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<document id="1FBD4D311F103AE75BE154F65CCA782B" ID-CLB-Dataset="4048" ID-DOI="10.11646/zootaxa.5048.4.2" ID-GBIF-Dataset="ad9f7049-4351-4deb-945a-409f9bbb5076" ID-ISSN="1175-5326" ID-Zenodo-Dep="5556598" ID-ZooBank="6A450390-1C69-42CD-9682-2A766CB31338" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1633693424633" checkinUser="plazi" docAuthor="Monastyrskii, Alexander L. &amp; Lien, Vu Van" docDate="2021" docId="36019464FFF0B60DB7F2FC7DFAF8F93A" docLanguage="en" docName="zootaxa.5048.4.2.pdf" docOrigin="Zootaxa 5048 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pantoporia bieti subsp. aurantina Monastyrskii &amp; To 2021, subspecies nov." docType="treatment" docVersion="8" lastPageNumber="496" masterDocId="CA38EC1CFFFAB607B765FF8AFFE2FFDB" masterDocTitle="New taxa and new records of butterflies from Vietnam (Lepidoptera, Papilionoidea)" masterLastPageNumber="510" masterPageNumber="486" pageNumber="496" updateTime="1699459186830" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="894082E454BDC96AFAB6782B91F842B6">New taxa and new records of butterflies from Vietnam (Lepidoptera, Papilionoidea)</mods:title>
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<mods:namePart id="E13CBF6554A07829FE5914771298FEEC">Monastyrskii, Alexander L.</mods:namePart>
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<mods:affiliation id="AB62A0547A9803D997111603D026B465">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam https: // orcid. org / 0000 - 0001 - 7645 - 2458 &amp; Fauna &amp; Flora International, Vietnam Programme, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="19AB035AD926838C7C89AF0A6F0A3DAB">Lien, Vu Van</mods:namePart>
<mods:nameIdentifier id="AE4A5485A84A039C5FFA0904054E0156" type="ORCID">0000-0001-7645-2458</mods:nameIdentifier>
<mods:affiliation id="95E748EB473D9FE48ACFF8A396A0907F">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam https: // orcid. org / 0000 - 0001 - 7645 - 2458 &amp; vulien @ gmail. com; https: // orcid. org / 0000 - 0003 - 2389 - 9402</mods:affiliation>
<mods:nameIdentifier id="7D0CC35201742F520A9934A0F39537AB" type="email">vulien@gmail.com</mods:nameIdentifier>
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<treatment id="36019464FFF0B60DB7F2FC7DFAF8F93A" ID-DOI="http://doi.org/10.5281/zenodo.5570235" ID-GBIF-Taxon="188639114" ID-Zenodo-Dep="5570235" LSID="urn:lsid:plazi:treatment:36019464FFF0B60DB7F2FC7DFAF8F93A" httpUri="http://treatment.plazi.org/id/36019464FFF0B60DB7F2FC7DFAF8F93A" lastPageNumber="496" pageId="10" pageNumber="496">
<subSubSection id="F6B276F9FFF0B60DB7F2FC7DFC7BFBCA" box="[151,921,1014,1041]" pageId="10" pageNumber="496" type="nomenclature">
<paragraph id="BE172572FFF0B60DB7F2FC7DFC7BFBCA" blockId="10.[151,921,1014,1077]" box="[151,921,1014,1041]" pageId="10" pageNumber="496">
<heading id="E55F921EFFF0B60DB7F2FC7DFC7BFBCA" bold="true" box="[151,921,1014,1041]" fontSize="11" level="1" pageId="10" pageNumber="496" reason="1">
<emphasis id="8CDCF960FFF0B60DB7F2FC7DFC7BFBCA" bold="true" box="[151,921,1014,1041]" pageId="10" pageNumber="496">
<taxonomicName id="79A85EF1FFF0B60DB7F2FC7DFD2DFBCA" authority="Monastyrskii &amp; To" authorityName="Monastyrskii &amp; To" authorityYear="2021" box="[151,719,1014,1041]" class="Insecta" family="Nymphalidae" genus="Pantoporia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="subSpecies" species="bieti" status="subsp. nov." subSpecies="aurantina">
<emphasis id="8CDCF960FFF0B60DB7F2FC7DFE38FBCA" bold="true" box="[151,474,1014,1041]" italics="true" pageId="10" pageNumber="496">Pantoporia bieti aurantina</emphasis>
Monastyrskii &amp; To
</taxonomicName>
,
<taxonomicNameLabel id="97EF441BFFF0B60DB5B9FC7CFC7BFBCA" box="[732,921,1014,1041]" pageId="10" pageNumber="496" rank="subSpecies">subspecies nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFF0B60DB7F2FB91FD07FA8E" pageId="10" pageNumber="496" type="description">
<paragraph id="BE172572FFF0B60DB7F2FB91FEC9FBEE" blockId="10.[151,921,1014,1077]" box="[151,299,1051,1077]" pageId="10" pageNumber="496">
(
<figureCitation id="269339F7FFF0B60DB7FAFB91FEC0FBEE" box="[159,290,1051,1077]" captionStart="FIGURE 7" captionStartId="11.[151,250,1424,1448]" captionTargetBox="[156,1437,191,1394]" captionTargetId="figure-20@11.[150,1437,181,1400]" captionTargetPageId="11" captionText="FIGURE 7.—Pantoporia bieti aurantina Monastyrskii &amp; To, subspec. nov. AB—♀HT, central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; CD—P. b. paona Tytler, 1915, Type, ♀, Kirbari, Naga Hills; EF—P. b. bieti (Oberthür, 1894), Type, ♂, western China, Ta-Tsien-Lou" figureDoi="http://doi.org/10.5281/zenodo.5556618" httpUri="https://zenodo.org/record/5556618/files/figure.png" pageId="10" pageNumber="496">Figs. 7A, B</figureCitation>
)
</paragraph>
<paragraph id="BE172572FFF0B60DB7F2FBE8FD1CFBA6" blockId="10.[151,1437,1122,1761]" box="[151,766,1122,1149]" pageId="10" pageNumber="496">
<emphasis id="8CDCF960FFF0B60DB7F2FBE8FEC9FBA7" bold="true" box="[151,299,1122,1148]" pageId="10" pageNumber="496">Description:</emphasis>
Female: Length of forewing: HT
<quantity id="79508897FFF0B60DB5C2FBE8FD1BFBA6" box="[679,761,1122,1149]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="496" unit="mm" value="25.0">25 mm</quantity>
.
</paragraph>
<paragraph id="BE172572FFF0B60DB7A2FB0DFD62FB32" blockId="10.[151,1437,1122,1761]" pageId="10" pageNumber="496">
<emphasis id="8CDCF960FFF0B60DB7A2FB0DFE31FB7B" bold="true" box="[199,467,1158,1185]" pageId="10" pageNumber="496">Both wings upperside.</emphasis>
(
<figureCitation id="269339F7FFF0B60DB687FB0DFDBBFB7A" box="[482,601,1159,1185]" captionStart="FIGURE 7" captionStartId="11.[151,250,1424,1448]" captionTargetBox="[156,1437,191,1394]" captionTargetId="figure-20@11.[150,1437,181,1400]" captionTargetPageId="11" captionText="FIGURE 7.—Pantoporia bieti aurantina Monastyrskii &amp; To, subspec. nov. AB—♀HT, central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; CD—P. b. paona Tytler, 1915, Type, ♀, Kirbari, Naga Hills; EF—P. b. bieti (Oberthür, 1894), Type, ♂, western China, Ta-Tsien-Lou" figureDoi="http://doi.org/10.5281/zenodo.5556618" httpUri="https://zenodo.org/record/5556618/files/figure.png" pageId="10" pageNumber="496">Figure 7A</figureCitation>
) Ground colour is black; all spots and bands are uniformly bright orange, including large discal cell streak merged with the streak beyond discal cell; lower postdiscal large spots in cells A1+A2 and Cu
<subScript id="222C2737FFF0B60DB626FB6BFEB0FB34" attach="left" box="[323,338,1249,1263]" fontSize="6" pageId="10" pageNumber="496">1a</subScript>
are connected in cell Cu
<subScript id="222C2737FFF0B60DB502FB6BFD9AFB34" attach="left" box="[615,632,1249,1263]" fontSize="6" pageId="10" pageNumber="496">1b</subScript>
;
</paragraph>
<paragraph id="BE172572FFF0B60DB7A2FB79FD07FA8E" blockId="10.[151,1437,1122,1761]" pageId="10" pageNumber="496">
<emphasis id="8CDCF960FFF0B60DB7A2FB79FE30FAD7" bold="true" box="[199,466,1266,1293]" pageId="10" pageNumber="496">Both wings underside.</emphasis>
(
<figureCitation id="269339F7FFF0B60DB685FB79FDB6FAD6" box="[480,596,1267,1293]" captionStart="FIGURE 7" captionStartId="11.[151,250,1424,1448]" captionTargetBox="[156,1437,191,1394]" captionTargetId="figure-20@11.[150,1437,181,1400]" captionTargetPageId="11" captionText="FIGURE 7.—Pantoporia bieti aurantina Monastyrskii &amp; To, subspec. nov. AB—♀HT, central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; CD—P. b. paona Tytler, 1915, Type, ♀, Kirbari, Naga Hills; EF—P. b. bieti (Oberthür, 1894), Type, ♂, western China, Ta-Tsien-Lou" figureDoi="http://doi.org/10.5281/zenodo.5556618" httpUri="https://zenodo.org/record/5556618/files/figure.png" pageId="10" pageNumber="496">Figure 7B</figureCitation>
) Ground colour of both wings dark reddish brown; all spots and bands are yellow; markings broad and distinct; wing patterns similar to those on the upperside except for a series of distinct pale violet subcostal spots in cells R
<subScript id="222C2737FFF0B60DB548FAC7FDD7FA80" attach="left" box="[557,565,1357,1371]" fontSize="6" pageId="10" pageNumber="496">1</subScript>
, R
<subScript id="222C2737FFF0B60DB533FAC7FDBCFA80" attach="left" box="[598,606,1357,1371]" fontSize="6" pageId="10" pageNumber="496">2</subScript>
, R
<subScript id="222C2737FFF0B60DB51AFAC7FD65FA80" attach="left" box="[639,647,1357,1371]" fontSize="6" pageId="10" pageNumber="496">5</subScript>
and M
<subScript id="222C2737FFF0B60DB5B3FAC7FD3CFA80" attach="left" box="[726,734,1357,1371]" fontSize="6" pageId="10" pageNumber="496">1</subScript>
.
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFF0B60DB7A2FAD4FE9FF942" pageId="10" pageNumber="496" type="diagnosis">
<paragraph id="BE172572FFF0B60DB7A2FAD4FE9FF942" blockId="10.[151,1437,1122,1761]" pageId="10" pageNumber="496">
<emphasis id="8CDCF960FFF0B60DB7A2FAD4FEA6FAA3" bold="true" box="[199,324,1374,1400]" pageId="10" pageNumber="496">Diagnosis:</emphasis>
The new taxon may be distinguished from individuals of other subspecies distributed in western
<collectingCountry id="C6BF65E2FFF0B60DB7F2FA09FF38FA46" box="[151,218,1411,1437]" name="China" pageId="10" pageNumber="496">China</collectingCountry>
(the nominate subspecies) (
<figureCitation id="269339F7FFF0B60DB575FA09FD40FA46" box="[528,674,1411,1437]" captionStart="FIGURE 7" captionStartId="11.[151,250,1424,1448]" captionTargetBox="[156,1437,191,1394]" captionTargetId="figure-20@11.[150,1437,181,1400]" captionTargetPageId="11" captionText="FIGURE 7.—Pantoporia bieti aurantina Monastyrskii &amp; To, subspec. nov. AB—♀HT, central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; CD—P. b. paona Tytler, 1915, Type, ♀, Kirbari, Naga Hills; EF—P. b. bieti (Oberthür, 1894), Type, ♂, western China, Ta-Tsien-Lou" figureDoi="http://doi.org/10.5281/zenodo.5556618" httpUri="https://zenodo.org/record/5556618/files/figure.png" pageId="10" pageNumber="496">Figure 7EF</figureCitation>
), Naga Hills and northern
<collectingCountry id="C6BF65E2FFF0B60DB4A3FA09FBC7FA46" box="[966,1061,1411,1437]" name="Vietnam" pageId="10" pageNumber="496">Vietnam</collectingCountry>
(
<taxonomicName id="79A85EF1FFF0B60DB356FA09FB03FA47" box="[1075,1249,1411,1437]" pageId="10" pageNumber="496" rank="subSpecies" subSpecies="paona">
subspec.
<emphasis id="8CDCF960FFF0B60DB3FEFA09FB03FA47" box="[1179,1249,1411,1436]" italics="true" pageId="10" pageNumber="496">paona</emphasis>
</taxonomicName>
) (
<figureCitation id="269339F7FFF0B60DB39DFA09FA73FA46" box="[1272,1425,1411,1437]" captionStart="FIGURE 7" captionStartId="11.[151,250,1424,1448]" captionTargetBox="[156,1437,191,1394]" captionTargetId="figure-20@11.[150,1437,181,1400]" captionTargetPageId="11" captionText="FIGURE 7.—Pantoporia bieti aurantina Monastyrskii &amp; To, subspec. nov. AB—♀HT, central Vietnam, Gia Lai province, Kon Ka Kinh N.P.; CD—P. b. paona Tytler, 1915, Type, ♀, Kirbari, Naga Hills; EF—P. b. bieti (Oberthür, 1894), Type, ♂, western China, Ta-Tsien-Lou" figureDoi="http://doi.org/10.5281/zenodo.5556618" httpUri="https://zenodo.org/record/5556618/files/figure.png" pageId="10" pageNumber="496">Figure 7CD</figureCitation>
). It has well developed, broad, bright orange markings on the upperside of both wings, while the other subspecies have whitish (eastern
<collectingRegion id="7C6CEB90FFF0B60DB6EAFA41FE2FFA3E" box="[399,461,1483,1509]" country="China" name="Xizang" pageId="10" pageNumber="496">Tibet</collectingRegion>
;
<bibRefCitation id="DA395883FFF0B60DB6BCFA41FD95FA3E" author="Bozano, G. C." box="[473,631,1483,1509]" pageId="10" pageNumber="496" pagination="1 - 77" refId="ref11908" refString="Bozano, G. C. (2008) In: Bozano, G. C. (Ed.), Guide to the Butterflies of the Palearctic Region. Nymphalidae Part III. Subfamily Limenitidinae, Tribe Neptini. Omnes Artes, Milano, pp. 1 - 77." type="book chapter" year="2008">Bozano, 2008</bibRefCitation>
) or yellowish markings (the nominate
<taxonomicName id="79A85EF1FFF0B60DB34AFA41FB2EFA3E" box="[1071,1228,1483,1509]" pageId="10" pageNumber="496" rank="subSpecies" subSpecies="from">subspec. from</taxonomicName>
central
<collectingCountry id="C6BF65E2FFF0B60DB24CFA41FA8EFA3E" box="[1321,1388,1483,1509]" name="China" pageId="10" pageNumber="496">China</collectingCountry>
and
<taxonomicName id="79A85EF1FFF0B60DB7F2FA65FEA4F9D3" box="[151,326,1519,1545]" pageId="10" pageNumber="496" rank="subSpecies" subSpecies="paona">
subspec.
<emphasis id="8CDCF960FFF0B60DB665FA65FEA4F9D3" box="[256,326,1519,1544]" italics="true" pageId="10" pageNumber="496">paona</emphasis>
</taxonomicName>
from northeastern
<collectingCountry id="C6BF65E2FFF0B60DB547FA65FDBEF9D2" box="[546,604,1519,1545]" name="India" pageId="10" pageNumber="496">India</collectingCountry>
(ex
<taxonomicName id="79A85EF1FFF0B60DB5F7FA65FCFBF9D2" box="[658,793,1519,1545]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="10" pageNumber="496" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
collection of the NHM). The most unusual character of the new taxon is the well-developed series of subcostal spots on the underside of the forewing. Individuals of other subspecies either lack this character or it is unclear. In addition, subcostal spots on the forewing are not typical for
<taxonomicName id="79A85EF1FFF0B60DB7F2F9D1FEF5F9AF" authorityName="Hubner" authorityYear="1819" box="[151,279,1627,1652]" class="Insecta" family="Nymphalidae" genus="Pantoporia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="genus">
<emphasis id="8CDCF960FFF0B60DB7F2F9D1FEF5F9AF" box="[151,279,1627,1652]" italics="true" pageId="10" pageNumber="496">Pantoporia</emphasis>
</taxonomicName>
. This character is usually observed in some
<taxonomicName id="79A85EF1FFF0B60DB463F9D1FCACF9AF" box="[774,846,1627,1652]" class="Insecta" family="Nymphalidae" genus="Neptis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="genus">
<emphasis id="8CDCF960FFF0B60DB463F9D1FCACF9AF" box="[774,846,1627,1652]" italics="true" pageId="10" pageNumber="496">Neptis</emphasis>
</taxonomicName>
with orange markings, such as the
<taxonomicName id="79A85EF1FFF0B60DB3BDF9D1FAA7F9AE" authorityName="Oberthur" authorityYear="1876" box="[1240,1349,1627,1653]" class="Insecta" family="Nymphalidae" genus="Neptis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="species" species="armandia">
<emphasis id="8CDCF960FFF0B60DB3BDF9D1FAA7F9AE" box="[1240,1349,1627,1653]" italics="true" pageId="10" pageNumber="496">armandia</emphasis>
</taxonomicName>
,
<emphasis id="8CDCF960FFF0B60DB236F9D1FA7EF9AE" box="[1363,1436,1627,1653]" italics="true" pageId="10" pageNumber="496">themis</emphasis>
and
<emphasis id="8CDCF960FFF0B60DB7A2F9F5FEC1F942" box="[199,291,1663,1689]" italics="true" pageId="10" pageNumber="496">antilope</emphasis>
groups.
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFF0B60DB7A2F929FA6BF966" box="[199,1417,1698,1725]" pageId="10" pageNumber="496" type="materials_examined">
<paragraph id="BE172572FFF0B60DB7A2F929FA6BF966" blockId="10.[151,1437,1122,1761]" box="[199,1417,1698,1725]" pageId="10" pageNumber="496">
<emphasis id="8CDCF960FFF0B60DB7A2F929FE95F967" bold="true" box="[199,375,1698,1725]" pageId="10" pageNumber="496">Type material:</emphasis>
<materialsCitation id="0EC02F2FFFF0B60DB61BF929FA6BF966" ID-GBIF-Occurrence="3383539338" box="[382,1417,1699,1725]" collectingDate="2016-04" collectorName="Kon Ka Kinh N. P." country="Vietnam" location="Gia Lai province" pageId="10" pageNumber="496" specimenCount="1" specimenCount-female="1" stateProvince="Gia Lai" typeStatus="holotype">
<typeStatus id="61139BD0FFF0B60DB61BF929FE09F966" box="[382,491,1699,1725]" pageId="10" pageNumber="496" type="holotype">Holotype</typeStatus>
: CENTRAL
<collectingCountry id="C6BF65E2FFF0B60DB519F929FCE2F966" box="[636,768,1699,1725]" name="Vietnam" pageId="10" pageNumber="496">VIETNAM</collectingCountry>
,
<specimenCount id="A8AEEEFBFFF0B60DB46EF929FCC2F966" box="[779,800,1699,1725]" pageId="10" pageNumber="496" type="female"></specimenCount>
<collectingRegion id="7C6CEB90FFF0B60DB442F929FC0AF966" box="[807,1000,1699,1725]" country="Vietnam" name="Gia Lai" pageId="10" pageNumber="496">Gia Lai province</collectingRegion>
,
<collectorName id="135D40A4FFF0B60DB496F929FB20F966" box="[1011,1218,1699,1725]" pageId="10" pageNumber="496">Kon Ka Kinh N.P.</collectorName>
,
<date id="CA1603B2FFF0B60DB3A8F929FACBF966" box="[1229,1321,1699,1725]" pageId="10" pageNumber="496" value="2016-04">
<collectingDate id="DA52FA5AFFF0B60DB3A8F929FACBF966" box="[1229,1321,1699,1725]" pageId="10" pageNumber="496" value="2016-04">IV.2016</collectingDate>
</date>
, TQ leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F6B276F9FFF0B60DB7A2F94CFAF8F93A" box="[199,1306,1734,1761]" pageId="10" pageNumber="496" type="distribution">
<paragraph id="BE172572FFF0B60DB7A2F94CFAF8F93A" blockId="10.[151,1437,1122,1761]" box="[199,1306,1734,1761]" pageId="10" pageNumber="496">
<emphasis id="8CDCF960FFF0B60DB7A2F94CFE81F93B" bold="true" box="[199,355,1734,1760]" pageId="10" pageNumber="496">Distribution:</emphasis>
Currently only recorded from Central
<collectingCountry id="C6BF65E2FFF0B60DB471F94DFC94F93A" box="[788,886,1735,1761]" name="Vietnam" pageId="10" pageNumber="496">Vietnam</collectingCountry>
,
<collectingRegion id="7C6CEB90FFF0B60DB4E4F94DFBA0F93A" box="[897,1090,1735,1761]" country="Vietnam" name="Gia Lai" pageId="10" pageNumber="496">Gia Lai province</collectingRegion>
, Kon Ka Kinh N.P.
</paragraph>
</subSubSection>
</treatment>
</document>