213 lines
25 KiB
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213 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="6CB2427055C6516F980825736C3921CC" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole andohahela Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="AD047203-0321-4E69-B9BB-EF246CA61BFC" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>949</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>185</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="165090176" LSID="urn:lsid:zoobank.org:act:AD047203-0321-4E69-B9BB-EF246CA61BFC" httpUri="http://treatment.plazi.org/id/6CB2427055C6516F980825736C3921CC" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="6CB24270-55C6-516F-9808-25736C3921CC" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole andohahela" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="andohahela" status="sp. nov.">Pheidole andohahela</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">Figs 22A-F</figureCitation>
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<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63D</figureCitation>
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<figureCitation captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Toliara; Anosy Region, Parc National Andohahela, Col de Tanatana;
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<geoCoordinate degrees="24.74969" direction="south" orientation="latitude" precision="1" value="-24.74969">-24.74969</geoCoordinate>
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,
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<geoCoordinate degrees="46.84949" direction="east" orientation="longitude" precision="1" value="46.84949">46.84949</geoCoordinate>
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; alt. 400 m; 9 Mar 2015; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF36798; CASENT0700917 (CASC).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
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• 1w.; same data as for holotype, CASENT0923281 (CASC).
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</paragraph>
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<caption doi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1" start="Figure 22" startId="F22">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 22.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole andohahela" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="andohahela">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole andohahela</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0923281) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0700917).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" pageId="0" pageNumber="1">Toliara</emphasis>
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: • 1w., 1s.; Anosy Region, Parc National Andohahela, Col de Tanatana;
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<geoCoordinate degrees="24.74969" direction="south" orientation="latitude" precision="1" value="-24.74969">-24.74969</geoCoordinate>
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,
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<geoCoordinate degrees="46.84949" direction="east" orientation="longitude" precision="1" value="46.84949">46.84949</geoCoordinate>
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; alt. 400 m; 9 Mar 2015; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF36803 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Minute species.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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HL <0.95 mm and WL <0.8 mm; head in full-face view sub-oval; body dark brown; sides of head with very dense, relatively short, suberect pilosity; entire head distinctly sculptured; scape, when laid back, exceeding the midlength of head by two-fifths of its length; inner hypostomal teeth distinct, large, with sharp apex; outer hypostomal teeth lobe-like with apex directed distinctly outward; inner and outer hypostomal teeth not closely spaced and not connected by concavity.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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HL <0.5 mm and WL <0.6 mm, scape, when laid back, surpassing the posterior head margin by two-fifths of its length, propodeal spines reduced to small lobes, head elongate and oval and body dark brown; head sculpture shiny and predominantly smooth, vertex and frons with sparse, short, and transverse rugulae; anterolateral sides of propodeum and anepisternum with indistinct and sparse rugoreticulae.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 2): HL: 0.87, 0.92; HW: 0.87, 0.92; SL: 0.64, 0.63; EL: 0.12, 0.11; WL: 0.77, 0.79; PSL: 0.1, 0.11; MTL: 0.53, 0.56; PNW: 0.36, 0.39; PTW: 0.11, 0.12; PPW: 0.28, 0.29; CI: 100.0, 99.9; SI: 72.9, 67.8; PSLI: 11.9, 12.4; PPI: 40.1, 40.7; PNI: 41.3, 42.0; MTI: 60.8, 60.5.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view sub-oval, slightly widening posteriorly, with anterior and posterior sides convex (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with very dense, relatively short, suberect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with thick, dense, longitudinal, and interrupted rugae, interspaces smooth; lateral sides with thick, dense, and irregular rugae, interspaces shiny with dense and distinct rugoreticulae. Occipital lobes with thick, irregular rugae and smooth to rugoreticulate interspaces. Area posterolateral from eyes rugoreticulate to rugofoveolate with additional longitudinal and thick rugae. Gena with relatively dense, thick, and longitudinal rugae and rugoreticulate interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by two-fifths of its length; pilosity subdecumbent to erect (Fig. B, D). Inner hypostomal teeth distinct, large, closely spaced, triangular, with sharp apex directed upward; outer hypostomal teeth lobe-like, more narrow and lower than inner hypostomal teeth, apex directed outward; inner and outer hypostomal teeth not closely spaced and not connected by concavity (Fig.
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<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63D</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines short, with moderately wide base and acute apex; humeral area laterally weakly produced (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22D</figureCitation>
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). Surface shiny with fine rugofoveolae; anterolateral sides of pronotum with smooth notches. Pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
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Shiny with fine foveolae;node smooth to finely foveolate, low, triangular, with rounded and thin apex, in rear view node dorsoventrally straight to slightly convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
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Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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Shiny and smooth; pilosity dense, long, and erect (Fig. D, F).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Dark brown; legs, gaster and antennae yellowish brown (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22D, F</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 2): HL: 0.48, 0.49; HW: 0.39, 0.43; SL: 0.55, 0.57; EL: 0.09, 0.09; WL: 0.58, 0.58; PSL: 0.05, 0.05; MTL: 0.4, 0.42; PNW: 0.28, 0.28; PTW: 0.07, 0.08; PPW: 0.12, 0.14; CI: 122.4, 115.0; SI: 140.3, 132.8; PSLI: 10.8, 9.4; PPI: 55.6, 55.6; PNI: 70.9, 66.0; MTI: 102.0, 98.4.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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Cephalic margin indistinctly convex or straight (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22A</figureCitation>
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). Pilosity relatively dense, long, decumbent to suberect. Sculpture shiny and smooth; vertex and frons with sparse, short, and transverse rugulae; antennal sockets with few thick, curved outward rugae and smooth interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity dense, subdecumbent to erect (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22A, C</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum moderately high and short, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines reduce to small tubercles (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22C</figureCitation>
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). Sculpture shiny and smooth; anterolateral sides of propodeum and anepisternum with indistinct and sparse rugoreticulae. Pilosity moderately sparse, long, and erect (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22C, E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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With sparse, erect pilosity (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22C, E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Dark brown, legs and antenna yellowish brown (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pheidole andohahela sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923281) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0700917)." figureDoi="10.3897/zookeys.949.51269.figure22" httpUri="https://binary.pensoft.net/fig/432191" pageId="0" pageNumber="1">22C, E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
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<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The species was collected between 400-775 m in elevation, in rainforest. Nests were located in rotten logs and rotten sticks on the ground.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="comments">
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<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole andohahela" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="andohahela">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole andohahela</emphasis>
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</taxonomicName>
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sp. nov. is known only from its terra typica: Col de Tanatana in Parc National Andohahela, Toliara and is most similar to parapatric
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<taxonomicName lsidName="P. lavasoa" pageId="0" pageNumber="1" rank="species" species="lavasoa">
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<emphasis italics="true" pageId="0" pageNumber="1">P. lavasoa</emphasis>
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</taxonomicName>
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sp. nov. recorded so far only from Grand Lavasoa, Toliara. Major workers of both taxa are extremely similar and can be distinguished based on slight differences in the setosity of lateral sides of head (which is sparser and longer in
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<taxonomicName lsidName="P. andohahela" pageId="0" pageNumber="1" rank="species" species="andohahela">
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<emphasis italics="true" pageId="0" pageNumber="1">P. andohahela</emphasis>
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</taxonomicName>
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sp. nov.) and shape of the hypostomal teeth (
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<taxonomicName lsidName="P. andohahela" pageId="0" pageNumber="1" rank="species" species="andohahela">
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<emphasis italics="true" pageId="0" pageNumber="1">P. andohahela</emphasis>
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</taxonomicName>
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sp. nov. has inner hypostomal teeth with sharp apices and outer hypostomal teeth that are lobe-like and distinctly directed outward). Minor workers are a better resource for species separation. Minor workers of
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<taxonomicName lsidName="P. andohahela" pageId="0" pageNumber="1" rank="species" species="andohahela">
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<emphasis italics="true" pageId="0" pageNumber="1">P. andohahela</emphasis>
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</taxonomicName>
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sp. nov. have transverse rugulae on vertex and frons, while anterolateral sides of their propodeum, and anepisternum have indistinct and sparse rugoreticulae; body colour is dark brown.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |