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<document id="2146498EFF15CDB7492C0948511B399F" ID-DOI="10.5281/zenodo.158093" ID-GBIF-Dataset="dc3ceda3-b7ae-43ac-b9ae-f5c235acf210" ID-ISSN="1175­5326" ID-Zenodo-Dep="158093" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461026429590" checkinUser="plazi" docAuthor="Pinto-Da-Rocha, Ricardo" docDate="2004" docId="3E4B87CEFFAABF6DDE54FD30243BFE44" docLanguage="en" docName="zt00640.pdf" docOrigin="Zootaxa 640" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Fissiphallius martensi Pinto-Da-Rocha, 2004, new species" docType="treatment" docVersion="9" lastPageNumber="6" masterDocId="C272FFB6FFABBF68DF5CFFCC2676FFBA" masterDocTitle="A new species of Fissiphalliidae from Brazilian Amazon Rain Forest (Arachnida: Opiliones)" masterLastPageNumber="6" masterPageNumber="1" pageNumber="2" updateTime="1698572253321" updateUser="plazi">
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<mods:title id="1EACE85A059FC3C8D0EE941AC479997D">A new species of Fissiphalliidae from Brazilian Amazon Rain Forest (Arachnida: Opiliones)</mods:title>
</mods:titleInfo>
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<mods:namePart id="6E8D2B0B304A7B7B46FBE1362791907E">Pinto-Da-Rocha, Ricardo</mods:namePart>
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<mods:title id="C9252DC912D88F0EAB65F088E275A7D6">Zootaxa</mods:title>
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<mods:part id="154F976DD58C69C33FB12C6F70E27B32">
<mods:date id="EEF3B0AEF6947A67AE9E276399D6D70C">2004</mods:date>
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<mods:identifier id="0C23565C9D9821D64D5A32DBEEFB2D69" type="DOI">10.5281/zenodo.158093</mods:identifier>
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<treatment id="3E4B87CEFFAABF6DDE54FD30243BFE44" ID-DOI="http://doi.org/10.5281/zenodo.6270864" ID-GBIF-Taxon="119606528" ID-Zenodo-Dep="6270864" LSID="urn:lsid:plazi:treatment:3E4B87CEFFAABF6DDE54FD30243BFE44" httpUri="http://treatment.plazi.org/id/3E4B87CEFFAABF6DDE54FD30243BFE44" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2">
<subSubSection id="FEF86553FFAABF69DE54FD3027F2FC83" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="B65D36D8FFAABF69DE54FD3024DBFCAC" blockId="1.[264,685,764,825]" box="[264,685,764,790]" pageId="1" pageNumber="2">
<heading id="ED1581B4FFAABF69DE54FD3024DBFCAC" bold="true" box="[264,685,764,790]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="8496EACAFFAABF69DE54FD3024DBFCAC" bold="true" box="[264,685,764,790]" pageId="1" pageNumber="2">
<taxonomicName id="71E24D5BFFAABF69DE54FD302463FCAC" box="[264,533,764,790]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="8496EACAFFAABF69DE54FD302463FCAC" bold="true" box="[264,533,764,790]" italics="true" pageId="1" pageNumber="2">Fissiphallius martensi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FA557B1FFAABF69DD41FD3024DBFCAC" box="[541,685,764,790]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B65D36D8FFAABF69DE54FCD327F2FC83" blockId="1.[264,685,764,825]" box="[264,388,799,825]" pageId="1" pageNumber="2">(Figs. 1­8)</paragraph>
</subSubSection>
<subSubSection id="FEF86553FFAABF69DE54FCA22722FB4B" pageId="1" pageNumber="2" type="description">
<paragraph id="B65D36D8FFAABF69DE54FCA22722FB4B" blockId="1.[264,1325,878,1905]" pageId="1" pageNumber="2">
<emphasis id="8496EACAFFAABF69DE54FCA227B4FC32" bold="true" box="[264,450,878,904]" pageId="1" pageNumber="2">Type­material:</emphasis>
<typeStatus id="6959887AFFAABF69DE90FCA3244AFC33" box="[460,572,879,905]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
, male (MCN­1147), from Reserva Ducke, Manaus, Amazonas,
<collectingCountry id="CEF57648FFAABF69DE54FC5B2703FC0B" box="[264,373,919,945]" name="Brazil" pageId="1" pageNumber="2">BRAZIL</collectingCountry>
,
<date id="C25C1018FFAABF69DEDEFC5B2425FC0B" box="[386,595,919,945]" pageId="1" pageNumber="2" value="1992-08-06" valueMax="1992-08-09" valueMin="1992-08-06">69 August 1992</date>
, A.D. Brescovit
<emphasis id="8496EACAFFAABF69DC41FC5B253EFC0A" box="[797,840,919,944]" italics="true" pageId="1" pageNumber="2">leg.</emphasis>
.
<typeStatus id="6959887AFFAABF69DC05FC5B25A6FC0B" box="[857,976,919,945]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
: same locality as
<typeStatus id="6959887AFFAABF69DBF9FC5B2379FC0B" box="[1189,1295,919,945]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
,
<specimenCount id="A0E4FD51FFAABF69DA40FC5B272DFC63" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
,
<date id="C25C1018FFAABF69DE37FC732423FC63" box="[363,597,959,985]" pageId="1" pageNumber="2" value="1990-12-11">11 December 1990</date>
, F. Apolinário
<emphasis id="8496EACAFFAABF69DC48FC732530FC62" box="[788,838,959,984]" italics="true" pageId="1" pageNumber="2">leg.,</emphasis>
in termite nest (INPA);
<specimenCount id="A0E4FD51FFAABF69DB26FC732285FC63" box="[1146,1267,959,985]" pageId="1" pageNumber="2" type="female">2 females</specimenCount>
,
<date id="C25C1018FFAABF69DA58FC7327D1FBBB" pageId="1" pageNumber="2" value="1992-08-06" valueMax="1992-08-09" valueMin="1992-08-06">6­9 August 1992</date>
, A.D. Brescovit
<emphasis id="8496EACAFFAABF69DD20FC2B24D1FBBA" box="[636,679,999,1024]" italics="true" pageId="1" pageNumber="2">leg.</emphasis>
(MCN­1147);
<specimenCount id="A0E4FD51FFAABF69DC3FFC2B25CDFBBB" box="[867,955,999,1025]" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
, same data (MZSP­23458);
<specimenCount id="A0E4FD51FFAABF69DA40FC2B2739FB93" pageId="1" pageNumber="2" type="male">3 males</specimenCount>
,
<specimenCount id="A0E4FD51FFAABF69DE3CFBC327A0FB93" box="[352,470,1039,1065]" pageId="1" pageNumber="2" type="female">2 females</specimenCount>
<date id="C25C1018FFAABF69DEBEFBC324A0FB93" box="[482,726,1039,1065]" pageId="1" pageNumber="2" value="1991-08-15" valueMax="1991-08-23" valueMin="1991-08-15">1523 August 1991</date>
, A.D. Brescovit
<emphasis id="8496EACAFFAABF69DCF5FBC325A3FB92" box="[937,981,1039,1064]" italics="true" pageId="1" pageNumber="2">leg.</emphasis>
(MCN­1103); same data,
<specimenCount id="A0E4FD51FFAABF69DA40FBC32732FBEB" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
,
<specimenCount id="A0E4FD51FFAABF69DE08FBFB27C9FBEB" box="[340,447,1079,1105]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
(MZSP­23459);
<specimenCount id="A0E4FD51FFAABF69DDCEFBFB249EFBEB" box="[658,744,1079,1105]" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
,
<specimenCount id="A0E4FD51FFAABF69DDA5FBFB2510FBEB" box="[761,870,1079,1105]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
, A.D. Brescovit
<emphasis id="8496EACAFFAABF69DB6AFBFB2217FBEA" box="[1078,1121,1079,1104]" italics="true" pageId="1" pageNumber="2">leg.</emphasis>
(MCN­1224);
<specimenCount id="A0E4FD51FFAABF69DA40FBFB2732FBC3" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
,
<specimenCount id="A0E4FD51FFAABF69DE0CFB9327CFFBC3" box="[336,441,1119,1145]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
, same data (MZSP­23460);
<specimenCount id="A0E4FD51FFAABF69DC59FB93252EFBC3" box="[773,856,1119,1145]" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
,
<date id="C25C1018FFAABF69DC38FB93224EFBC3" box="[868,1080,1119,1145]" pageId="1" pageNumber="2" value="1992-12-07">7 December 1992</date>
, H. Höfer &amp; T. Gasnier (INPA);
<specimenCount id="A0E4FD51FFAABF69DEF2FB4B2464FB1B" box="[430,530,1159,1185]" pageId="1" pageNumber="2" type="male">2 males</specimenCount>
,
<date id="C25C1018FFAABF69DD78FB4B2483FB1B" box="[548,757,1159,1185]" pageId="1" pageNumber="2" value="1994-01-18">18 January 1994</date>
, A.D. Brescovit
<emphasis id="8496EACAFFAABF69DC91FB4B258EFB1A" box="[973,1016,1159,1184]" italics="true" pageId="1" pageNumber="2">leg.</emphasis>
(MCN­1212);
<specimenCount id="A0E4FD51FFAABF69DBEBFB4B2351FB1B" box="[1207,1319,1159,1185]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
, Reserva Campina, Manaus, Amazonas,
<date id="C25C1018FFAABF69DDB1FB6325BBFB73" box="[749,973,1199,1225]" pageId="1" pageNumber="2" value="1991-09-02">2 September 1991</date>
, A.D. Brescovit
<emphasis id="8496EACAFFAABF69DBC1FB6322BFFB72" box="[1181,1225,1199,1224]" italics="true" pageId="1" pageNumber="2">leg.</emphasis>
(MCN­ 1213).
</paragraph>
</subSubSection>
<subSubSection id="FEF86553FFAABF69DE64FB322727FAFB" pageId="1" pageNumber="2" type="etymology">
<paragraph id="B65D36D8FFAABF69DE64FB322727FAFB" blockId="1.[264,1325,878,1905]" pageId="1" pageNumber="2">
<emphasis id="8496EACAFFAABF69DE64FB3227C9FAA2" bold="true" box="[312,447,1278,1304]" pageId="1" pageNumber="2">Etymology</emphasis>
: Named in honor of Jochen Martens, who discovered the family
<taxonomicName id="71E24D5BFFAABF69DBE4FB33273BFAFB" class="Arachnida" family="Fissiphalliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="family">Fissiphalliidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FEF86553FFAABF69DE64FA8224CBF98B" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="B65D36D8FFAABF69DE64FA8224CBF98B" blockId="1.[264,1325,878,1905]" pageId="1" pageNumber="2">
<emphasis id="8496EACAFFAABF69DE64FA8227CCFAD2" bold="true" box="[312,442,1358,1384]" pageId="1" pageNumber="2">Diagnosis:</emphasis>
<taxonomicName id="71E24D5BFFAABF69DE9DFA832432FAD2" box="[449,580,1359,1384]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="martensi">
<emphasis id="8496EACAFFAABF69DE9DFA832432FAD2" box="[449,580,1359,1384]" italics="true" pageId="1" pageNumber="2">F. martensi</emphasis>
</taxonomicName>
differs from other species of the genus/family (
<emphasis id="8496EACAFFAABF69DB2EFA832731FA2A" italics="true" pageId="1" pageNumber="2">
<taxonomicName id="71E24D5BFFAABF69DB2EFA83237CFAD2" box="[1138,1290,1359,1384]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="spinulatus">F. spinulatus</taxonomicName>
,
<taxonomicName id="71E24D5BFFAABF69DA4AFA832731FA2A" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="stumi">F. stumi</taxonomicName>
</emphasis>
and
<taxonomicName id="71E24D5BFFAABF69DED5FABB244EFA2A" box="[393,568,1399,1424]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="sympatricus">
<emphasis id="8496EACAFFAABF69DED5FABB244EFA2A" box="[393,568,1399,1424]" italics="true" pageId="1" pageNumber="2">F. sympatricus</emphasis>
</taxonomicName>
) by having sulcus II straight (V­shaped in other species), the presence of a long spine on eye mound (small tubercles in others), spines on coxa IV, lateral margin of dorsal scute, tergites, anal operculum and sternites (other species smooth or with short tubercules), genital operculum shorter than estigmatic area (longer in others), and ventral plate of penis sword­like.
</paragraph>
</subSubSection>
<subSubSection id="FEF86553FFAABF6CDE64F9F2279DFA29" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2" type="description">
<paragraph id="B65D36D8FFAABF69DE64F9F22496F9E2" blockId="1.[264,1325,878,1905]" box="[312,736,1598,1624]" pageId="1" pageNumber="2">
<emphasis id="8496EACAFFAABF69DE64F9F22496F9E2" bold="true" box="[312,736,1598,1624]" pageId="1" pageNumber="2">
Description of the male (
<typeStatus id="6959887AFFAABF69DD35F9F224AEF9E2" box="[617,728,1598,1624]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
)
</emphasis>
</paragraph>
<paragraph id="B65D36D8FFAABF69DE64F9AB2408F913" blockId="1.[264,1325,878,1905]" pageId="1" pageNumber="2">Measurements (mm): Total length 2.48; cephalothorax length 0.84; maximal width 2.24; cephalothorax width 1.06.</paragraph>
<paragraph id="B65D36D8FFAABF6ADE64F97B226FFE3C" blockId="1.[264,1325,878,1905]" lastBlockId="2.[264,1323,284,550]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">Dorsum (Fig. 1): Anterior margin with 1­2 tubercles on each side. Eye mound close to anterior margin, with developed eyes, with large and straight anteriorly directed spine. Cephalothorax with 5 small tubercles between eye mound and groove I. Area I with 7 tubercles; II with 14; III with 13; IV with 12; posterior margin with 9 tubercles. Lateral margin with 1 pointed tubercle between sulci IIIII. Free­tergites IIII with 1 enlarged lateral tubercle; I with a larger median tubercle and 78 smaller laterals; II with 12 lateral tubercles; III with 3 long median tubercles. Anal operculum with 3 large tubercles (larger than those on the free tergites), 7 smaller ones on posterior margin.</paragraph>
<paragraph id="B65D36D8FFA9BF6ADE64FE582431FD9C" blockId="2.[264,1323,284,550]" pageId="2" pageNumber="3">Venter: Genital operculum shorter than stigmatic area. Coxa IIV with 6 pointed tubercles; tracheal openings covered by tubercles (Fig. 2); posterior margin and free tergites with 1 row of wide tubercles. Sternite IV with 1 large and curved posteriorly spine fused with anal operculum.</paragraph>
<paragraph id="B65D36D8FFA9BF6ADE54F8C62717F8DD" blockId="2.[264,1322,1802,1895]" pageId="2" pageNumber="3">
<emphasis id="8496EACAFFA9BF6ADE54F8C627CCF898" bold="true" box="[264,442,1802,1826]" pageId="2" pageNumber="3">FIGURES. 14.</emphasis>
Male
<typeStatus id="6959887AFFA9BF6ADEA3F8C7242AF898" box="[511,604,1803,1826]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
of
<taxonomicName id="71E24D5BFFA9BF6ADD23F8C7251AF898" box="[639,876,1803,1826]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="8496EACAFFA9BF6ADD23F8C7251AF898" box="[639,876,1803,1826]" italics="true" pageId="2" pageNumber="3">Fissiphallius martensi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FA557B1FFA9BF6ADC28F8C725B0F898" box="[884,966,1803,1826]" pageId="2" pageNumber="3" rank="species">
sp.
<date id="C25C1018FFA9BF6ADCC4F8C725B0F898" box="[920,966,1803,1826]" pageId="2" pageNumber="3">nov.</date>
</taxonomicNameLabel>
1, dorsal view of habitus; 2, ventral view of coxa IV and sternites; 3, left pedipalp; 4, tibia II. Scale bar: figures 12 =
<quantity id="711A9B3DFFA9BF6ADBFCF8E12297F8FE" box="[1184,1249,1837,1860]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="mm" value="1.0">1 mm</quantity>
; 34 =
<quantity id="711A9B3DFFA9BF6ADE54F89C272BF8DD" box="[264,349,1872,1895]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="3" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
<paragraph id="B65D36D8FFA8BF6BDE54F8FD22DDF8D4" blockId="3.[264,1323,1841,1902]" pageId="3" pageNumber="4">
<emphasis id="8496EACAFFA8BF6BDE54F8FD27CDF8F3" bold="true" box="[264,443,1841,1865]" pageId="3" pageNumber="4">FIGURES. 58.</emphasis>
Penis (
<typeStatus id="6959887AFFA8BF6BDD56F8FE241DF8F3" box="[522,619,1842,1865]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
) of
<taxonomicName id="71E24D5BFFA8BF6BDDCAF8FE25F2F8F3" box="[662,900,1842,1865]" class="Arachnida" family="Zalmoxidae" genus="Fissiphallius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="8496EACAFFA8BF6BDDCAF8FE25F2F8F3" box="[662,900,1842,1865]" italics="true" pageId="3" pageNumber="4">Fissiphallius martensi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FA557B1FFA8BF6BDCD0F8FE25A9F8F3" box="[908,991,1842,1865]" pageId="3" pageNumber="4" rank="species">
sp.
<date id="C25C1018FFA8BF6BDCEDF8FE25A9F8F3" box="[945,991,1842,1865]" pageId="3" pageNumber="4">nov.</date>
</taxonomicNameLabel>
57, scanning of distal part: 5, dorsal; 6, lateral; 7, ventral; 8, lateral view of penis. Scale 57 =
<quantity id="711A9B3DFFA8BF6BDCE7F89B226BF8D4" box="[955,1053,1879,1902]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="0.05">0.05 mm</quantity>
; 8 =
<quantity id="711A9B3DFFA8BF6BDB0EF89B22D0F8D4" box="[1106,1190,1879,1902]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
<paragraph id="B65D36D8FFAFBF6CDE64FED0257AFE14" blockId="4.[264,1325,284,1270]" pageId="4" pageNumber="5">Chelicera: small­tuberculate; 1 tubercle on posterior of bulla; teeth with 4 tubercles. Pedipalp: coxa with 1 ventral tubercle; femur with 5 ventral tubercles (two basal with large setae), 1 subapical on prolateral region; patella with 1 prolateral tubercle; tibia ectal IiIi, mesal IIII; tarsus ectal iIIi/IIi and mesal iIIi.</paragraph>
<paragraph id="B65D36D8FFAFBF6CDE64FE702503FD24" blockId="4.[264,1325,284,1270]" pageId="4" pageNumber="5">Legs: Legs IIV tuberculate. Coxa II with 2 dorsal tubercles; IV visible dorsally to groove IV, with 6 large and thin tubercles and several small. Femur IIV with 1 dorsal row of tubercles; IV with 1 ventral row of 6 large tubercles increasing apicad. Patella IV with 2 large ventrosubapical tubercles. Tibia II swollen ventrally (Fig. 4), others normal; IV with 1 ventral row of large tubercles; 1 retrolateral, 1 prolateral. Metatarsus IV tuberculate, calcaneus swollen. Tarsal segmentation: 4(2), 7(3), 5, 5.</paragraph>
<paragraph id="B65D36D8FFAFBF6CDE64FD60228AFC84" blockId="4.[264,1325,284,1270]" pageId="4" pageNumber="5">Penis (Figs. 58): Truncus without projections. Ventral plate sword­like with 3 pairs of setae (basal pair shorter) on rutrum and 1 pair above pergula, median keel (= pergula) with two pairs of lateral setae. Capsula externa flattened, with 2 apical branches fused (arrowlike on lateral view). Stylus thin and long, hidden by capsula externa and ventral plate.</paragraph>
<paragraph id="B65D36D8FFAFBF6CDE64FC80252BFBBC" blockId="4.[264,1325,284,1270]" pageId="4" pageNumber="5">Colour: Yellowish. Cephalothorax with reticulate brown patches. Areas IIV with dark brown patches laterally. Lateral margin with darker stripe close to posterior border. Free tergites and posterior border with 2 transverse stripes. Femur­Tibia brownish. Tibia II whitish on ventral swollen part. Coxa IV with brownish stripe. Patela IV brownish on distal third. Anal operculum with 2 brownish patches.</paragraph>
<paragraph id="B65D36D8FFAFBF6CDE64FBD82594FB94" blockId="4.[264,1325,284,1270]" box="[312,994,1044,1070]" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDE64FBD8249EFB94" bold="true" box="[312,744,1044,1070]" pageId="4" pageNumber="5">
Description of the female
<typeStatus id="6959887AFFAFBF6CDD24FBD8249EFB94" box="[632,744,1044,1070]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
</emphasis>
(INPA,
<date id="C25C1018FFAFBF6CDC16FBD825ACFB94" box="[842,986,1044,1070]" pageId="4" pageNumber="5" value="1990-12-11">11.XII.1990</date>
)
</paragraph>
<paragraph id="B65D36D8FFAFBF6CDE64FBF022B6FB4C" blockId="4.[264,1325,284,1270]" pageId="4" pageNumber="5">Measurements (mm): Total length 1.9; cephalothorax length 0.74; maximal width 1.7; cephalothorax width 1.02. As male, except for the following: lateral margin with pointed tubercle close to groove II; free­tergite II with 1 median tubercle; anal operculum with 8 tubercles (3 medians larger than others); free sternites with small tubercules; area IV without long tubercle; legs covered by tubercles, smaller in male; tibia II not swollen.</paragraph>
<paragraph id="B65D36D8FFAFBF6CDE54FA9F279DFA29" blockId="4.[264,1323,1363,1427]" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDE54FA9F2709FAD1" bold="true" box="[264,383,1363,1387]" pageId="4" pageNumber="5">TABLE I.</emphasis>
Measurements of appendages of male
<typeStatus id="6959887AFFAFBF6CDC1EFA9825E9FAD1" box="[834,927,1364,1387]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
and female
<typeStatus id="6959887AFFAFBF6CDB69FA9822E7FAD1" box="[1077,1169,1364,1387]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
(parenthized) (INPA,
<date id="C25C1018FFAFBF6CDE06FAB727ABFA28" box="[346,477,1403,1426]" pageId="4" pageNumber="5" value="1990-12-11">11.XII.1990</date>
).
</paragraph>
</subSubSection>
<paragraph id="B65D36D8FFAFBF6CDEF0FA75236AF919" pageId="4" pageNumber="5">
<table id="C4E2C478FFAF4097DE51FA75236AF919" box="[269,1308,1465,1699]" gridcols="8" gridrows="6" pageId="4" pageNumber="5">
<tr id="08D2349AFFAF4097DE51FA75236AFA74" box="[269,1308,1465,1486]" gridrow="0" pageId="4" pageNumber="5" rowspan-0="1">
<th id="4B035DE6FFAF4097DED4FA752784FA74" box="[392,498,1465,1486]" gridcol="1" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDEF0FA7527BBFA74" bold="true" box="[428,461,1465,1486]" pageId="4" pageNumber="5">TR</emphasis>
</th>
<th id="4B035DE6FFAF4097DD53FA75240FFA74" box="[527,633,1465,1486]" gridcol="2" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDD69FA752422FA74" bold="true" box="[565,596,1465,1486]" pageId="4" pageNumber="5">FE</emphasis>
</th>
<th id="4B035DE6FFAF4097DDCAFA752576FA74" box="[662,768,1465,1486]" gridcol="3" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDDE0FA7524ACFA74" bold="true" box="[700,730,1465,1486]" pageId="4" pageNumber="5">PT</emphasis>
</th>
<th id="4B035DE6FFAF4097DC41FA7525F1FA74" box="[797,903,1465,1486]" gridcol="4" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDC19FA752528FA74" bold="true" box="[837,862,1465,1486]" pageId="4" pageNumber="5">TI</emphasis>
</th>
<th id="4B035DE6FFAF4097DCF8FA752278FA74" box="[932,1038,1465,1486]" gridcol="5" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDC9AFA75259BFA74" bold="true" box="[966,1005,1465,1486]" pageId="4" pageNumber="5">MT</emphasis>
</th>
<th id="4B035DE6FFAF4097DB77FA7522E3FA74" box="[1067,1173,1465,1486]" gridcol="6" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDB0CFA752219FA74" bold="true" box="[1104,1135,1465,1486]" pageId="4" pageNumber="5">TA</emphasis>
</th>
<th id="4B035DE6FFAF4097DBEEFA75236AFA74" box="[1202,1308,1465,1486]" gridcol="7" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDB91FA752377FA74" bold="true" box="[1229,1281,1465,1486]" pageId="4" pageNumber="5">Total</emphasis>
</th>
</tr>
<tr id="08D2349AFFAF4097DE51FA2F236AFA43" box="[269,1308,1507,1529]" gridrow="1" pageId="4" pageNumber="5">
<th id="4B035DE6FFAF4097DE51FA2F271EFA43" box="[269,360,1507,1529]" gridcol="0" gridrow="1" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDE51FA2F2760FA42" bold="true" box="[269,278,1507,1528]" pageId="4" pageNumber="5">I</emphasis>
</th>
<td id="4B035DE6FFAF4097DED4FA2F2784FA43" box="[392,498,1507,1529]" gridcol="1" gridrow="1" pageId="4" pageNumber="5">0.44 (0.24)</td>
<td id="4B035DE6FFAF4097DD53FA2F240FFA43" box="[527,633,1507,1529]" gridcol="2" gridrow="1" pageId="4" pageNumber="5">1.14 (0.84)</td>
<td id="4B035DE6FFAF4097DDCAFA2F2576FA43" box="[662,768,1507,1529]" gridcol="3" gridrow="1" pageId="4" pageNumber="5">0.44 (0.4)</td>
<td id="4B035DE6FFAF4097DC41FA2F25F1FA43" box="[797,903,1507,1529]" gridcol="4" gridrow="1" pageId="4" pageNumber="5">0.7 (0.6)</td>
<td id="4B035DE6FFAF4097DCF8FA2F2278FA43" box="[932,1038,1507,1529]" gridcol="5" gridrow="1" pageId="4" pageNumber="5">1.14 (1.02)</td>
<td id="4B035DE6FFAF4097DB77FA2F22E3FA43" box="[1067,1173,1507,1529]" gridcol="6" gridrow="1" pageId="4" pageNumber="5">0.72 (0.64)</td>
<td id="4B035DE6FFAF4097DBEEFA2F236AFA43" box="[1202,1308,1507,1529]" gridcol="7" gridrow="1" pageId="4" pageNumber="5">4.58 (3.74)</td>
</tr>
<tr id="08D2349AFFAF4097DE51F9C2236AF999" box="[269,1308,1550,1571]" gridrow="2" pageId="4" pageNumber="5">
<th id="4B035DE6FFAF4097DE51F9C2271EF999" box="[269,360,1550,1571]" gridcol="0" gridrow="2" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDE51F9C22756F999" bold="true" box="[269,288,1550,1571]" pageId="4" pageNumber="5">II</emphasis>
</th>
<td id="4B035DE6FFAF4097DED4F9C22784F999" box="[392,498,1550,1571]" gridcol="1" gridrow="2" pageId="4" pageNumber="5">0.4 (0.32)</td>
<td id="4B035DE6FFAF4097DD53F9C2240FF999" box="[527,633,1550,1571]" gridcol="2" gridrow="2" pageId="4" pageNumber="5">1.5 (1.2)</td>
<td id="4B035DE6FFAF4097DDCAF9C22576F999" box="[662,768,1550,1571]" gridcol="3" gridrow="2" pageId="4" pageNumber="5">0.58 (0.56)</td>
<td id="4B035DE6FFAF4097DC41F9C225F1F999" box="[797,903,1550,1571]" gridcol="4" gridrow="2" pageId="4" pageNumber="5">1.16 (0.92)</td>
<td id="4B035DE6FFAF4097DCF8F9C22278F999" box="[932,1038,1550,1571]" gridcol="5" gridrow="2" pageId="4" pageNumber="5">1.72 (1.34)</td>
<td id="4B035DE6FFAF4097DB77F9C222E3F999" box="[1067,1173,1550,1571]" gridcol="6" gridrow="2" pageId="4" pageNumber="5">1.62 (1.24)</td>
<td id="4B035DE6FFAF4097DBEEF9C2236AF999" box="[1202,1308,1550,1571]" gridcol="7" gridrow="2" pageId="4" pageNumber="5">6.98 (5.58)</td>
</tr>
<tr id="08D2349AFFAF4097DE51F9F4236AF9F4" box="[269,1308,1592,1614]" gridrow="3" pageId="4" pageNumber="5">
<th id="4B035DE6FFAF4097DE51F9F4271EF9F4" box="[269,360,1592,1614]" gridcol="0" gridrow="3" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDE51F9F4275FF9F7" bold="true" box="[269,297,1592,1613]" pageId="4" pageNumber="5">III</emphasis>
</th>
<td id="4B035DE6FFAF4097DED4F9F42784F9F4" box="[392,498,1592,1614]" gridcol="1" gridrow="3" pageId="4" pageNumber="5">0.28 (0.36)</td>
<td id="4B035DE6FFAF4097DD53F9F4240FF9F4" box="[527,633,1592,1614]" gridcol="2" gridrow="3" pageId="4" pageNumber="5">1.24 (0.96)</td>
<td id="4B035DE6FFAF4097DDCAF9F42576F9F4" box="[662,768,1592,1614]" gridcol="3" gridrow="3" pageId="4" pageNumber="5">0.56 (0.48)</td>
<td id="4B035DE6FFAF4097DC41F9F425F1F9F4" box="[797,903,1592,1614]" gridcol="4" gridrow="3" pageId="4" pageNumber="5">0.94 (0.8)</td>
<td id="4B035DE6FFAF4097DCF8F9F42278F9F4" box="[932,1038,1592,1614]" gridcol="5" gridrow="3" pageId="4" pageNumber="5">1.46 (1.3)</td>
<td id="4B035DE6FFAF4097DB77F9F422E3F9F4" box="[1067,1173,1592,1614]" gridcol="6" gridrow="3" pageId="4" pageNumber="5">0.94 (0.84)</td>
<td id="4B035DE6FFAF4097DBEEF9F4236AF9F4" box="[1202,1308,1592,1614]" gridcol="7" gridrow="3" pageId="4" pageNumber="5">5.42 (4.74)</td>
</tr>
<tr id="08D2349AFFAF4097DE51F9AF236AF9C2" box="[269,1308,1635,1656]" gridrow="4" pageId="4" pageNumber="5">
<th id="4B035DE6FFAF4097DE51F9AF271EF9C2" box="[269,360,1635,1656]" gridcol="0" gridrow="4" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDE51F9AF275EF9C2" bold="true" box="[269,296,1635,1656]" pageId="4" pageNumber="5">IV</emphasis>
</th>
<td id="4B035DE6FFAF4097DED4F9AF2784F9C2" box="[392,498,1635,1656]" gridcol="1" gridrow="4" pageId="4" pageNumber="5">0.48 (0.44)</td>
<td id="4B035DE6FFAF4097DD53F9AF240FF9C2" box="[527,633,1635,1656]" gridcol="2" gridrow="4" pageId="4" pageNumber="5">1.7 (1.32)</td>
<td id="4B035DE6FFAF4097DDCAF9AF2576F9C2" box="[662,768,1635,1656]" gridcol="3" gridrow="4" pageId="4" pageNumber="5">0.76 (0.58)</td>
<td id="4B035DE6FFAF4097DC41F9AF25F1F9C2" box="[797,903,1635,1656]" gridcol="4" gridrow="4" pageId="4" pageNumber="5">1.74 (1.28)</td>
<td id="4B035DE6FFAF4097DCF8F9AF2278F9C2" box="[932,1038,1635,1656]" gridcol="5" gridrow="4" pageId="4" pageNumber="5">1.84 (1.62)</td>
<td id="4B035DE6FFAF4097DB77F9AF22E3F9C2" box="[1067,1173,1635,1656]" gridcol="6" gridrow="4" pageId="4" pageNumber="5">0.94 (0.96)</td>
<td id="4B035DE6FFAF4097DBEEF9AF236AF9C2" box="[1202,1308,1635,1656]" gridcol="7" gridrow="4" pageId="4" pageNumber="5">7.46 (6.2)</td>
</tr>
<tr id="08D2349AFFAF4097DE51F942236AF919" box="[269,1308,1678,1699]" gridrow="5" pageId="4" pageNumber="5">
<th id="4B035DE6FFAF4097DE51F942271EF919" box="[269,360,1678,1699]" gridcol="0" gridrow="5" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDE51F942271EF919" bold="true" box="[269,360,1678,1699]" pageId="4" pageNumber="5">Pedipalp</emphasis>
</th>
<td id="4B035DE6FFAF4097DED4F9422784F919" box="[392,498,1678,1699]" gridcol="1" gridrow="5" pageId="4" pageNumber="5">0.22 (0.24)</td>
<td id="4B035DE6FFAF4097DD53F942240FF919" box="[527,633,1678,1699]" gridcol="2" gridrow="5" pageId="4" pageNumber="5">0.7 (0.66)</td>
<td id="4B035DE6FFAF4097DDCAF9422576F919" box="[662,768,1678,1699]" gridcol="3" gridrow="5" pageId="4" pageNumber="5">0.44 (0.4)</td>
<td id="4B035DE6FFAF4097DC41F94225F1F919" box="[797,903,1678,1699]" gridcol="4" gridrow="5" pageId="4" pageNumber="5">0.46 (0.42)</td>
<td id="4B035DE6FFAF4097DCF8F9422278F919" box="[932,1038,1678,1699]" gridcol="5" gridrow="5" pageId="4" pageNumber="5">­­­­</td>
<td id="4B035DE6FFAF4097DB77F94222E3F919" box="[1067,1173,1678,1699]" gridcol="6" gridrow="5" pageId="4" pageNumber="5">0.5 (0.4)</td>
<td id="4B035DE6FFAF4097DBEEF942236AF919" box="[1202,1308,1678,1699]" gridcol="7" gridrow="5" pageId="4" pageNumber="5">2.32 (2.12)</td>
</tr>
</table>
</paragraph>
<subSubSection id="FEF86553FFAFBF6DDE64F8C92400FE3C" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="distribution">
<paragraph id="B65D36D8FFAFBF6CDE64F8C92403F8F2" blockId="4.[264,1324,1797,1864]" pageId="4" pageNumber="5">
<emphasis id="8496EACAFFAFBF6CDE64F8C927ACF8A5" bold="true" box="[312,474,1797,1823]" pageId="4" pageNumber="5">Distribution:</emphasis>
Recorded only from Manaus, Amazonas,
<collectingCountry id="CEF57648FFAFBF6CDCB8F8CA225AF89A" box="[996,1068,1798,1824]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
in Terra Firme from Central Amazonia Rain Forest.
</paragraph>
<paragraph id="B65D36D8FFAEBF6DDE64FED72400FE3C" blockId="5.[264,1323,283,510]" pageId="5" pageNumber="6">
<emphasis id="8496EACAFFAEBF6DDE64FED7247FFE8F" bold="true" box="[312,521,283,309]" pageId="5" pageNumber="6">Systematic note:</emphasis>
Despite the striking morphological differences present in
<emphasis id="8496EACAFFAEBF6DDB8DFED02749FEE7" italics="true" pageId="5" pageNumber="6">F. m a r ­ tensi</emphasis>
, I preferred to avoid the creation of a new genus in a little known Amazonian family composed of only four species.
</paragraph>
</subSubSection>
<subSubSection id="FEF86553FFAEBF6DDE64FE5F243BFE44" pageId="5" pageNumber="6" type="description">
<paragraph id="B65D36D8FFAEBF6DDE64FE5F243BFE44" blockId="5.[264,1323,283,510]" pageId="5" pageNumber="6">
<emphasis id="8496EACAFFAEBF6DDE64FE5F2785FE17" bold="true" box="[312,499,403,429]" pageId="5" pageNumber="6">Biological note</emphasis>
: Most of specimens were collected from bush and tree branches by beating. One was in an arboreal termite nest. None was obtained from soil samples or surface litter layers (
<quantity id="711A9B3DFFAEBF6DDE89FE282436FE44" box="[469,576,484,510]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="15.0" metricValueMin="0.0" pageId="5" pageNumber="6" unit="cm" value="7.5" valueMax="15.0" valueMin="0.0">015 cm</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>