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<document id="1F660F009CF5D4A5D087BE48F4B70C06" ID-CLB-Dataset="26833" ID-DOI="10.11646/zootaxa.4604.2.3" ID-GBIF-Dataset="f5b155f4-c251-4daf-b267-322b95115f91" ID-ISSN="1175-5326" ID-Zenodo-Dep="2807064" ID-ZooBank="0BA07364-39ED-4349-98C5-27431A90CEAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557812978353" checkinUser="plazi" docAuthor="Jaschhof, Mathias &amp; Jaschhof, Catrin" docDate="2019" docId="4C4087808A47FFEC23A46933FDC46B19" docLanguage="en" docName="zootaxa.4604.2.3.pdf" docOrigin="Zootaxa 4604 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Camptomyia Kieffer 1894" docType="treatment" docVersion="7" lastPageNumber="288" masterDocId="B079FFF88A42FFEB23336E06FFB86822" masterDocTitle="New and rarely found species of asynaptine Porricondylinae (Diptera: Cecidomyiidae) in northern Europe" masterLastPageNumber="300" masterPageNumber="281" pageNumber="286" updateTime="1698728559376" updateUser="ExternalLinkService">
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<mods:title id="F980018765D7860496531D2B3ABDCDB0">New and rarely found species of asynaptine Porricondylinae (Diptera: Cecidomyiidae) in northern Europe</mods:title>
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<paragraph id="C45636968A47FFEE23A46933FE2A6F6D" blockId="5.[151,402,1845,1871]" box="[151,402,1845,1871]" pageId="5" pageNumber="286">
<heading id="9F1E81FA8A47FFEE23A46933FE2A6F6D" bold="true" box="[151,402,1845,1871]" fontSize="11" level="1" pageId="5" pageNumber="286" reason="1">
<taxonomicName id="03E94D158A47FFEE23A46933FE2A6F6D" ID-CoL="3GJR" ID-ENA="1068926" authority="Kieffer" authorityName="Kieffer" authorityYear="1894" box="[151,402,1845,1871]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A47FFEE23A46933FE2A6F6D" bold="true" box="[151,402,1845,1871]" pageId="5" pageNumber="286">
<emphasis id="F69DEA848A47FFEE23A46933FE886F6D" bold="true" box="[151,304,1845,1871]" italics="true" pageId="5" pageNumber="286">Camptomyia</emphasis>
Kieffer
</emphasis>
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=
<emphasis id="F69DEA848A47FFEE239F697CFE936FB0" box="[172,299,1914,1938]" italics="true" pageId="5" pageNumber="286">Niladmirara</emphasis>
Fedotova
<emphasis id="F69DEA848A47FFEE22A6697CFE4B6FB0" bold="true" box="[405,499,1914,1938]" pageId="5" pageNumber="286">syn. nov.</emphasis>
</paragraph>
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<paragraph id="C45636968A47FFED23A469C6FC1A6B19" blockId="5.[151,1436,1983,2011]" lastBlockId="6.[151,1437,153,2015]" lastPageId="6" lastPageNumber="287" pageId="5" pageNumber="286">
Major works on
<taxonomicName id="03E94D158A47FFEE226069C7FE5A6FF9" authorityName="Kieffer" authorityYear="1894" box="[339,482,1985,2011]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A47FFEE226069C7FE5A6FF9" box="[339,482,1985,2011]" italics="true" pageId="5" pageNumber="286">Camptomyia</emphasis>
</taxonomicName>
in Europe are those by
<bibRefCitation id="A0784B678A47FFEE21C269C7FC106FF9" author="Mamaev, B. M." box="[753,936,1984,2011]" pageId="5" pageNumber="286" pagination="1677 - 1690" refId="ref13562" refString="Mamaev, B. M. (1961) Gall midges of the USSR. New species of the genus Camptomyia Kieffer (Itonididae, Diptera). Zoologicheskii Zhurnal, 40, 1677 - 1690. [in Russian]" type="journal article" year="1961">Mamaev (1961)</bibRefCitation>
,
<bibRefCitation id="A0784B678A47FFEE208769C6FBD86FF8" author="Panelius, S." box="[948,1120,1983,2010]" pageId="5" pageNumber="286" pagination="1 - 157" refId="ref13706" refString="Panelius, S. (1965) A revision of the European gall midges of the subfamily Porricondylinae (Diptera: Itonididae). Acta Zoologica Fennica, 113, 1 - 157." type="journal article" year="1965">Panelius (1965</bibRefCitation>
: 85ff.),
<bibRefCitation id="A0784B678A47FFEE278B69C6FADF6FF8" author="Spungis, V." box="[1208,1383,1984,2010]" pageId="5" pageNumber="286" pagination="54 - 74" refId="ref13788" refString="Spungis, V. (1989) A revision of the European gall midge species of the genus Camptomyia Kieffer (Diptera, Cecidomyiidae). Latvijas Entomologs, 32, 54 - 74. [in Russian]" type="journal article" year="1989">Spungis (1989)</bibRefCitation>
, and
<bibRefCitation id="A0784B678A44FFED23A46E9FFE7F6891" author="Jaschhof, M. &amp; Jaschhof, C." box="[151,455,153,179]" pageId="6" pageNumber="287" pagination="1 - 392" refId="ref13243" refString="Jaschhof, M. &amp; Jaschhof, C. (2013) The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement, 20, 1 - 392." type="journal article" year="2013">Jaschhof &amp; Jaschhof (2013</bibRefCitation>
: 328ff.).Also
<bibRefCitation id="A0784B678A44FFED21506E9FFC256891" author="Mamaev, B. M. &amp; Zaitzev, A. I." box="[611,925,153,179]" pageId="6" pageNumber="287" pagination="1 - 12" refId="ref13603" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1998) Ten new species of gall midges of the genus Camptomyia Kieffer (Diptera, Cecidomyiidae) from Far East Palaearctic region. Far Eastern Entomologist, 62, 1 - 12." type="journal article" year="1998">Mamaev &amp; Zaitzev’s (1998)</bibRefCitation>
descriptions of ten new
<taxonomicName id="03E94D158A44FFED279A6E9FFA806891" authorityName="Kieffer" authorityYear="1894" box="[1193,1336,153,179]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED279A6E9FFA806891" box="[1193,1336,153,179]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
from the Far East of
<collectingCountry id="BCFE76068A44FFED222A6EBBFEDC68F5" box="[281,356,189,215]" name="Russia" pageId="6" pageNumber="287">Russia</collectingCountry>
have a bearing on the European fauna, as one of those species was later shown to occur in
<collectingCountry id="BCFE76068A44FFED26506EBBFF0768D9" name="Sweden" pageId="6" pageNumber="287">Sweden</collectingCountry>
and others might be unrecognized synonyms. About 26 different
<taxonomicName id="03E94D158A44FFED20946EE7FB8E68D9" authorityName="Kieffer" authorityYear="1894" box="[935,1078,225,251]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED20946EE7FB8E68D9" box="[935,1078,225,251]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
have previously been shown to occur in Europe, with almost all of these found in the northern parts of the continent (
<bibRefCitation id="A0784B678A44FFED27616F03FADE693D" author="Gagne, R. J. &amp; Jaschhof, M." box="[1106,1382,261,287]" pageId="6" pageNumber="287" refId="ref13081" refString="Gagne, R. J. &amp; Jaschhof, M. (2017) A Catalog of the Cecidomyiidae (Diptera) of the World. Digital. Version 3. 4 th Edition. Available from: http: // www. ars. usda. gov / ARSUserFiles / 80420580 / Gagne _ 2017 _ World _ Cat _ 4 th _ ed. pdf (accessed 15 January 2019)" type="book" year="2017">Gagné &amp; Jaschhof 2017</bibRefCitation>
). Although European
<taxonomicName id="03E94D158A44FFED226E6F2FFE546961" authorityName="Kieffer" authorityYear="1894" box="[349,492,297,323]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED226E6F2FFE546961" box="[349,492,297,323]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
have repeatedly been the focus of taxonomic revisions in the past few decades, there remain a number of unresolved issues with species named in the past. As is known, closely related
<taxonomicName id="03E94D158A44FFED263E6F4BFA246945" authorityName="Kieffer" authorityYear="1894" box="[1293,1436,333,359]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED263E6F4BFA246945" box="[1293,1436,333,359]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
differ mainly in male genitalic characters (the morphology of larvae is generally poorly researched). It is, therefore, crucial that genitalic structures with relevance for species delimation and identification are properly illustrated―an aspiration that is all too often not met in species descriptions published before 2013. In our revision of
<taxonomicName id="03E94D158A44FFED26636FBFFF5B69D5" authorityName="Kieffer" authorityYear="1894" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED26636FBFFF5B69D5" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
in
<collectingCountry id="BCFE76068A44FFED223B6FDBFED969D5" box="[264,353,477,503]" name="Sweden" pageId="6" pageNumber="287">Sweden</collectingCountry>
(
<bibRefCitation id="A0784B678A44FFED225C6FDBFD2569D5" author="Jaschhof, M. &amp; Jaschhof, C." box="[367,669,477,503]" pageId="6" pageNumber="287" pagination="1 - 392" refId="ref13243" refString="Jaschhof, M. &amp; Jaschhof, C. (2013) The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement, 20, 1 - 392." type="journal article" year="2013">Jaschhof &amp; Jaschhof 2013</bibRefCitation>
: 328ff.) we had reason to re-illustrate 14 species named by previous authors, and, regrettably, similar redescriptions are needed for almost all the remaining species in Europe, as well as those described from elsewhere. However, several circumstances impede a comprehensive revision of named
<taxonomicName id="03E94D158A44FFED23A46C4FFE9E6A41" authorityName="Kieffer" authorityYear="1894" box="[151,294,585,611]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED23A46C4FFE9E6A41" box="[151,294,585,611]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
, first and foremost the circumstance that quite a number of species lack properly prepared specimens, a prerequisite for making both informative illustrations and qualified taxonomic decisions. The resources required to collect fresh specimens would be substantial, considering that the species in question are scattered across several continents. (Even greater efforts would have to be made to obtain fresh specimens for integrative taxonomy.) Those resources are not on the horizon, so the only feasible approach is to revise
<taxonomicName id="03E94D158A44FFED20D26CDFFBC86AD1" authorityName="Kieffer" authorityYear="1894" box="[993,1136,729,755]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED20D26CDFFBC86AD1" box="[993,1136,729,755]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
step-by-step as new material accumulates. This is done here: apart from describing four new
<taxonomicName id="03E94D158A44FFED20BE6CFBFBA46B35" authorityName="Kieffer" authorityYear="1894" box="[909,1052,765,791]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED20BE6CFBFBA46B35" box="[909,1052,765,791]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
, we redescribe three of the named species whose identification has previously been proven problematic.
</paragraph>
<paragraph id="C45636968A44FFED23F46D43FE1D6CC9" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="287">
Male genitalic structures are even more diverse in
<taxonomicName id="03E94D158A44FFED21C96D43FC316B7D" authorityName="Kieffer" authorityYear="1894" box="[762,905,837,863]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED21C96D43FC316B7D" box="[762,905,837,863]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
than in
<taxonomicName id="03E94D158A44FFED20D06D43FBF36B7D" authorityName="Loew" authorityYear="1850" box="[995,1099,837,863]" class="Insecta" family="Cecidomyiidae" genus="Asynapta" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED20D06D43FBF36B7D" box="[995,1099,837,863]" italics="true" pageId="6" pageNumber="287">Asynapta</emphasis>
</taxonomicName>
. This is true for all the genitalic components but especially for the parameres. To get an idea of the parameral diversity in European
<taxonomicName id="03E94D158A44FFED26356D6FFA2D6BA1" authorityName="Kieffer" authorityYear="1894" box="[1286,1429,873,899]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED26356D6FFA2D6BA1" box="[1286,1429,873,899]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
, one should quickly look over the 15 species figured in our 2013 book (
<bibRefCitation id="A0784B678A44FFED20F26D8BFB4A6B85" author="Jaschhof, M. &amp; Jaschhof, C." box="[961,1266,909,935]" pageId="6" pageNumber="287" pagination="1 - 392" refId="ref13243" refString="Jaschhof, M. &amp; Jaschhof, C. (2013) The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement, 20, 1 - 392." type="journal article" year="2013">Jaschhof &amp; Jaschhof 2013</bibRefCitation>
: figs 162–176) and be aware that the range of variation indicated there is considerably greater with extra-European species. It is important to note here that the presence of two parameral pairs is obvious, or at least discernible, in most
<taxonomicName id="03E94D158A44FFED263E6DD3FA246BCD" authorityName="Kieffer" authorityYear="1894" box="[1293,1436,981,1007]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED263E6DD3FA246BCD" box="[1293,1436,981,1007]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
(
<figureCitation id="5CD22A138A44FFED23AC6DFFFF506C31" box="[159,232,1017,1043]" captionStart="FIGURES 5–9" captionStartId="8.[151,264,1537,1562]" captionTargetBox="[191,1250,567,1460]" captionTargetId="figure@8.[153,1286,525,1497]" captionTargetPageId="8" captionText="FIGURES 5–9. Camptomyia hedmarki, male, holotype. 5: Genitalia, ventral. 6: Apex of gonostylus, ventral. 7: Zoom of central parts of genitalia, ventral. 8: Fourth flagellomere, lateral. 9: Aedeagus and parameres, ventral. Scales for 5, 0.1 mm, for 6, 0.025 mm, for 7–9, 0.05 mm. Arrows refer to characters described in the diagnosis. Abbreviations: dpm = dorsal parameres, vpm = ventral parameres." httpUri="https://zenodo.org/record/2807076/files/figure.png" pageId="6" pageNumber="287">Figs 9</figureCitation>
,
<figureCitation id="5CD22A138A44FFED23C76DFEFEAA6C31" box="[244,274,1016,1043]" captionStart="FIGURES 10–15" captionStartId="10.[151,264,1677,1702]" captionTargetBox="[189,1250,382,1586]" captionTargetId="figure@10.[153,1286,342,1627]" captionTargetPageId="10" captionText="FIGURES 10–15. Morphology of Camptomyia spp. 10–12: C. oldhammeri, holotype; 13–15: C. rhynchostylata, holo- and paratype. 10: Genitalia, ventral. 11: Ninth tergite, dorsal. 12: Fourth flagellomere, lateral. 13: Fourth flagellomere, lateral, paratype. 14: Genitalia, ventral, holotype. 15: Aedeagus and parameres, ventral, holotype. Scales 0.05 mm. Arrows refer to characters described in the diagnoses." httpUri="https://zenodo.org/record/2807078/files/figure.png" pageId="6" pageNumber="287">15</figureCitation>
,
<figureCitation id="5CD22A138A44FFED222D6DFFFE846C31" box="[286,316,1017,1043]" captionStart="FIGURES 18–24" captionStartId="14.[151,264,1625,1650]" captionTargetBox="[254,1326,349,1563]" captionTargetId="figure@14.[227,1360,310,1595]" captionTargetPageId="14" captionText="FIGURES 18–24. Morphology of Asynaptini spp. 18: Genitalia of Asynapta baltica, ventral, specimen from Sweden. 19: Aedeagus and parameres of A. baltica, ventral, specimen from Latvia. 20: Genitalia of Asynapta inflata, ventral, specimen from Sweden. 21–24: Camptomyia fulva, specimen from Sweden. 21: Fourth flagellomere, lateral. 22: Genitalia, ventral. 23: Ninth tergite, dorsal. 24: Aedeagus and parameres, ventral. Scales for 18, 20–22, 0.05 mm, for 19, 23–24, 0.025 mm. Arrows refer to characters described in the diagnoses." httpUri="https://zenodo.org/record/2807090/files/figure.png" pageId="6" pageNumber="287">24</figureCitation>
,
<figureCitation id="5CD22A138A44FFED227B6DFEFEDF6C31" box="[328,359,1016,1043]" captionStart="FIGURES 25–33" captionStartId="17.[151,264,1540,1565]" captionTargetBox="[262,1331,251,1459]" captionTargetId="figure@17.[227,1360,211,1496]" captionTargetPageId="17" captionText="FIGURES 25–33. Morphologyof Camptomyia spp. 25–27:C. piptopori, specimen from Sweden; 28–33:C. ulmicola, specimens from Sweden. 25: Aedeagus and parameres, ventral. 26: Genitalia, ventral. 27: Ninth tergite, dorsal. 28: Fourth flagellomere, lateral. 29: Genitalia, ventral. 30: Gonostylus, dorsal. 31: Ninth tergite, dorsal. 32: Aedeagus and parameres, ventral. 33: Apex of aedeagus and parameres, ventral, variation. Scales for 25–26, 28–29, 0.05 mm, for 27, 30–32, 0,025 mm. Arrows refer to characters described in the diagnoses." httpUri="https://zenodo.org/record/2807092/files/figure.png" pageId="6" pageNumber="287">25</figureCitation>
); absence of one of the pairs, which is not unusual (
<figureCitation id="5CD22A138A44FFED209F6DFFFBBB6C31" box="[940,1027,1017,1043]" captionStart="FIGURES 10–15" captionStartId="10.[151,264,1677,1702]" captionTargetBox="[189,1250,382,1586]" captionTargetId="figure@10.[153,1286,342,1627]" captionTargetPageId="10" captionText="FIGURES 10–15. Morphology of Camptomyia spp. 10–12: C. oldhammeri, holotype; 13–15: C. rhynchostylata, holo- and paratype. 10: Genitalia, ventral. 11: Ninth tergite, dorsal. 12: Fourth flagellomere, lateral. 13: Fourth flagellomere, lateral, paratype. 14: Genitalia, ventral, holotype. 15: Aedeagus and parameres, ventral, holotype. Scales 0.05 mm. Arrows refer to characters described in the diagnoses." httpUri="https://zenodo.org/record/2807078/files/figure.png" pageId="6" pageNumber="287">Figs 10</figureCitation>
,
<figureCitation id="5CD22A138A44FFED273C6DFFFB966C31" box="[1039,1070,1017,1043]" captionStart="FIGURES 25–33" captionStartId="17.[151,264,1540,1565]" captionTargetBox="[262,1331,251,1459]" captionTargetId="figure@17.[227,1360,211,1496]" captionTargetPageId="17" captionText="FIGURES 25–33. Morphologyof Camptomyia spp. 25–27:C. piptopori, specimen from Sweden; 28–33:C. ulmicola, specimens from Sweden. 25: Aedeagus and parameres, ventral. 26: Genitalia, ventral. 27: Ninth tergite, dorsal. 28: Fourth flagellomere, lateral. 29: Genitalia, ventral. 30: Gonostylus, dorsal. 31: Ninth tergite, dorsal. 32: Aedeagus and parameres, ventral. 33: Apex of aedeagus and parameres, ventral, variation. Scales for 25–26, 28–29, 0.05 mm, for 27, 30–32, 0,025 mm. Arrows refer to characters described in the diagnoses." httpUri="https://zenodo.org/record/2807092/files/figure.png" pageId="6" pageNumber="287">32</figureCitation>
), is explained here by secondary loss. The aedeagus of
<taxonomicName id="03E94D158A44FFED22BA6A1BFDA06C15" authorityName="Kieffer" authorityYear="1894" box="[393,536,1053,1079]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED22BA6A1BFDA06C15" box="[393,536,1053,1079]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
has, despite considerable interspecific variation, a uniform basic structure, which consists of a well-developed, sclerotized rod—the aedeagal apodeme—and the elongate, sclerotized aedeagal apex (
<figureCitation id="5CD22A138A44FFED23936A63FF5D6C5D" box="[160,229,1125,1151]" captionStart="FIGURES 5–9" captionStartId="8.[151,264,1537,1562]" captionTargetBox="[191,1250,567,1460]" captionTargetId="figure@8.[153,1286,525,1497]" captionTargetPageId="8" captionText="FIGURES 5–9. Camptomyia hedmarki, male, holotype. 5: Genitalia, ventral. 6: Apex of gonostylus, ventral. 7: Zoom of central parts of genitalia, ventral. 8: Fourth flagellomere, lateral. 9: Aedeagus and parameres, ventral. Scales for 5, 0.1 mm, for 6, 0.025 mm, for 7–9, 0.05 mm. Arrows refer to characters described in the diagnosis. Abbreviations: dpm = dorsal parameres, vpm = ventral parameres." httpUri="https://zenodo.org/record/2807076/files/figure.png" pageId="6" pageNumber="287">Fig. 9</figureCitation>
). The apodeme is often extended basally to provide space for muscle insertion, with the extension usually resembling a T (
<figureCitation id="5CD22A138A44FFED227F6A8FFE286C81" box="[332,400,1161,1187]" captionStart="FIGURES 5–9" captionStartId="8.[151,264,1537,1562]" captionTargetBox="[191,1250,567,1460]" captionTargetId="figure@8.[153,1286,525,1497]" captionTargetPageId="8" captionText="FIGURES 5–9. Camptomyia hedmarki, male, holotype. 5: Genitalia, ventral. 6: Apex of gonostylus, ventral. 7: Zoom of central parts of genitalia, ventral. 8: Fourth flagellomere, lateral. 9: Aedeagus and parameres, ventral. Scales for 5, 0.1 mm, for 6, 0.025 mm, for 7–9, 0.05 mm. Arrows refer to characters described in the diagnosis. Abbreviations: dpm = dorsal parameres, vpm = ventral parameres." httpUri="https://zenodo.org/record/2807076/files/figure.png" pageId="6" pageNumber="287">Fig. 9</figureCitation>
). The construction of the aedeagal apex is highly variable, both regarding its dimensions and equipment with fine-structures, of which some may lead to remarkably elaborate designs (
<bibRefCitation id="A0784B678A44FFED279E6AABFF6B6CC9" author="Jaschhof, M. &amp; Jaschhof, C." pageId="6" pageNumber="287" pagination="1 - 392" refId="ref13243" refString="Jaschhof, M. &amp; Jaschhof, C. (2013) The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement, 20, 1 - 392." type="journal article" year="2013">Jaschhof &amp; Jaschhof 2013</bibRefCitation>
: figs 162B, 170E).
</paragraph>
<paragraph id="C45636968A44FFED23F46AF3FB8F6EB9" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="287">
The diversity found in male genitalic structures of
<taxonomicName id="03E94D158A44FFED21CC6AF3FC366D2D" authorityName="Kieffer" authorityYear="1894" box="[767,910,1269,1295]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED21CC6AF3FC366D2D" box="[767,910,1269,1295]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
has so far proved to be too intricate as to be of use for building a subgeneric classification that is both comprehensive and in accordance with the natural relationships between species. Even a subdivision into groups of similar-looking species, which would facilitate the task of identification, has not yet been achieved. The difficulty is, in brief, to recognize meaningful patterns in a world of overwhelming diversity. On top of this, it is obvious that only fragments of this diversity have been described to date, so that the time is simply not ripe for clustering species in distinct groups, let alone subgenera. Against this background
<bibRefCitation id="A0784B678A44FFED22106BCBFE516DCA" author="Mamaev, B. M." box="[291,489,1485,1512]" pageId="6" pageNumber="287" pagination="1677 - 1690" refId="ref13562" refString="Mamaev, B. M. (1961) Gall midges of the USSR. New species of the genus Camptomyia Kieffer (Itonididae, Diptera). Zoologicheskii Zhurnal, 40, 1677 - 1690. [in Russian]" type="journal article" year="1961">Mamaev’s (1961)</bibRefCitation>
proposal of dividing
<taxonomicName id="03E94D158A44FFED21E86BCBFCD26DC5" authorityName="Kieffer" authorityYear="1894" box="[731,874,1485,1511]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED21E86BCBFCD26DC5" box="[731,874,1485,1511]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
into five subgenera, including
<emphasis id="F69DEA848A44FFED27FB6BCBFF676E29" italics="true" pageId="6" pageNumber="287">
<taxonomicName id="03E94D158A44FFED27FB6BCBFAEF6DC5" authority="Jaschhof &amp; Jaschhof, 2019" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[1224,1367,1485,1511]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus" sensu="stricto">Camptomyia</taxonomicName>
<taxonomicNameLabel id="EDAE57FF8A44FFED266D6BC8FF676E29" pageId="6" pageNumber="287" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
, must be regarded as premature. Aside from the fact that as many as five subgenera were needed to accommodate only 11 species,
<bibRefCitation id="A0784B678A44FFED22856813FDC76E12" author="Mamaev, B. M." box="[438,639,1557,1584]" pageId="6" pageNumber="287" pagination="1677 - 1690" refId="ref13562" refString="Mamaev, B. M. (1961) Gall midges of the USSR. New species of the genus Camptomyia Kieffer (Itonididae, Diptera). Zoologicheskii Zhurnal, 40, 1677 - 1690. [in Russian]" type="journal article" year="1961">Mamaev’s (1961)</bibRefCitation>
subgeneric definitions are vague, with the exception of
<emphasis id="F69DEA848A44FFED263E6813FE9B6E71" italics="true" pageId="6" pageNumber="287">
<taxonomicName id="03E94D158A44FFED263E6813FA246E0D" authority="Jaschhof &amp; Jaschhof, 2019" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[1293,1436,1557,1583]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus" sensu="stricto">Camptomyia</taxonomicName>
<taxonomicNameLabel id="EDAE57FF8A44FFED23A4683CFE9B6E71" box="[151,291,1594,1619]" pageId="6" pageNumber="287" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
, which is largely identical with the
<taxonomicName id="03E94D158A44FFED2198683FFC8D6E71" baseAuthorityName="Loew" baseAuthorityYear="1851" box="[683,821,1593,1619]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="corticalis">
<emphasis id="F69DEA848A44FFED2198683FFC8D6E71" box="[683,821,1593,1619]" italics="true" pageId="6" pageNumber="287">C. corticalis</emphasis>
</taxonomicName>
group of species by
<bibRefCitation id="A0784B678A44FFED2724683FFAF06E71" author="Jaschhof, M. &amp; Jaschhof, C." box="[1047,1352,1593,1619]" pageId="6" pageNumber="287" pagination="1 - 392" refId="ref13243" refString="Jaschhof, M. &amp; Jaschhof, C. (2013) The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement, 20, 1 - 392." type="journal article" year="2013">Jaschhof &amp; Jaschhof (2013</bibRefCitation>
:
<specimenCount id="D2EFFD1F8A44FFED2661683FFA296E76" box="[1362,1425,1593,1620]" pageId="6" pageNumber="287" type="generic">332f.</specimenCount>
).
<bibRefCitation id="A0784B678A44FFED23A4685BFEFE6E55" author="Spungis, V." box="[151,326,1629,1655]" pageId="6" pageNumber="287" pagination="54 - 74" refId="ref13788" refString="Spungis, V. (1989) A revision of the European gall midge species of the genus Camptomyia Kieffer (Diptera, Cecidomyiidae). Latvijas Entomologs, 32, 54 - 74. [in Russian]" type="journal article" year="1989">Spungis (1989)</bibRefCitation>
, in reference to
<bibRefCitation id="A0784B678A44FFED22CF685BFD7A6E5A" author="Mamaev, B. M." box="[508,706,1629,1656]" pageId="6" pageNumber="287" pagination="1677 - 1690" refId="ref13562" refString="Mamaev, B. M. (1961) Gall midges of the USSR. New species of the genus Camptomyia Kieffer (Itonididae, Diptera). Zoologicheskii Zhurnal, 40, 1677 - 1690. [in Russian]" type="journal article" year="1961">Mamaev’s (1961)</bibRefCitation>
proposal, remarked that it would require another five or six subgenera to accomodate all the 24 different
<taxonomicName id="03E94D158A44FFED215B6887FD4F6EB9" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[616,759,1665,1691]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED215B6887FD4F6EB9" box="[616,759,1665,1691]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
known to him from Europe.
</paragraph>
<paragraph id="C45636968A44FFEC23F468A3FED168F5" blockId="6.[151,1437,153,2015]" lastBlockId="7.[151,1437,152,827]" lastPageId="7" lastPageNumber="288" pageId="6" pageNumber="287">
Similarly unhelpful seems to us the practice to separate single, somehow conspicuous species from
<taxonomicName id="03E94D158A44FFED266368A3FF5B6EC1" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED266368A3FF5B6EC1" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
and classify these in discrete, new genera. Examples of this strategy are
<taxonomicName id="03E94D158A44FFED271268CFFB1F6EC1" authorityName="Fedotova &amp; Sidorenko" authorityYear="2005" box="[1057,1191,1737,1763]" class="Insecta" family="Cecidomyiidae" genus="Eudokimyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED271268CFFB1F6EC1" box="[1057,1191,1737,1763]" italics="true" pageId="6" pageNumber="287">Eudokimyia</emphasis>
</taxonomicName>
and
<taxonomicName id="03E94D158A44FFED27D368CFFADB6EC1" authorityName="Fedotova &amp; Sidorenko" authorityYear="2005" box="[1248,1379,1737,1763]" class="Insecta" family="Cecidomyiidae" genus="Tenepidosis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED27D368CFFADB6EC1" box="[1248,1379,1737,1763]" italics="true" pageId="6" pageNumber="287">Tenepidosis</emphasis>
</taxonomicName>
, two genera introduced by
<bibRefCitation id="A0784B678A44FFED22A268EBFD516F25" author="Fedotova, Z. A. &amp; Sidorenko, V. S." box="[401,745,1772,1799]" pageId="6" pageNumber="287" pagination="89 - 127" refId="ref13033" refString="Fedotova, Z. A. &amp; Sidorenko, V. S. (2005) New species of gall midges of the subfamily Porricondylinae from the Russian Far East (Diptera: Cecidomyiidae). International Journal of Dipterological Research, 16, 89 - 127." type="journal article" year="2005">Fedotova &amp; Sidorenko (2005)</bibRefCitation>
for two
<taxonomicName id="03E94D158A44FFED207C68EBFC666F25" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[847,990,1773,1799]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED207C68EBFC666F25" box="[847,990,1773,1799]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
with needle-like copulatory organ, apparently the parameres intimately merged with the aedeagus. This most simplified
<typeStatus id="1B5288348A44FFED270A6917FBD16F09" box="[1081,1129,1809,1835]" pageId="6" pageNumber="287">type</typeStatus>
of copulatory organ is also found in the Nearctic
<taxonomicName id="03E94D158A44FFED22BE6933FCE36F6D" authority="Hedin &amp; Johnson, 1968" authorityName="Hedin &amp; Johnson" authorityYear="1968" box="[397,859,1845,1871]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="pseudotsugae">
<emphasis id="F69DEA848A44FFED22BE6933FDFE6F6D" box="[397,582,1845,1871]" italics="true" pageId="6" pageNumber="287">C. pseudotsugae</emphasis>
Hedin &amp; Johnson, 1968
</taxonomicName>
, as well as in an undescribed
<taxonomicName id="03E94D158A44FFED279A6933FA806F6D" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[1193,1336,1845,1871]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED279A6933FA806F6D" box="[1193,1336,1845,1871]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
we have seen from
<collectingCountry id="BCFE76068A44FFED2238695FFE786F51" box="[267,448,1881,1907]" name="Czech Republic" pageId="6" pageNumber="287">Czech Republic</collectingCountry>
. There are many other parameral
<typeStatus id="1B5288348A44FFED2002695FFCD46F51" box="[817,876,1881,1907]" pageId="6" pageNumber="287">types</typeStatus>
in
<taxonomicName id="03E94D158A44FFED20BD695FFBA56F51" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[910,1053,1881,1907]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED20BD695FFBA56F51" box="[910,1053,1881,1907]" italics="true" pageId="6" pageNumber="287">Camptomyia</emphasis>
</taxonomicName>
, including several really outstanding, but none correlates with a particular
<typeStatus id="1B5288348A44FFED2151697BFD2A6FB5" box="[610,658,1917,1943]" pageId="6" pageNumber="287">type</typeStatus>
of, for instance, gonostylar or gonocoxal structure (an exception here is, as mentioned above, the
<taxonomicName id="03E94D158A44FFED22E769A7FD866F99" baseAuthorityName="Loew" baseAuthorityYear="1851" box="[468,574,1953,1979]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="corticalis">
<emphasis id="F69DEA848A44FFED22E769A7FD866F99" box="[468,574,1953,1979]" italics="true" pageId="6" pageNumber="287">corticalis</emphasis>
</taxonomicName>
group, whose species are likely to form a monophylum). Randomly chosen combinations of characters, as they underlie the generic concepts of
<taxonomicName id="03E94D158A44FFED20A969C3FB986FFD" authorityName="Fedotova &amp; Sidorenko" authorityYear="2005" box="[922,1056,1989,2015]" class="Insecta" family="Cecidomyiidae" genus="Eudokimyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED20A969C3FB986FFD" box="[922,1056,1989,2015]" italics="true" pageId="6" pageNumber="287">Eudokimyia</emphasis>
</taxonomicName>
and
<taxonomicName id="03E94D158A44FFED276469C3FB626FFD" authorityName="Fedotova &amp; Sidorenko" authorityYear="2005" box="[1111,1242,1989,2015]" class="Insecta" family="Cecidomyiidae" genus="Tenepidosis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A44FFED276469C3FB626FFD" box="[1111,1242,1989,2015]" italics="true" pageId="6" pageNumber="287">Tenepidosis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="A0784B678A44FFEC262469C3FEF56891" author="Fedotova, Z. A. &amp; Sidorenko, V. S." lastPageId="7" lastPageNumber="288" pageId="6" pageNumber="287" pagination="89 - 127" refId="ref13033" refString="Fedotova, Z. A. &amp; Sidorenko, V. S. (2005) New species of gall midges of the subfamily Porricondylinae from the Russian Far East (Diptera: Cecidomyiidae). International Journal of Dipterological Research, 16, 89 - 127." type="journal article" year="2005">Fedotova &amp; Sidorenko 2005</bibRefCitation>
), could be employed to define genera in any quantity desired―an approach rendering the category of genus pointless.
</paragraph>
<paragraph id="C45636968A45FFEC23F46EE7FA886961" blockId="7.[151,1437,152,827]" pageId="7" pageNumber="288">
All the genus-group names introduced by both
<bibRefCitation id="A0784B678A45FFEC21E86EE7FC3768DE" author="Mamaev, B. M." box="[731,911,225,252]" pageId="7" pageNumber="288" pagination="1677 - 1690" refId="ref13562" refString="Mamaev, B. M. (1961) Gall midges of the USSR. New species of the genus Camptomyia Kieffer (Itonididae, Diptera). Zoologicheskii Zhurnal, 40, 1677 - 1690. [in Russian]" type="journal article" year="1961">Mamaev (1961)</bibRefCitation>
and
<bibRefCitation id="A0784B678A45FFEC20F46EE7FAA268D9" author="Fedotova, Z. A. &amp; Sidorenko, V. S." box="[967,1306,224,251]" pageId="7" pageNumber="288" pagination="89 - 127" refId="ref13033" refString="Fedotova, Z. A. &amp; Sidorenko, V. S. (2005) New species of gall midges of the subfamily Porricondylinae from the Russian Far East (Diptera: Cecidomyiidae). International Journal of Dipterological Research, 16, 89 - 127." type="journal article" year="2005">Fedotova &amp; Sidorenko (2005)</bibRefCitation>
are, for the reasons explained above, regarded as synonyms of
<taxonomicName id="03E94D158A45FFEC21EA6F03FCD0693D" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[729,872,261,287]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A45FFEC21EA6F03FCD0693D" box="[729,872,261,287]" italics="true" pageId="7" pageNumber="288">Camptomyia</emphasis>
</taxonomicName>
(see
<bibRefCitation id="A0784B678A45FFEC20976F03FB0F693D" author="Gagne, R. J. &amp; Jaschhof, M." box="[932,1207,261,287]" pageId="7" pageNumber="288" refId="ref13081" refString="Gagne, R. J. &amp; Jaschhof, M. (2017) A Catalog of the Cecidomyiidae (Diptera) of the World. Digital. Version 3. 4 th Edition. Available from: http: // www. ars. usda. gov / ARSUserFiles / 80420580 / Gagne _ 2017 _ World _ Cat _ 4 th _ ed. pdf (accessed 15 January 2019)" type="book" year="2017">Gagné &amp; Jaschhof 2017</bibRefCitation>
), which does not interfere with the fact that they continue being available should their usefulness be shown in future studies.
</paragraph>
<paragraph id="C45636968A45FFEC23F46F4BFDC46B19" blockId="7.[151,1437,152,827]" pageId="7" pageNumber="288">
The latest, in our view superfluous name introduced for
<taxonomicName id="03E94D158A45FFEC207C6F4BFC666945" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[847,990,333,359]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A45FFEC207C6F4BFC666945" box="[847,990,333,359]" italics="true" pageId="7" pageNumber="288">Camptomyia</emphasis>
</taxonomicName>
is
<emphasis id="F69DEA848A45FFEC27306F4BFB376945" box="[1027,1167,333,359]" italics="true" pageId="7" pageNumber="288">Niladmirara</emphasis>
<bibRefCitation id="A0784B678A45FFEC27AB6F4BFAF46945" author="Fedotova, Z. A." box="[1176,1356,333,359]" pageId="7" pageNumber="288" pagination="1 - 12" refId="ref12979" refString="Fedotova, Z. A. (2018) New taxa of the gall midges of the tribe Asynaptini (Diptera: Cecidomyiidae, Porricondylinae) from the Russian Far East. Far Eastern Entomologist, 362, 1 - 12. https: // doi. org / 10.25221 / fee. 362.1" type="journal article" year="2018">Fedotova, 2018</bibRefCitation>
. In her generic definition
<bibRefCitation id="A0784B678A45FFEC22506F77FDAE69A9" author="Fedotova, Z. A." box="[355,534,369,395]" pageId="7" pageNumber="288" pagination="1 - 12" refId="ref12979" refString="Fedotova, Z. A. (2018) New taxa of the gall midges of the tribe Asynaptini (Diptera: Cecidomyiidae, Porricondylinae) from the Russian Far East. Far Eastern Entomologist, 362, 1 - 12. https: // doi. org / 10.25221 / fee. 362.1" type="journal article" year="2018">Fedotova (2018</bibRefCitation>
: 1) referred to eight characters (seven male genitalic) that purportedly differ from the conditions found in
<taxonomicName id="03E94D158A45FFEC22AD6F93FD95698D" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[414,557,405,431]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A45FFEC22AD6F93FD95698D" box="[414,557,405,431]" italics="true" pageId="7" pageNumber="288">Camptomyia</emphasis>
</taxonomicName>
—clearly a statement that is not in accordance with the facts. The truth is rather that each of the putative peculiarities of
<emphasis id="F69DEA848A45FFEC21686FBFFD5F69F1" box="[603,743,441,467]" italics="true" pageId="7" pageNumber="288">Niladmirara</emphasis>
are found here and there in
<taxonomicName id="03E94D158A45FFEC27106FBFFB0A69F1" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[1059,1202,441,467]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A45FFEC27106FBFFB0A69F1" box="[1059,1202,441,467]" italics="true" pageId="7" pageNumber="288">Camptomyia</emphasis>
</taxonomicName>
. The new genus was mainly justified by having two pairs of biramous parameres, which is correctly observed, but as argued here, is a plesiomorphous condition found throughout the asynaptine clade, including
<taxonomicName id="03E94D158A45FFEC20DE6C07FBC46A39" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[1005,1148,513,539]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A45FFEC20DE6C07FBC46A39" box="[1005,1148,513,539]" italics="true" pageId="7" pageNumber="288">Camptomyia</emphasis>
</taxonomicName>
. On these grounds
<emphasis id="F69DEA848A45FFEC26666C07FF5D6A1D" italics="true" pageId="7" pageNumber="288">Niladmirara</emphasis>
is treated here as a new junior synonym of
<taxonomicName id="03E94D158A45FFEC21E06C23FCDA6A1D" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2019" box="[723,866,549,575]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="F69DEA848A45FFEC21E06C23FCDA6A1D" box="[723,866,549,575]" italics="true" pageId="7" pageNumber="288">Camptomyia</emphasis>
</taxonomicName>
. From a technical point of view one has to remark that the
<typeStatus id="1B5288348A45FFEC23C36C4FFE986A41" box="[240,288,585,611]" pageId="7" pageNumber="288">type</typeStatus>
species of
<emphasis id="F69DEA848A45FFEC22A86C4FFD9F6A41" box="[411,551,585,611]" italics="true" pageId="7" pageNumber="288">Niladmirara</emphasis>
,
<emphasis id="F69DEA848A45FFEC21076C4CFD186A41" box="[564,672,585,611]" italics="true" pageId="7" pageNumber="288">N. metula</emphasis>
<bibRefCitation id="A0784B678A45FFEC21946C4FFCEF6A41" author="Fedotova, Z. A." box="[679,855,585,611]" pageId="7" pageNumber="288" pagination="1 - 12" refId="ref12979" refString="Fedotova, Z. A. (2018) New taxa of the gall midges of the tribe Asynaptini (Diptera: Cecidomyiidae, Porricondylinae) from the Russian Far East. Far Eastern Entomologist, 362, 1 - 12. https: // doi. org / 10.25221 / fee. 362.1" type="journal article" year="2018">Fedotova, 2018</bibRefCitation>
, is based on a single male whose genitalic structures are severely deformed, apparently as a result of improper handling during the mounting process. In particular the parameres are disrupted and wrested from their natural connections to other genitalic structures, which is evidenced by
<bibRefCitation id="A0784B678A45FFEC238F6CB3FE326AED" author="Fedotova, Z. A." box="[188,394,693,719]" pageId="7" pageNumber="288" pagination="1 - 12" refId="ref12979" refString="Fedotova, Z. A. (2018) New taxa of the gall midges of the tribe Asynaptini (Diptera: Cecidomyiidae, Porricondylinae) from the Russian Far East. Far Eastern Entomologist, 362, 1 - 12. https: // doi. org / 10.25221 / fee. 362.1" type="journal article" year="2018">Fedotova’s (2018)</bibRefCitation>
line drawing (fig. 1) and micrographs (figs 25–26). Specimens like this are normally disregarded in taxonomic studies to avoid treating artifacts as putative diagnostic characters. Reviewers of
<bibRefCitation id="A0784B678A45FFEC26126CDFFF5A6B35" author="Fedotova, Z. A." pageId="7" pageNumber="288" pagination="1 - 12" refId="ref12979" refString="Fedotova, Z. A. (2018) New taxa of the gall midges of the tribe Asynaptini (Diptera: Cecidomyiidae, Porricondylinae) from the Russian Far East. Far Eastern Entomologist, 362, 1 - 12. https: // doi. org / 10.25221 / fee. 362.1" type="journal article" year="2018">Fedotova’s (2018)</bibRefCitation>
work would likely have passed a pertinent comment, provided that someone had been given the chance to evaluate the manuscript before publication.
</paragraph>
</subSubSection>
</treatment>
</document>
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