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<document id="B188655B0AEC34EA90A2A8ED33CC0289" ID-CLB-Dataset="4048" ID-DOI="10.11646/zootaxa.5048.4.2" ID-GBIF-Dataset="ad9f7049-4351-4deb-945a-409f9bbb5076" ID-ISSN="1175-5326" ID-Zenodo-Dep="5556598" ID-ZooBank="6A450390-1C69-42CD-9682-2A766CB31338" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1633693424633" checkinUser="plazi" docAuthor="Monastyrskii, Alexander L. &amp; Lien, Vu Van" docDate="2021" docId="36019464FFFFB600B7F2F9A8FA79FAA2" docLanguage="en" docName="zootaxa.5048.4.2.pdf" docOrigin="Zootaxa 5048 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Talbotia naganum subsp. aurelia Monastyrskii &amp; Vu 2021, subspec. nov." docType="treatment" docVersion="7" lastPageNumber="493" masterDocId="CA38EC1CFFFAB607B765FF8AFFE2FFDB" masterDocTitle="New taxa and new records of butterflies from Vietnam (Lepidoptera, Papilionoidea)" masterLastPageNumber="510" masterPageNumber="486" pageNumber="491" updateTime="1699459186830" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5110CC6C51FE5CA186F0CC3599870200">New taxa and new records of butterflies from Vietnam (Lepidoptera, Papilionoidea)</mods:title>
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<mods:name id="F66B9F638239DC7B2FA3A9DE3C20FE3F" type="personal">
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<mods:namePart id="7CADF688DDF7238625A6724EB4CC9C5D">Monastyrskii, Alexander L.</mods:namePart>
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<mods:affiliation id="A27DC54DF49E5B19CF5EAD57E1BAF0C0">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam https: // orcid. org / 0000 - 0001 - 7645 - 2458 &amp; Fauna &amp; Flora International, Vietnam Programme, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="C88678B025FFAC184D30BFB38FA61687">Lien, Vu Van</mods:namePart>
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<mods:affiliation id="E9A4889BFD4A8BE1B13E32F507EB4374">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam https: // orcid. org / 0000 - 0001 - 7645 - 2458 &amp; vulien @ gmail. com; https: // orcid. org / 0000 - 0003 - 2389 - 9402</mods:affiliation>
<mods:nameIdentifier id="3F87C1A02C5DB2B2AAD74482EF94E523" type="email">vulien@gmail.com</mods:nameIdentifier>
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<mods:date id="041B5113C98F9DBF437B511C04B3ED24">2021</mods:date>
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<treatment id="36019464FFFFB600B7F2F9A8FA79FAA2" ID-DOI="http://doi.org/10.5281/zenodo.5556616" ID-GBIF-Taxon="188639100" ID-Zenodo-Dep="5556616" LSID="urn:lsid:plazi:treatment:36019464FFFFB600B7F2F9A8FA79FAA2" httpUri="http://treatment.plazi.org/id/36019464FFFFB600B7F2F9A8FA79FAA2" lastPageId="7" lastPageNumber="493" pageId="5" pageNumber="491">
<subSubSection id="F6B276F9FFFFB602B7F2F9A8FC61F9E6" box="[151,899,1570,1597]" pageId="5" pageNumber="491" type="nomenclature">
<paragraph id="BE172572FFFFB602B7F2F9A8FC61F9E6" blockId="5.[151,899,1570,1633]" box="[151,899,1570,1597]" pageId="5" pageNumber="491">
<heading id="E55F921EFFFFB602B7F2F9A8FC61F9E6" bold="true" box="[151,899,1570,1597]" fontSize="11" level="1" pageId="5" pageNumber="491" reason="1">
<emphasis id="8CDCF960FFFFB602B7F2F9A8FC61F9E6" bold="true" box="[151,899,1570,1597]" pageId="5" pageNumber="491">
<taxonomicName id="79A85EF1FFFFB602B7F2F9A8FD31F9E6" authority="Monastyrskii &amp; Vu" authorityName="Monastyrskii &amp; Vu" authorityYear="2021" box="[151,723,1570,1597]" class="Insecta" family="Pieridae" genus="Talbotia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="491" phylum="Arthropoda" rank="subSpecies" species="naganum" status="subsp. nov." subSpecies="aurelia">
<emphasis id="8CDCF960FFFFB602B7F2F9A8FE30F9E6" bold="true" box="[151,466,1570,1597]" italics="true" pageId="5" pageNumber="491">Talbotia naganum aurelia</emphasis>
Monastyrskii &amp; Vu
</taxonomicName>
,
<taxonomicNameLabel id="97EF441BFFFFB602B5BAF9A9FC61F9E6" box="[735,899,1571,1597]" pageId="5" pageNumber="491" rank="subSpecies">subspec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFFFB600B7F2F9CDFE39FCE2" lastPageId="7" lastPageNumber="493" pageId="5" pageNumber="491" type="description">
<paragraph id="BE172572FFFFB602B7F2F9CDFF09F9BA" blockId="5.[151,899,1570,1633]" box="[151,235,1607,1633]" pageId="5" pageNumber="491">
(
<figureCitation id="269339F7FFFFB602B7C5F9CDFF01F9BA" box="[160,227,1607,1633]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[157,1438,181,1838]" captionTargetId="figure-17@6.[148,1439,181,1841]" captionTargetPageId="6" captionText="FIGURE 4. Talbotia naganum aurelia Monastyrskii &amp; Vu, subspec. nov. AB—HT♂: central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; EF—PT♀ same location as the male. Talbotia naganum pamsi Vitalis de Salvaza, CD—♂: northern Vietnam, Lang Son province, Mau Son; GH—♀: northern Vietnam, Vinh Phuc province, Tam Dao N.P." figureDoi="http://doi.org/10.5281/zenodo.5556608" httpUri="https://zenodo.org/record/5556608/files/figure.png" pageId="5" pageNumber="491">Fig. 4</figureCitation>
)
</paragraph>
<paragraph id="BE172572FFFFB602B7F2F904FD99F916" blockId="5.[151,1437,1678,2035]" pageId="5" pageNumber="491">
<emphasis id="8CDCF960FFFFB602B7F2F904FE97F972" bold="true" box="[151,373,1678,1705]" pageId="5" pageNumber="491">Description: Male.</emphasis>
Length of forewing:
<specimenCount id="A8AEEEFBFFFFB602B506F905FD94F972" box="[611,630,1679,1705]" pageId="5" pageNumber="491" type="male"></specimenCount>
HT
<quantity id="79508897FFFFB602B5C1F905FD14F972" box="[676,758,1679,1705]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="491" unit="mm" value="30.0">30 mm</quantity>
;
<specimenCount id="A8AEEEFBFFFFB602B464F905FCF1F972" box="[769,787,1679,1705]" pageId="5" pageNumber="491" type="male"></specimenCount>
PTs 30, 27.5, 31, 29, 26, 28, 27, 29, 28.5, 29, 27, 27.5, 30, 27.5, 25, 30, 29.5 mm (n=17; mean=28.32)
</paragraph>
<paragraph id="BE172572FFFFB600B7A2F95CFD7EFE9A" blockId="5.[151,1437,1678,2035]" lastBlockId="7.[151,1437,151,1401]" lastPageId="7" lastPageNumber="493" pageId="5" pageNumber="491">
<emphasis id="8CDCF960FFFFB602B7A2F95CFE5AF92B" bold="true" box="[199,440,1750,1776]" pageId="5" pageNumber="491">Forewing upperside</emphasis>
(
<figureCitation id="269339F7FFFFB602B6AFF95DFDA4F92A" box="[458,582,1751,1777]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[157,1438,181,1838]" captionTargetId="figure-17@6.[148,1439,181,1841]" captionTargetPageId="6" captionText="FIGURE 4. Talbotia naganum aurelia Monastyrskii &amp; Vu, subspec. nov. AB—HT♂: central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; EF—PT♀ same location as the male. Talbotia naganum pamsi Vitalis de Salvaza, CD—♂: northern Vietnam, Lang Son province, Mau Son; GH—♀: northern Vietnam, Vinh Phuc province, Tam Dao N.P." figureDoi="http://doi.org/10.5281/zenodo.5556608" httpUri="https://zenodo.org/record/5556608/files/figure.png" pageId="5" pageNumber="491">Figure 4A</figureCitation>
). Ground colour uniformly white with blackened costal edge at basal and sub-basal areas; black broad submarginal band bordered distal part of costa, apex and half of termen extends from costa to the cell Cu
<subScript id="222C2737FFFFB602B608F8BBFE9FF8E4" attach="left" box="[365,381,1841,1855]" fontSize="6" pageId="5" pageNumber="491">1a</subScript>
with connection to large black submarginal spot in cell M
<subScript id="222C2737FFFFB602B36DF8BBFBF2F8E4" attach="left" box="[1032,1040,1841,1855]" fontSize="6" pageId="5" pageNumber="491">3</subScript>
, though rarely submarginal band is not connected with black spot in cell M
<subScript id="222C2737FFFFB602B53DF8DFFD82F8B8" attach="left" box="[600,608,1877,1891]" fontSize="6" pageId="5" pageNumber="491">3</subScript>
; discocellular vein between M
<subScript id="222C2737FFFFB602B4D8F8DFFC27F8B8" attach="left" box="[957,965,1877,1891]" fontSize="6" pageId="5" pageNumber="491">2</subScript>
and M
<subScript id="222C2737FFFFB602B373F8DFFBFCF8B8" attach="left" box="[1046,1054,1877,1891]" fontSize="6" pageId="5" pageNumber="491">3</subScript>
is ornamented by one black spot; submarginal area in cell Cu
<subScript id="222C2737FFFFB602B6ADF8F3FE3AF85C" attach="left" box="[456,472,1913,1927]" fontSize="6" pageId="5" pageNumber="491">1b</subScript>
sometimes bears blackish round spot varied in size.
<emphasis id="8CDCF960FFFFB602B37AF8ECFAEEF85B" bold="true" box="[1055,1292,1894,1920]" pageId="5" pageNumber="491">Forewing underside</emphasis>
(
<figureCitation id="269339F7FFFFB602B27FF8EDFA6DF85A" box="[1306,1423,1895,1921]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[157,1438,181,1838]" captionTargetId="figure-17@6.[148,1439,181,1841]" captionTargetPageId="6" captionText="FIGURE 4. Talbotia naganum aurelia Monastyrskii &amp; Vu, subspec. nov. AB—HT♂: central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; EF—PT♀ same location as the male. Talbotia naganum pamsi Vitalis de Salvaza, CD—♂: northern Vietnam, Lang Son province, Mau Son; GH—♀: northern Vietnam, Vinh Phuc province, Tam Dao N.P." figureDoi="http://doi.org/10.5281/zenodo.5556608" httpUri="https://zenodo.org/record/5556608/files/figure.png" pageId="5" pageNumber="491">Figure 4B</figureCitation>
). Ground colour uniformly white with yellowish ochreous margin from mid costa to mid termen which broadens at the apical area; large, usually rectangular black spot presented within centre of cell M
<subScript id="222C2737FFFFB602B322F84BFBADF814" attach="left" box="[1095,1103,1985,1999]" fontSize="6" pageId="5" pageNumber="491">3</subScript>
; sometimes, small, blackened spots presented in cells Cu
<subScript id="222C2737FFFFB602B6AEF86FFE39F828" attach="left" box="[459,475,2021,2035]" fontSize="6" pageId="5" pageNumber="491">1b</subScript>
and M
<subScript id="222C2737FFFFB602B54AF86FFDD5F828" attach="left" box="[559,567,2021,2035]" fontSize="6" pageId="5" pageNumber="491">1</subScript>
. Basal and sub-basal part of discal cell densely covered by blackish scales; such dusty black area prominent at lower part of discal cell along cubitus; distinct black spot on the discocellular vein between M
<subScript id="222C2737FFFDB600B634FF47FEBBFF00" attach="left" box="[337,345,205,219]" fontSize="6" pageId="7" pageNumber="493">2</subScript>
and M
<subScript id="222C2737FFFDB600B6C8FF47FE57FF00" attach="left" box="[429,437,205,219]" fontSize="6" pageId="7" pageNumber="493">3</subScript>
diffuses along cubitus.
<emphasis id="8CDCF960FFFDB600B5AFFF30FC22FF0F" bold="true" box="[714,960,186,212]" pageId="7" pageNumber="493">Hindwing upperside</emphasis>
(
<figureCitation id="269339F7FFFDB600B4B4FF31FBAEFF0E" box="[977,1100,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[157,1438,181,1838]" captionTargetId="figure-17@6.[148,1439,181,1841]" captionTargetPageId="6" captionText="FIGURE 4. Talbotia naganum aurelia Monastyrskii &amp; Vu, subspec. nov. AB—HT♂: central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; EF—PT♀ same location as the male. Talbotia naganum pamsi Vitalis de Salvaza, CD—♂: northern Vietnam, Lang Son province, Mau Son; GH—♀: northern Vietnam, Vinh Phuc province, Tam Dao N.P." figureDoi="http://doi.org/10.5281/zenodo.5556608" httpUri="https://zenodo.org/record/5556608/files/figure.png" pageId="7" pageNumber="493">Figure 4A</figureCitation>
). Ground colour of hindwing uniformly whitish with very pale creamy tinge which is the result of intensive yellow ground colour of the hindwing under surface.
<emphasis id="8CDCF960FFFDB600B65BFE88FDD0FEC7" bold="true" box="[318,562,258,284]" pageId="7" pageNumber="493">Hindwing underside</emphasis>
(
<figureCitation id="269339F7FFFDB600B524FE89FD5AFEC6" box="[577,696,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[157,1438,181,1838]" captionTargetId="figure-17@6.[148,1439,181,1841]" captionTargetPageId="6" captionText="FIGURE 4. Talbotia naganum aurelia Monastyrskii &amp; Vu, subspec. nov. AB—HT♂: central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; EF—PT♀ same location as the male. Talbotia naganum pamsi Vitalis de Salvaza, CD—♂: northern Vietnam, Lang Son province, Mau Son; GH—♀: northern Vietnam, Vinh Phuc province, Tam Dao N.P." figureDoi="http://doi.org/10.5281/zenodo.5556608" httpUri="https://zenodo.org/record/5556608/files/figure.png" pageId="7" pageNumber="493">Figure 4B</figureCitation>
). Ground colour of the hindwing uniformly intensive yellow with light golden tinge and whitish scales on veins.
</paragraph>
<caption id="EAD775FAFFFCB601B7F2F8C3FBBFF872" ID-DOI="http://doi.org/10.5281/zenodo.5556608" ID-Zenodo-Dep="5556608" httpUri="https://zenodo.org/record/5556608/files/figure.png" pageId="6" pageNumber="492" startId="6.[151,250,1865,1889]" targetBox="[157,1438,181,1838]" targetPageId="6">
<paragraph id="BE172572FFFCB601B7F2F8C3FBBFF872" blockId="6.[151,1437,1865,1962]" pageId="6" pageNumber="492">
<emphasis id="8CDCF960FFFCB601B7F2F8C3FEF0F8BA" bold="true" box="[151,274,1865,1889]" pageId="6" pageNumber="492">FIGURE 4.</emphasis>
<taxonomicName id="79A85EF1FFFCB601B67DF8C3FD0AF8BA" authority="Monastyrskii &amp; Vu" authorityName="Monastyrskii &amp; Vu" authorityYear="2021" box="[280,744,1865,1890]" class="Insecta" family="Pieridae" genus="Talbotia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="492" phylum="Arthropoda" rank="subSpecies" species="naganum" status="subsp. nov." subSpecies="aurelia">
<emphasis id="8CDCF960FFFCB601B67DF8C3FDFFF8BA" box="[280,541,1865,1889]" italics="true" pageId="6" pageNumber="492">Talbotia naganum aurelia</emphasis>
Monastyrskii &amp; Vu
</taxonomicName>
,
<emphasis id="8CDCF960FFFCB601B594F8C0FC2FF8BA" bold="true" box="[753,973,1865,1890]" pageId="6" pageNumber="492">
<taxonomicNameLabel id="97EF441BFFFCB601B594F8C0FC9FF8B9" box="[753,893,1866,1890]" pageId="6" pageNumber="492" rank="subSpecies">subspec. nov.</taxonomicNameLabel>
AB—
</emphasis>
HT♂: central Vietnam, Gia Lai province, Kon Ka Kinh N.P.;
<emphasis id="8CDCF960FFFCB601B64EF8E4FEBAF85D" bold="true" box="[299,344,1902,1926]" pageId="6" pageNumber="492">EF</emphasis>
—PT♀ same location as the male.
<taxonomicName id="79A85EF1FFFCB601B5DDF8E7FB69F85E" authority="Vitalis de Salvaza, C" authorityName="Vitalis de Salvaza, C" box="[696,1163,1901,1925]" class="Insecta" family="Pieridae" genus="Talbotia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="492" phylum="Arthropoda" rank="subSpecies" species="naganum" subSpecies="pamsi">
<emphasis id="8CDCF960FFFCB601B5DDF8E7FC51F85E" box="[696,947,1901,1925]" italics="true" pageId="6" pageNumber="492">Talbotia naganum pamsi</emphasis>
Vitalis de Salvaza,
<emphasis id="8CDCF960FFFCB601B31FF8E7FB69F85E" bold="true" box="[1146,1163,1901,1925]" pageId="6" pageNumber="492">C</emphasis>
</taxonomicName>
<emphasis id="8CDCF960FFFCB601B3EEF8E7FB49F85E" bold="true" box="[1163,1195,1901,1925]" pageId="6" pageNumber="492">D</emphasis>
—♂: northern Vietnam, Lang Son province, Mau Son;
<emphasis id="8CDCF960FFFCB601B6ABF81BFDE1F872" bold="true" box="[462,515,1937,1961]" pageId="6" pageNumber="492">GH</emphasis>
—♀: northern Vietnam, Vinh Phuc province, Tam Dao N.P.
</paragraph>
</caption>
<paragraph id="BE172572FFFDB600B7A2FEC1FBF7FEBD" blockId="7.[151,1437,151,1401]" box="[199,1045,330,358]" pageId="7" pageNumber="493">
<emphasis id="8CDCF960FFFDB600B7A2FEC1FEFCFEBE" bold="true" box="[199,286,331,357]" pageId="7" pageNumber="493">Female</emphasis>
. Length of forewing: PTs 28, 27, 26, 27,
<quantity id="79508897FFFDB600B588FEC1FCDEFEBE" box="[749,828,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="493" unit="mm" value="23.0">23 mm</quantity>
(n=5; mean= 26,2)
</paragraph>
<paragraph id="BE172572FFFDB600B7A2FEE4FE39FCE2" blockId="7.[151,1437,151,1401]" pageId="7" pageNumber="493">
<emphasis id="8CDCF960FFFDB600B7A2FEE4FE56FE53" bold="true" box="[199,436,366,392]" pageId="7" pageNumber="493">Forewing upperside</emphasis>
(
<figureCitation id="269339F7FFFDB600B6A7FEE5FDD1FE52" box="[450,563,367,393]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[157,1438,181,1838]" captionTargetId="figure-17@6.[148,1439,181,1841]" captionTargetPageId="6" captionText="FIGURE 4. Talbotia naganum aurelia Monastyrskii &amp; Vu, subspec. nov. AB—HT♂: central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; EF—PT♀ same location as the male. Talbotia naganum pamsi Vitalis de Salvaza, CD—♂: northern Vietnam, Lang Son province, Mau Son; GH—♀: northern Vietnam, Vinh Phuc province, Tam Dao N.P." figureDoi="http://doi.org/10.5281/zenodo.5556608" httpUri="https://zenodo.org/record/5556608/files/figure.png" pageId="7" pageNumber="493">Figure 4E</figureCitation>
) Ground colour of the forewing varies from uniformly white to uniformly very pale yellowish unequally dusted with black scales; pattern includes a thick black border around the wing perimeter with small gap in cell Cu
<subScript id="222C2737FFFDB600B6DCFE43FE2BFE0C" attach="left" box="[441,457,457,471]" fontSize="6" pageId="7" pageNumber="493">1b</subScript>
, which includes a black oval spot; discal cell densely covered by black scales or it is fully black; discocellular veins covered by distinct black spot and thick black band connects discal cell with black border in cell M
<subScript id="222C2737FFFDB600B62FFD9BFEB0FDC4" attach="left" box="[330,338,529,543]" fontSize="6" pageId="7" pageNumber="493">3</subScript>
.
<emphasis id="8CDCF960FFFDB600B63AFE74FDAEFDC3" bold="true" box="[351,588,510,536]" pageId="7" pageNumber="493">Forewing underside</emphasis>
(
<figureCitation id="269339F7FFFDB600B53FFE75FD29FDC2" box="[602,715,511,537]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[157,1438,181,1838]" captionTargetId="figure-17@6.[148,1439,181,1841]" captionTargetPageId="6" captionText="FIGURE 4. Talbotia naganum aurelia Monastyrskii &amp; Vu, subspec. nov. AB—HT♂: central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; EF—PT♀ same location as the male. Talbotia naganum pamsi Vitalis de Salvaza, CD—♂: northern Vietnam, Lang Son province, Mau Son; GH—♀: northern Vietnam, Vinh Phuc province, Tam Dao N.P." figureDoi="http://doi.org/10.5281/zenodo.5556608" httpUri="https://zenodo.org/record/5556608/files/figure.png" pageId="7" pageNumber="493">Figure 4F</figureCitation>
) Ground colour of the white, densely dusted with blackish scales especially within discal cell; yellowish ochreous border spread from mid costa to mid termen becoming broader at the apical area; black band within discal cell curve behind discocellular vein and broadened at termen within cell M
<subScript id="222C2737FFFDB600B7D5FDF7FF5AFD50" attach="left" box="[176,184,637,651]" fontSize="6" pageId="7" pageNumber="493">3</subScript>
; another black band extends within cells 2A and Cu
<subScript id="222C2737FFFDB600B467FDF7FCF0FD50" attach="left" box="[770,786,637,651]" fontSize="6" pageId="7" pageNumber="493">1b</subScript>
from the basal area to the termen where it terminates as a large oval spot in cell Cu
<subScript id="222C2737FFFDB600B6CBFD2BFE5CFD74" attach="left" box="[430,446,673,687]" fontSize="6" pageId="7" pageNumber="493">1b</subScript>
.
<emphasis id="8CDCF960FFFDB600B6A9FD04FD5DFD73" bold="true" box="[460,703,654,680]" pageId="7" pageNumber="493">Hindwing upperside</emphasis>
(
<figureCitation id="269339F7FFFDB600B5A8FD05FCA0FD72" box="[717,834,655,681]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[157,1438,181,1838]" captionTargetId="figure-17@6.[148,1439,181,1841]" captionTargetPageId="6" captionText="FIGURE 4. Talbotia naganum aurelia Monastyrskii &amp; Vu, subspec. nov. AB—HT♂: central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; EF—PT♀ same location as the male. Talbotia naganum pamsi Vitalis de Salvaza, CD—♂: northern Vietnam, Lang Son province, Mau Son; GH—♀: northern Vietnam, Vinh Phuc province, Tam Dao N.P." figureDoi="http://doi.org/10.5281/zenodo.5556608" httpUri="https://zenodo.org/record/5556608/files/figure.png" pageId="7" pageNumber="493">Figure 4E</figureCitation>
). Ground colour of hindwing varies from pale creamy (specimens of dsf) to yellowish; series of marginal blackish triangle spots from vein Cu
<subScript id="222C2737FFFDB600B30AFD4FFB9DFD08" attach="left" box="[1135,1151,709,723]" fontSize="6" pageId="7" pageNumber="493">1b</subScript>
to M
<subScript id="222C2737FFFDB600B3D8FD4FFB27FD08" attach="left" box="[1213,1221,709,723]" fontSize="6" pageId="7" pageNumber="493">2</subScript>
and spot in cell Rs usually weakly developed; in some specimens of dsf these spots fully disappeared; basal area dusted with black scales.
<emphasis id="8CDCF960FFFDB600B78EFD70FE03FCCF" bold="true" box="[235,481,762,788]" pageId="7" pageNumber="493">Hindwing underside</emphasis>
(
<figureCitation id="269339F7FFFDB600B697FD71FD8AFCCE" box="[498,616,763,789]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[157,1438,181,1838]" captionTargetId="figure-17@6.[148,1439,181,1841]" captionTargetPageId="6" captionText="FIGURE 4. Talbotia naganum aurelia Monastyrskii &amp; Vu, subspec. nov. AB—HT♂: central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; EF—PT♀ same location as the male. Talbotia naganum pamsi Vitalis de Salvaza, CD—♂: northern Vietnam, Lang Son province, Mau Son; GH—♀: northern Vietnam, Vinh Phuc province, Tam Dao N.P." figureDoi="http://doi.org/10.5281/zenodo.5556608" httpUri="https://zenodo.org/record/5556608/files/figure.png" pageId="7" pageNumber="493">Figure 4F</figureCitation>
). Ground colour of the hindwing uniformly intensive yellow with light golden tinge; fringe is white.
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFFDB600B7A2FCC8FA6DFBA6" pageId="7" pageNumber="493" type="diagnosis">
<paragraph id="BE172572FFFDB600B7A2FCC8FA6DFBA6" blockId="7.[151,1437,151,1401]" pageId="7" pageNumber="493">
<emphasis id="8CDCF960FFFDB600B7A2FCC8FE14FC87" bold="true" box="[199,502,834,861]" pageId="7" pageNumber="493">Diagnosis and discussion:</emphasis>
The bright golden yellow colouration and unique pattern of the hindwing undersides are the main distinctive characteristics in both sexes of the new taxon. The subspecies
<emphasis id="8CDCF960FFFDB600B320FCEDFB6AFC5B" box="[1093,1160,871,896]" italics="true" pageId="7" pageNumber="493">pamsi</emphasis>
is distributed in northern
<collectingCountry id="C6BF65E2FFFDB600B7F2FC01FF14FC7E" box="[151,246,907,933]" name="Vietnam" pageId="7" pageNumber="493">Vietnam</collectingCountry>
and the northern part of central
<collectingCountry id="C6BF65E2FFFDB600B532FC01FD5BFC7E" box="[599,697,907,933]" name="Vietnam" pageId="7" pageNumber="493">Vietnam</collectingCountry>
, and the intensity of the yellow colouration increases from north to south. Other subspecies found in
<collectingCountry id="C6BF65E2FFFDB600B56DFC25FDA9FC12" box="[520,587,943,969]" name="China" pageId="7" pageNumber="493">China</collectingCountry>
(
<taxonomicName id="79A85EF1FFFDB600B53CFC25FCEBFC13" box="[601,777,943,969]" pageId="7" pageNumber="493" rank="subSpecies" subSpecies="cisseis">
subspec.
<emphasis id="8CDCF960FFFDB600B5A5FC25FCEBFC13" box="[704,777,943,968]" italics="true" pageId="7" pageNumber="493">cisseis</emphasis>
</taxonomicName>
) and
<collectingCountry id="C6BF65E2FFFDB600B422FC25FC7BFC12" box="[839,921,943,969]" name="Taiwan" pageId="7" pageNumber="493">Taiwan</collectingCountry>
(
<taxonomicName id="79A85EF1FFFDB600B4C2FC25FB87FC12" box="[935,1125,943,969]" pageId="7" pageNumber="493" rank="subSpecies" subSpecies="karumii">
subspec.
<emphasis id="8CDCF960FFFDB600B36BFC25FB87FC12" box="[1038,1125,943,969]" italics="true" pageId="7" pageNumber="493">karumii</emphasis>
</taxonomicName>
) have much paler hindwing undersides. The females of
<emphasis id="8CDCF960FFFDB600B6B5FC59FDAFFC37" box="[464,589,979,1004]" italics="true" pageId="7" pageNumber="493">T. n. pamsi</emphasis>
and the other subspecies mentioned above have prominent marginal black spots on the upperside of the hindwing while these spots are usually weakly developed or absent in the
<emphasis id="8CDCF960FFFDB600B20DFC7DFF05FBEE" italics="true" pageId="7" pageNumber="493">T. n. aurelia</emphasis>
. In addition, all black markings on the underside of the forewing are larger and darker. Gia Lai province in the Vietnamese Central Highlands is the southern-most border of the subspecies distribution. Comparison of populations demonstrates intensification of yellow coloration on the underside of both wings from north to south.
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFFDB600B7A2FB0DFF23FAEA" pageId="7" pageNumber="493" type="materials_examined">
<paragraph id="BE172572FFFDB600B7A2FB0DFF23FAEA" blockId="7.[151,1437,151,1401]" pageId="7" pageNumber="493">
<emphasis id="8CDCF960FFFDB600B7A2FB0DFE94FB7B" bold="true" box="[199,374,1158,1185]" pageId="7" pageNumber="493">Type material:</emphasis>
<materialsCitation id="0EC02F2FFFFDB600B618FB0DFD48FB1E" ID-GBIF-Occurrence="3383539309" collectingDate="2018-05-19" collectionCode="ALM" collectorName="Kon Ka Kinh N. P." country="Vietnam" elevation="800" location="Man Giang district" pageId="7" pageNumber="493" specimenCount="1" specimenCount-male="1" stateProvince="Gia Lai" typeStatus="holotype">
<typeStatus id="61139BD0FFFDB600B618FB0DFE08FB7A" box="[381,490,1159,1185]" pageId="7" pageNumber="493" type="holotype">Holotype</typeStatus>
: CENTRAL
<collectingCountry id="C6BF65E2FFFDB600B51DFB0DFD1CFB7A" box="[632,766,1159,1185]" name="Vietnam" pageId="7" pageNumber="493">VIETNAM</collectingCountry>
:
<specimenCount id="A8AEEEFBFFFDB600B46DFB0DFCFFFB7A" box="[776,797,1159,1185]" pageId="7" pageNumber="493" type="male"></specimenCount>
<collectingRegion id="7C6CEB90FFFDB600B446FB0DFC00FB7A" box="[803,994,1159,1185]" country="Vietnam" name="Gia Lai" pageId="7" pageNumber="493">Gia Lai province</collectingRegion>
,
<location id="BB7773A9FFFDB600B489FB0DFB23FB7A" LSID="urn:lsid:plazi:treatment:36019464FFFFB600B7F2F9A8FA79FAA2:BB7773A9FFFDB600B489FB0DFB23FB7A" box="[1004,1217,1159,1185]" country="Vietnam" name="Man Giang district" pageId="7" pageNumber="493" stateProvince="Gia Lai">Man Giang district</location>
,
<collectorName id="135D40A4FFFDB600B3AEFB0DFA7AFB7A" box="[1227,1432,1159,1185]" pageId="7" pageNumber="493">Kon Ka Kinh N.P.</collectorName>
,
<date id="CA1603B2FFFDB600B7F2FB21FEEEFB1E" box="[151,268,1195,1221]" pageId="7" pageNumber="493" value="2018-05-19">
<collectingDate id="DA52FA5AFFFDB600B7F2FB21FEEEFB1E" box="[151,268,1195,1221]" pageId="7" pageNumber="493" value="2018-05-19">19.V.2018</collectingDate>
</date>
, forest stream,
<elevation id="3585C241FFFDB600B6D8FB21FDD3FB1E" box="[445,561,1195,1221]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="493" unit="m" value="800.0">
<quantity id="79508897FFFDB600B6D8FB21FDE4FB1E" box="[445,518,1195,1221]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="493" unit="m" value="800.0">800 m</quantity>
asl
</elevation>
,
<collectionCode id="D8B9BDB7FFFDB600B558FB21FD99FB1E" box="[573,635,1195,1221]" country="0" httpUri="http://grbio.org/cool/icw9-2ehs" name="Museum National Historie Naturelle" pageId="7" pageNumber="493">ALM</collectionCode>
leg
</materialsCitation>
.
<materialsCitation id="0EC02F2FFFFDB600B5D3FB21FAA0FB1E" ID-GBIF-Occurrence="3383539327" box="[694,1346,1194,1221]" collectingDate="2018-05-19" collectionCode="ALM" collectorName="Kon Ka Kinh N. P." country="Vietnam" elevation="800" location="Man Giang district" pageId="7" pageNumber="493" specimenCount="16" specimenCount-female="5" specimenCount-male="11" stateProvince="Gia Lai" typeStatus="paratype">
<typeStatus id="61139BD0FFFDB600B5D3FB21FCC1FB1E" box="[694,803,1195,1221]" pageId="7" pageNumber="493" type="paratype">Paratypes</typeStatus>
<specimenCount id="A8AEEEFBFFFDB600B449FB21FCBCFB1E" box="[812,862,1195,1221]" pageId="7" pageNumber="493" type="male">11♂</specimenCount>
,
<specimenCount id="A8AEEEFBFFFDB600B409FB20FC6DFB1E" box="[876,911,1194,1221]" pageId="7" pageNumber="493" type="female">5♀</specimenCount>
from the same locality as HT;
<date id="CA1603B2FFFDB600B39DFB21FAA0FB1E" box="[1272,1346,1195,1221]" pageId="7" pageNumber="493" value="2018-05-18" valueMax="2018-05-19" valueMin="2018-05-18">1819.</date>
</materialsCitation>
<materialsCitation id="0EC02F2FFFFDB600B224FB21FE7DFB32" ID-GBIF-Occurrence="3383539332" collectingDate="1999-04-11" collectingDateMax="2019-04-13" collectingDateMin="1999-04-11" collectionCode="V" country="Vietnam" pageId="7" pageNumber="493" specimenCount="1" stateProvince="Gia Lai" typeStatus="paratype">
<collectionCode id="D8B9BDB7FFFDB600B224FB21FAB5FB1E" box="[1345,1367,1195,1221]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="493" type="Museum">V</collectionCode>
.2018;
<date id="CA1603B2FFFDB600B7F2FB45FEF5FB32" box="[151,279,1231,1257]" pageId="7" pageNumber="493" value="2019-04-13">
<collectingDate id="DA52FA5AFFFDB600B7F2FB45FEF5FB32" box="[151,279,1231,1257]" pageId="7" pageNumber="493" value="2019-04-13">13.IV.2019</collectingDate>
</date>
;
<date id="CA1603B2FFFDB600B647FB45FE7DFB32" box="[290,415,1231,1257]" pageId="7" pageNumber="493" value="1999-04-11">
<collectingDate id="DA52FA5AFFFDB600B647FB45FE7DFB32" box="[290,415,1231,1257]" pageId="7" pageNumber="493" value="1999-04-11">11.IV.1999</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0EC02F2FFFFDB600B6CEFB45FB30FB32" ID-GBIF-Occurrence="3383539335" box="[427,1234,1231,1257]" collectionCode="R" collectorName="Son Thanh N." country="Vietnam" location="Quang Nam province" pageId="7" pageNumber="493" specimenCount="6" specimenCount-male="6" stateProvince="Quang Nam" typeStatus="paratype">
<specimenCount id="A8AEEEFBFFFDB600B6CEFB45FE32FB32" box="[427,464,1231,1257]" pageId="7" pageNumber="493" type="male">6♂</specimenCount>
, CENTRAL
<collectingCountry id="C6BF65E2FFFDB600B507FB45FD04FB32" box="[610,742,1231,1257]" name="Vietnam" pageId="7" pageNumber="493">VIETNAM</collectingCountry>
,
<collectingRegion id="7C6CEB90FFFDB600B595FB45FC06FB32" box="[752,996,1231,1257]" country="Vietnam" name="Quang Nam" pageId="7" pageNumber="493">Quang Nam province</collectingRegion>
,
<collectorName id="135D40A4FFFDB600B48BFB45FB69FB32" box="[1006,1163,1231,1257]" pageId="7" pageNumber="493">
Son Thanh
<taxonomicNameLabel id="97EF441BFFFDB600B30BFB45FB69FB32" box="[1134,1163,1231,1257]" pageId="7" pageNumber="493" rank="subSpecies">N.</taxonomicNameLabel>
</collectorName>
<collectionCode id="D8B9BDB7FFFDB600B3EEFB45FB40FB32" box="[1163,1186,1231,1257]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="493">R</collectionCode>
.,
<date id="CA1603B2FFFDB600B3D4FB45FB30FB32" box="[1201,1234,1231,1257]" pageId="7" pageNumber="493" value="2014-05-30">30.</date>
</materialsCitation>
<materialsCitation id="0EC02F2FFFFDB600B3B4FB45FAC5FB32" ID-GBIF-Occurrence="3383539321" box="[1233,1319,1231,1257]" collectionCode="V" collectorName="Son Thanh N." country="Vietnam" isTailEnumeration="true" location="Quang Nam province" pageId="7" pageNumber="493" specimenCount="1" stateProvince="Quang Nam" typeStatus="paratype">
<collectionCode id="D8B9BDB7FFFDB600B3B4FB45FB04FB32" box="[1233,1254,1231,1257]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="493" type="Museum">V</collectionCode>
.2014
</materialsCitation>
;
<materialsCitation id="0EC02F2FFFFDB600B257FB45FE5AFAD6" ID-GBIF-Occurrence="3383539303" collectorName="Ngoc Linh N. P" country="Vietnam" location="Kon Tum province" pageId="7" pageNumber="493" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<collectingRegion id="7C6CEB90FFFDB600B257FB45FF1EFAD6" country="Vietnam" name="Kon Tum" pageId="7" pageNumber="493">Kon Tum province</collectingRegion>
,
<collectorName id="135D40A4FFFDB600B66CFB79FE5AFAD6" box="[265,440,1267,1293]" pageId="7" pageNumber="493">Ngoc Linh N.P</collectorName>
</materialsCitation>
.
<typeStatus id="61139BD0FFFDB600B6A1FB79FDD3FAD6" box="[452,561,1267,1293]" pageId="7" pageNumber="493" type="paratype">Paratypes</typeStatus>
<specimenCount id="A8AEEEFBFFFDB600B55FFB79FDBFFAD6" box="[570,605,1267,1293]" pageId="7" pageNumber="493" type="male">1♂</specimenCount>
<date id="CA1603B2FFFDB600B503FB79FD0BFAD6" box="[614,745,1267,1293]" pageId="7" pageNumber="493" value="2018-06-12">
12.
<collectionCode id="D8B9BDB7FFFDB600B5E3FB79FD45FAD6" box="[646,679,1267,1293]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="493">VI</collectionCode>
.2018
</date>
;
<specimenCount id="A8AEEEFBFFFDB600B593FB79FCE7FAD6" box="[758,773,1267,1293]" pageId="7" pageNumber="493" type="female"></specimenCount>
<date id="CA1603B2FFFDB600B460FB79FC69FAD6" box="[773,907,1267,1293]" pageId="7" pageNumber="493" value="1998-03-27">27.III.1998</date>
,
<materialsCitation id="0EC02F2FFFFDB600B4FCFB79FF5FFAEA" ID-GBIF-Occurrence="3383539306" collectionCode="VVL, ALM, PNT" country="Vietnam" elevation="2" pageId="7" pageNumber="493" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
forest valley at
<quantity id="79508897FFFDB600B329FB79FB4FFAD6" box="[1100,1197,1267,1293]" metricMagnitude="0" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="493" unit="m" value="1.6">
<elevation id="3585C241FFFDB600B329FB79FB4FFAD6" box="[1100,1197,1267,1293]" metricMagnitude="0" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="493" unit="m" value="1.6">1,600 m</elevation>
</quantity>
;
<collectionCode id="D8B9BDB7FFFDB600B3DFFB79FB14FAD6" box="[1210,1270,1267,1293]" pageId="7" pageNumber="493">VVL</collectionCode>
,
<collectionCode id="D8B9BDB7FFFDB600B267FB79FAA2FAD6" box="[1282,1344,1267,1293]" country="0" httpUri="http://grbio.org/cool/icw9-2ehs" name="Museum National Historie Naturelle" pageId="7" pageNumber="493">ALM</collectionCode>
&amp;
<collectionCode id="D8B9BDB7FFFDB600B20DFB79FA7EFAD6" box="[1384,1436,1267,1293]" pageId="7" pageNumber="493">PNT</collectionCode>
leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFFDB600B7A2FAB0FBBCFA8E" box="[199,1118,1338,1365]" pageId="7" pageNumber="493" type="distribution">
<paragraph id="BE172572FFFDB600B7A2FAB0FBBCFA8E" blockId="7.[151,1437,151,1401]" box="[199,1118,1338,1365]" pageId="7" pageNumber="493">
<emphasis id="8CDCF960FFFDB600B7A2FAB0FE81FA8F" bold="true" box="[199,355,1338,1364]" pageId="7" pageNumber="493">Distribution:</emphasis>
The new taxon is distributed from
<collectingRegion id="7C6CEB90FFFDB600B588FAB1FC97FA8E" box="[749,885,1339,1365]" country="Vietnam" name="Quang Nam" pageId="7" pageNumber="493">Quang Nam</collectingRegion>
to
<collectingRegion id="7C6CEB90FFFDB600B4FCFAB1FBB9FA8E" box="[921,1115,1339,1365]" country="Vietnam" name="Gia Lai" pageId="7" pageNumber="493">Gia Lai province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFFDB600B7A2FAD5FA79FAA2" box="[199,1435,1375,1401]" pageId="7" pageNumber="493" type="etymology">
<paragraph id="BE172572FFFDB600B7A2FAD5FA79FAA2" blockId="7.[151,1437,151,1401]" box="[199,1435,1375,1401]" pageId="7" pageNumber="493">
<emphasis id="8CDCF960FFFDB600B7A2FAD5FEAAFAA2" bold="true" box="[199,328,1375,1401]" pageId="7" pageNumber="493">Etymology</emphasis>
: The name
<emphasis id="8CDCF960FFFDB600B6A8FAD5FDFFFAA2" box="[461,541,1375,1401]" italics="true" pageId="7" pageNumber="493">aurelia</emphasis>
refers to the unusually bright golden yellow colour of the hindwing undersides.
</paragraph>
</subSubSection>
</treatment>
</document>