoreux, 1789)
</taxonomicName>
and
<taxonomicName id="D4DF4D5F0F59FFEF6FE3F918384E131D" authority="(Herbst, 1800)" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[945,1293,1584,1607]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="tunetanus">
<emphasis id="21ABEACE0F59FFEF6FE3F9183929131D" box="[945,1130,1584,1607]" italics="true" pageId="12" pageNumber="11">Buthus tunetanus</emphasis>
(Herbst, 1800)
</taxonomicName>
(
<figureCitation id="8BE42A590F59FFEF6949F9183830131D" box="[1307,1395,1584,1607]" captionStart="Figure 16" captionStartId="14.[192,268,970,993]" captionTargetBox="[191,1440,191,937]" captionTargetId="figure-10@14.[192,1439,192,936]" captionTargetPageId="14" captionText="Figure 16: Buthus tunetanus, ♂, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4673212" httpUri="https://zenodo.org/record/4673212/files/figure.png" pageId="12" pageNumber="11">Figs. 16</figureCitation>
and
<figureCitation id="8BE42A590F59FFEF6F6DF9663E19133F" box="[831,858,1614,1637]" captionStart="Figure 18" captionStartId="15.[192,268,1049,1072]" captionTargetBox="[191,1440,191,1021]" captionTargetId="figure-6@15.[192,1439,192,1020]" captionTargetPageId="15" captionText="Figure 18: Buthus tunetanus, ♀, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4673218" httpUri="https://zenodo.org/record/4673218/files/figure.png" pageId="12" pageNumber="11">18</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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