563 lines
49 KiB
XML
563 lines
49 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.98.86299" ID-Pensoft-Pub="1860-0743-2-435" ID-Pensoft-UUID="3DE923FDF8B4597695BC025E41200D64" ID-ZooBank="6EA087B23245455FAD1015016E8417D3" ModsDocID="1860-0743-98-2-435" checkinTime="1669095670786" checkinUser="pensoft" docAuthor="Conradie, Werner, Schmitz, Andreas, Lobon-Rovira, Javier, Becker, Francois S., Vaz Pinto, Pedro & Hauptfleisch, Morgan L." docDate="2022" docId="17D6C5F2F3015BFE8260B5DDFB4278F8" docLanguage="en" docName="ZoosystEvol 98(2): 435-453" docOrigin="Zoosystematics and Evolution 98 (2)" docPubDate="2022-11-21" docSource="http://dx.doi.org/10.3897/zse.98.86299" docTitle="Afroedura otjihipa Conradie & Schmitz & Lobón-Rovira & Becker & Vaz Pinto & Hauptfleisch 2022, sp. nov." docType="treatment" docUuid="B3818B24-7FF0-49B6-B6F9-F580BF50C0C0" docUuidSource="ZooBank" docVersion="2" id="3DE923FDF8B4597695BC025E41200D64" lastPageNumber="435" masterDocId="3DE923FDF8B4597695BC025E41200D64" masterDocTitle="Rock island melody remastered: two new species in the Afroedura bogerti Loveridge, 1944 group from Angola and Namibia" masterLastPageNumber="453" masterPageNumber="435" pageNumber="435" updateTime="1669095972909" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Rock island melody remastered: two new species in the Afroedura bogerti Loveridge, 1944 group from Angola and Namibia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Conradie, Werner</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0805-9683</mods:nameIdentifier>
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<mods:affiliation>Port Elizabeth Museum (Bayworld), P. O. Box 13147, Humewood 6013, South Africa & Department of Nature Conservation Management, Natural Resource Science and Management Cluster, Faculty of Science, George Campus, Nelson Mandela University, George, South Africa</mods:affiliation>
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<mods:nameIdentifier type="email">werner@bayworld.co.za</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schmitz, Andreas</mods:namePart>
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<mods:affiliation>Natural History Museum of Geneva, Route de Malagnou 1, C. P. 6434, 1211 Geneva 6, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lobon-Rovira, Javier</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4380-9427</mods:nameIdentifier>
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<mods:affiliation>CIBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, InBIO Laboratorio Associado, Campus de Vairao, Universidade do Porto, 4485 - 661 Vairao, Portugal & Departamento de Biologia, Faculdade de Ciencias, Universidade do Porto, 4099 - 002 Porto, Portugal</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Becker, Francois S.</mods:namePart>
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<mods:affiliation>BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661 Vairao, Portugal de Ciencias da Educacao da Huila (ISCED-Huila), Rua Sarmento Rodrigues, Lubango, Angola & National Museum of Namibia, Windhoek, Namibia</mods:affiliation>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vaz Pinto, Pedro</mods:namePart>
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<mods:affiliation>CIBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, InBIO Laboratorio Associado, Campus de Vairao, Universidade do Porto, 4485 - 661 Vairao, Portugal & Departamento de Biologia, Faculdade de Ciencias, Universidade do Porto, 4099 - 002 Porto, Portugal & School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Private Bag 3, Wits 2050, Johannesburg, South Africa & Fundacao Kissama, Luanda, Angola</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hauptfleisch, Morgan L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0654-0887</mods:nameIdentifier>
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<mods:affiliation>TwinLab CIBIO / ISCED - Instituto de Ciencias da Educacao da Huila, Rua Sarmento Rodrigues 2, C. P. 230, Lubango, Angola</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-11-21</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>98</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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<mods:start>435</mods:start>
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<mods:end>453</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.98.86299</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.98.86299</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-2-435</mods:identifier>
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<mods:identifier type="ZooBank">6EA087B23245455FAD1015016E8417D3</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:B3818B24-7FF0-49B6-B6F9-F580BF50C0C0" httpUri="http://treatment.plazi.org/id/17D6C5F2F3015BFE8260B5DDFB4278F8" lastPageNumber="435" pageId="0" pageNumber="435">
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<subSubSection pageId="0" pageNumber="435" type="nomenclature">
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<paragraph pageId="0" pageNumber="435">
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<taxonomicName LSID="https://zoobank.org/B3818B24-7FF0-49B6-B6F9-F580BF50C0C0" authority="Conradie & Schmitz & Lobón-Rovira & Becker & Vaz Pinto & Hauptfleisch, 2022" authorityName="Conradie & Schmitz & Lobón-Rovira & Becker & Vaz Pinto & Hauptfleisch" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura otjihipa" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="otjihipa" status="sp. nov.">Afroedura otjihipa</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="435">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="435" type="description">
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<paragraph pageId="0" pageNumber="435">
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitat photos of: A-B. Afroedura pundomontana sp. nov.: Morro do Pundo, 25 km west of Bocoio, Benguela Province, Angola; C-D. Afroedura otjihipa sp. nov.: Otjihipa Middleberg, Kunene Region, Namibia. Photos: A-B. Pedro Vaz Pinto; C-D. Francois Becker." figureDoi="10.3897/zse.98.86299.figure5" httpUri="https://binary.pensoft.net/fig/772233" pageId="0" pageNumber="435">Otjihipa Flat Gecko (English) Otjihipa Platgeitjie (Afrikaans) Figs 5C-D</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Live specimens of: A-B. Afroedura donveae from Omahua, Namibe Province, Angola (not sampled); C-D. Afroedura otjihipa sp. nov. (holotype female, NMNW R 11253) from Otjihipa Middleberg, Kunene Region, Namibia. Photos: A-B. Javier Lobon-Rovira; C-D. Francois Becker." figureDoi="10.3897/zse.98.86299.figure6" httpUri="https://binary.pensoft.net/fig/772234" pageId="0" pageNumber="435">, 6C-D</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Afroedura otjihipa sp. nov. (NMNW R 11253) from Otjihipa Middleberg, Kunene Region, Namibia. Scale bar: 1 cm. Photos: Francois Becker." figureDoi="10.3897/zse.98.86299.figure7" httpUri="https://binary.pensoft.net/fig/772235" pageId="0" pageNumber="435">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="435" type="synonym">
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<paragraph pageId="0" pageNumber="435">Synonym.</paragraph>
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<paragraph pageId="0" pageNumber="435">
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<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura bogerti" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="bogerti">Afroedura cf. bogerti</taxonomicName>
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-
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<bibRefCitation DOI="https://doi.org/10.2307/1447414" author="Branch, WR" journalOrPublisher="Struik, Pretoria" pageId="0" pageNumber="435" refId="B6" refString="Branch, WR, 1998. Field Guide to Snakes and other Reptiles of Southern Africa. Struik, Pretoria, DOI: https://doi.org/10.2307/1447414" title="Field Guide to Snakes and other Reptiles of Southern Africa." url="https://doi.org/10.2307/1447414" year="1998">Branch 1998</bibRefCitation>
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: 232;
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<bibRefCitation author="Griffin, M" journalOrPublisher="Directorate of Scientific Services, Ministry of Environment and Tourism, Windhoek" pageId="0" pageNumber="435" publicationUrl="https://www.lacerta.de/AF/Bibliografie/BIB_4908.pdf" refId="B19" refString="Griffin, M, 2002. Annotated Checklist and Provisional National Conservation Status of Namibian Reptiles. Technical Reports of Scientific Services. Number 1. Directorate of Scientific Services, Ministry of Environment and Tourism, Windhoek, https://www.lacerta.de/AF/Bibliografie/BIB_4908.pdf" title="Annotated Checklist and Provisional National Conservation Status of Namibian Reptiles. Technical Reports of Scientific Services. Number 1." url="https://www.lacerta.de/AF/Bibliografie/BIB_4908.pdf" year="2002">Griffin 2002</bibRefCitation>
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: 20, 2003:10;
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<bibRefCitation DOI="https://doi.org/10.1016/j.jaridenv.2012.05.003" author="Herrman, H-W" journalOrPublisher="Journal of Arid Environments" pageId="0" pageNumber="435" pagination="94 - 115" refId="B22" refString="Herrman, H-W, Branch, WR, 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94 - 115, DOI: https://doi.org/10.1016/j.jaridenv.2012.05.003" title="Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist." url="https://doi.org/10.1016/j.jaridenv.2012.05.003" volume="93" year="2013">Herrman and Branch 2013</bibRefCitation>
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: 5.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="435" type="holotype">
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<paragraph pageId="0" pageNumber="435">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="435">NMNW R11253, adult female, collected from Otjihipa Middleberg (-17.28314, 12.66506, 1,900 m a.s.l.), Kunene Region, Namibia, by Morgan Hauptfleisch, Francois Becker, Vera De Cauwer, Wessel Swanepoel and Ernst van Jaarsveld on 23 April 2021.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="435" type="paratype">
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<paragraph pageId="0" pageNumber="435">Paratype.</paragraph>
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<paragraph pageId="0" pageNumber="435">NMNW R11245, adult male (paired with female NMNW R11253 in same rock crack). Same collection details as holotype.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="435" type="etymology">
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<paragraph pageId="0" pageNumber="435">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="435">The new species is named in reference to the area it was collected, namely Otjihipa Mountains in northern Namibia.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="435" type="diagnosis">
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<paragraph pageId="0" pageNumber="435">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="435">
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A member of the greater '
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<taxonomicName baseAuthorityName="Hewitt" baseAuthorityYear="1925" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura transvaalica" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="transvaalica">
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<emphasis italics="true" pageId="0" pageNumber="435">Afroedura transvaalica</emphasis>
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</taxonomicName>
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' group as it possesses two pairs of enlarged scansors per digit and a strongly verticillate and flattened tail (
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3846.4.1" author="Jacobsen, NHG" journalOrPublisher="Zootaxa" pageId="0" pageNumber="435" pagination="451 - 501" refId="B24" refString="Jacobsen, NHG, Kuhn, AL, Jackman, TR, Bauer, AM, 2014. A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa. Zootaxa 3846 (4): 451 - 501, DOI: https://doi.org/10.11646/zootaxa.3846.4.1" title="A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa." url="https://doi.org/10.11646/zootaxa.3846.4.1" volume="3846" year="2014">Jacobsen et al. 2014</bibRefCitation>
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). Part of the
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<taxonomicName lsidName="A. bogerti" pageId="0" pageNumber="435" rank="species" species="bogerti">
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<emphasis italics="true" pageId="0" pageNumber="435">A. bogerti</emphasis>
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</taxonomicName>
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group which differs from other members of the '
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<taxonomicName baseAuthorityName="Hewitt" baseAuthorityYear="1925" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura transvaalica" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="transvaalica">
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<emphasis italics="true" pageId="0" pageNumber="435">Afroedura transvaalica</emphasis>
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</taxonomicName>
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' group by having less than 72 mid-body scale rows (vs. 97-102 in
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<taxonomicName lsidName="A. gorongosa" pageId="0" pageNumber="435" rank="species" species="gorongosa">
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<emphasis italics="true" pageId="0" pageNumber="435">A. gorongosa</emphasis>
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</taxonomicName>
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, 113-120 in
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<taxonomicName lsidName="A. loveridgei" pageId="0" pageNumber="435" rank="species" species="loveridgei">
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<emphasis italics="true" pageId="0" pageNumber="435">A. loveridgei</emphasis>
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</taxonomicName>
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, 102-119 in
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<taxonomicName lsidName="A. transvaalica" pageId="0" pageNumber="435" rank="species" species="transvaalica">
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<emphasis italics="true" pageId="0" pageNumber="435">A. transvaalica</emphasis>
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</taxonomicName>
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); rostral excluded from the nostril (in contact in
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<taxonomicName lsidName="A. gorongosa" pageId="0" pageNumber="435" rank="species" species="gorongosa">
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<emphasis italics="true" pageId="0" pageNumber="435">A. gorongosa</emphasis>
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</taxonomicName>
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); supranasals always in contact (separated by 1-3 granules in
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<taxonomicName lsidName="A. gorongosa" pageId="0" pageNumber="435" rank="species" species="gorongosa">
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<emphasis italics="true" pageId="0" pageNumber="435">A. gorongosa</emphasis>
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</taxonomicName>
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; always in broad contact in
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<taxonomicName lsidName="A. loveridgei" pageId="0" pageNumber="435" rank="species" species="loveridgei">
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<emphasis italics="true" pageId="0" pageNumber="435">A. loveridgei</emphasis>
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</taxonomicName>
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; usually in broad contact in
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<taxonomicName lsidName="A. transvaalica" pageId="0" pageNumber="435" rank="species" species="transvaalica">
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<emphasis italics="true" pageId="0" pageNumber="435">A. transvaalica</emphasis>
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</taxonomicName>
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~ 3-18%); and 15-16 scales between anterior borders of the eyes (19-22 in
|
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<taxonomicName lsidName="A. gorongosa" pageId="0" pageNumber="435" rank="species" species="gorongosa">
|
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<emphasis italics="true" pageId="0" pageNumber="435">A. gorongosa</emphasis>
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</taxonomicName>
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, 15-19 in
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<taxonomicName lsidName="A. loveridgei" pageId="0" pageNumber="435" rank="species" species="loveridgei">
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<emphasis italics="true" pageId="0" pageNumber="435">A. loveridgei</emphasis>
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</taxonomicName>
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, 15-20 in
|
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<taxonomicName lsidName="A. transvaalica" pageId="0" pageNumber="435" rank="species" species="transvaalica">
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<emphasis italics="true" pageId="0" pageNumber="435">A. transvaalica</emphasis>
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</taxonomicName>
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) (comparative data from
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4324.1.8" author="Branch, WR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="435" pagination="142 - 160" refId="B7" refString="Branch, WR, Guyton, JA, Schmitz, A, Barej, MF, Naskrecki, P, Farooq, H, Verburgt, L, Roedel, M-O, 2017. Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique. Zootaxa 4324 (1): 142 - 160, DOI: https://doi.org/10.11646/zootaxa.4324.1.8" title="Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique." url="https://doi.org/10.11646/zootaxa.4324.1.8" volume="4324" year="2017">Branch et al. 2017</bibRefCitation>
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,
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<bibRefCitation DOI="https://doi.org/10.3897/zse.97.57202" author="Branch, WR" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="435" pagination="55 - 82" refId="B8" refString="Branch, WR, Schmitz, A, Lobon-Rovira, J, Baptista, NL, Antonio, T, Conradie, W, 2021. Rock island melody: A revision of the Afroedura bogerti Loveridge, 1944 group, with the description of four new endemic species from Angola. Zoosystematics and Evolution 97 (1): 55 - 82, DOI: https://doi.org/10.3897/zse.97.57202" title="Rock island melody: A revision of the Afroedura bogerti Loveridge, 1944 group, with the description of four new endemic species from Angola." url="https://doi.org/10.3897/zse.97.57202" volume="97" year="2021">2021</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="435">
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<taxonomicName authorityName="Conradie & Schmitz & Lobón-Rovira & Becker & Vaz Pinto & Hauptfleisch" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura otjihipa" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="otjihipa">
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<emphasis italics="true" pageId="0" pageNumber="435">Afroedura otjihipa</emphasis>
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</taxonomicName>
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sp. nov. differs from other members of the
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<taxonomicName lsidName="A. bogerti" pageId="0" pageNumber="435" rank="species" species="bogerti">
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<emphasis italics="true" pageId="0" pageNumber="435">A. bogerti</emphasis>
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</taxonomicName>
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group by a combination of the following characteristics (see Tables
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<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Summary of morphological data for Afroedura bogerti, A. wulfhaackei (including members of the morphologically-indistinguishable subclades), A. donveae, A. vazpintorum (including isolated escarpment population), A. praedicta, A. pundomontana sp. nov. and A. otjihipa sp. nov. Values are given as a range with mean in parenthesis for scalation and mean + / - standard deviation for meristic ratios. M = male, F = female, n = sample size." httpUri="http://table.plazi.org/id/A05B6B175BAAE754ED3C075FFD08A1D8" pageId="0" pageNumber="435" tableUuid="A05B6B175BAAE754ED3C075FFD08A1D8">5</tableCitation>
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and
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<tableCitation captionStart="Table 7" captionStartId="T7" captionText="Table 7. Measurements (in mm) and scale counts for the type series of Afroedura otjihipa sp. nov." httpUri="http://table.plazi.org/id/4AA3093F14F3331B02665C8753C139FF" pageId="0" pageNumber="435" tableUuid="4AA3093F14F3331B02665C8753C139FF">7</tableCitation>
|
||
): 65-67 (mean 66.0) mid-body scale rows (64-78 [mean 72.8] in
|
||
<taxonomicName lsidName="A. donveae" pageId="0" pageNumber="435" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. donveae</emphasis>
|
||
</taxonomicName>
|
||
, 69-77 [mean 73.5] in
|
||
<taxonomicName lsidName="A. bogerti" pageId="0" pageNumber="435" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
, 73-78 [mean 74.8] in
|
||
<taxonomicName lsidName="A. praedicta" pageId="0" pageNumber="435" rank="species" species="praedicta">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. praedicta</emphasis>
|
||
</taxonomicName>
|
||
, 78-82 (mean 79.5) in
|
||
<taxonomicName lsidName="A. pundomontana" pageId="0" pageNumber="435" rank="species" species="pundomontana">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. pundomontana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; 76-88 [mean 79.3] in
|
||
<taxonomicName lsidName="A. wulfhaackei" pageId="0" pageNumber="435" rank="species" species="wulfhaackei">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. wulfhaackei</emphasis>
|
||
</taxonomicName>
|
||
, 73-86 [mean 80.3] in
|
||
<taxonomicName lsidName="A. vazpintorum" pageId="0" pageNumber="435" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
); supranasals always in contact (similar to
|
||
<taxonomicName lsidName="A. donveae" pageId="0" pageNumber="435" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. donveae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="A. vazpintorum" pageId="0" pageNumber="435" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="A. praedicta" pageId="0" pageNumber="435" rank="species" species="praedicta">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. praedicta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="A. pundomontana" pageId="0" pageNumber="435" rank="species" species="pundomontana">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. pundomontana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; in contact in ~ 33% of
|
||
<taxonomicName lsidName="A. bogerti" pageId="0" pageNumber="435" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
; in contact in ~ 57%
|
||
<emphasis italics="true" pageId="0" pageNumber="435">
|
||
of
|
||
<taxonomicName lsidName="A. wulfhaackei" pageId="0" pageNumber="435" rank="species" species="wulfhaackei">A. wulfhaackei</taxonomicName>
|
||
</emphasis>
|
||
); each tail verticil comprises 5 ventral and 6 dorsal rows of scales (mean 4 ventral and 5 dorsal in
|
||
<taxonomicName lsidName="A. bogerti" pageId="0" pageNumber="435" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="A. praedicta" pageId="0" pageNumber="435" rank="species" species="praedicta">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. praedicta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="A. wulfhaackei" pageId="0" pageNumber="435" rank="species" species="wulfhaackei">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. wulfhaackei</emphasis>
|
||
</taxonomicName>
|
||
; 4-5 (mean 4.4) ventral and 5-6 (mean 5.6) dorsal in
|
||
<taxonomicName lsidName="A. pundomontana" pageId="0" pageNumber="435" rank="species" species="pundomontana">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. pundomontana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; 5-6 [mean 5.5] ventral and 6-7 [mean 6.6] dorsal in
|
||
<taxonomicName lsidName="A. donveae" pageId="0" pageNumber="435" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. donveae</emphasis>
|
||
</taxonomicName>
|
||
; 5-6 [mean 5.0] ventral and 6-7 [mean 6.1] dorsal
|
||
<taxonomicName lsidName="A. vazpintorum" pageId="0" pageNumber="435" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
); ventral surfaces light cream and almost immaculate, with some scattered dark spots near lateral edges (similar to
|
||
<taxonomicName lsidName="A. donveae" pageId="0" pageNumber="435" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. donveae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="A. vazpintorum" pageId="0" pageNumber="435" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
; greyish with black spots in
|
||
<emphasis italics="true" pageId="0" pageNumber="435">
|
||
<taxonomicName lsidName="A. bogerti" pageId="0" pageNumber="435" rank="species" species="bogerti">A. bogerti</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="A. wulfhaackei" pageId="0" pageNumber="435" rank="species" species="wulfhaackei">A. wulfhaackei</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName lsidName="A. praedicta" pageId="0" pageNumber="435" rank="species" species="praedicta">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. praedicta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="A. pundomontana" pageId="0" pageNumber="435" rank="species" species="pundomontana">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. pundomontana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.); larger average adult size 58.2 mm SVL (versus 57.6 mm in
|
||
<taxonomicName lsidName="A. donveae" pageId="0" pageNumber="435" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. donveae</emphasis>
|
||
</taxonomicName>
|
||
, 51.7 mm in
|
||
<taxonomicName lsidName="A. wulfhaackei" pageId="0" pageNumber="435" rank="species" species="wulfhaackei">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. wulfhaackei</emphasis>
|
||
</taxonomicName>
|
||
, 51.3 mm in
|
||
<taxonomicName lsidName="A. vazpintorum" pageId="0" pageNumber="435" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
; 50.3 mm in
|
||
<taxonomicName lsidName="A. pundomontana" pageId="0" pageNumber="435" rank="species" species="pundomontana">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. pundomontana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., 50.0 mm in
|
||
<taxonomicName lsidName="A. bogerti" pageId="0" pageNumber="435" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
, 49.9 mm
|
||
<taxonomicName lsidName="A. praedicta" pageId="0" pageNumber="435" rank="species" species="praedicta">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. praedicta</emphasis>
|
||
</taxonomicName>
|
||
), and by having very distinct black-and-white tail banding (similar to
|
||
<taxonomicName lsidName="A. donveae" pageId="0" pageNumber="435" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. donveae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName authorityName="Conradie & Schmitz & Lobón-Rovira & Becker & Vaz Pinto & Hauptfleisch" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura otjihipa" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="otjihipa">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">Afroedura otjihipa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from its sister lowland species
|
||
<taxonomicName lsidName="A. donveae" pageId="0" pageNumber="435" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. donveae</emphasis>
|
||
</taxonomicName>
|
||
in having a brown or copper coloured (versus black) iris, a relatively broader head (mean HL/HW 1.1 versus 1.3), and in dorsal colour pattern (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Live specimens of: A-B. Afroedura donveae from Omahua, Namibe Province, Angola (not sampled); C-D. Afroedura otjihipa sp. nov. (holotype female, NMNW R 11253) from Otjihipa Middleberg, Kunene Region, Namibia. Photos: A-B. Javier Lobon-Rovira; C-D. Francois Becker." figureDoi="10.3897/zse.98.86299.figure6" httpUri="https://binary.pensoft.net/fig/772234" pageId="0" pageNumber="435">6</figureCitation>
|
||
): in
|
||
<taxonomicName lsidName="A. otjihipa" pageId="0" pageNumber="435" rank="species" species="otjihipa">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. otjihipa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. it is dominantly dark brown, the yellow appearing as small asymmetrical, irregular patches, and as irregular borders of four paired, asymmetrical, irregular, roughly triangular brown blotches, which merge at the scapular and sacral regions to form two additional bands (versus roughly symmetrical brown patterns on a mostly yellow background in
|
||
<taxonomicName lsidName="A. donveae" pageId="0" pageNumber="435" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. donveae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.98.86299.figure6" httpUri="https://binary.pensoft.net/fig/772234" pageId="0" pageNumber="435" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="435">
|
||
<emphasis bold="true" pageId="0" pageNumber="435">Figure 6.</emphasis>
|
||
Live specimens of:
|
||
<emphasis bold="true" pageId="0" pageNumber="435">A-B.</emphasis>
|
||
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura donveae" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">Afroedura donveae</emphasis>
|
||
</taxonomicName>
|
||
from Omahua, Namibe Province, Angola (not sampled);
|
||
<emphasis bold="true" pageId="0" pageNumber="435">C-D.</emphasis>
|
||
<taxonomicName authorityName="Conradie & Schmitz & Lobón-Rovira & Becker & Vaz Pinto & Hauptfleisch" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura otjihipa" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="otjihipa">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">Afroedura otjihipa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (holotype female, NMNW R11253) from Otjihipa Middleberg, Kunene Region, Namibia. Photos:
|
||
<emphasis bold="true" pageId="0" pageNumber="435">A-B.</emphasis>
|
||
Javier
|
||
<normalizedToken originalValue="Lobón-Rovira">Lobon-Rovira</normalizedToken>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="435">C-D</emphasis>
|
||
. Francois Becker.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="435" type="description">
|
||
<paragraph pageId="0" pageNumber="435">Holotype description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="435">
|
||
Adult female: SVL 57.9 mm; tail regenerated, with a small mid-ventral incision for the removal of liver sample. Measurements and meristic characters of holotype presented in Table
|
||
<tableCitation captionStart="Table 7" captionStartId="T7" captionText="Table 7. Measurements (in mm) and scale counts for the type series of Afroedura otjihipa sp. nov." httpUri="http://table.plazi.org/id/4AA3093F14F3331B02665C8753C139FF" pageId="0" pageNumber="435" tableUuid="4AA3093F14F3331B02665C8753C139FF">7</tableCitation>
|
||
. Head and body dorsoventrally depressed; HL 13.6 mm, HW 13.2 mm, head broadest posterior level of eye and 1.02 times longer than wide. Eyes large (3.2 mm wide), pupil vertical with indented margins; circumorbital scales small and smooth, bottom posterior scales with small upward pointing spines. Snout rounded, 5.7 mm long, longer than distance between eye and ear openings (4.8 mm). Scales on top of snout smooth, rounded, similar in size, with no intervening minute granules. Scales on snout slightly larger than those on back of head or nape. Scales on eyelids larger than those on the crown, 5 scales deep from circumorbital scale to crown. Nostril pierced between first supralabial and three nasal scales; rostral narrowly excluded from nostril; supranasals much larger than the smaller postnasals, ventral postnasal being about half the size of its dorsal counterpart, and all in broad contact with one other. Nostrils very slightly elevated. Rostral roughly rectangular, but with its upper edges elongated due to extensions toward the nostril, and the central point extends between the nasals. Seven supralabials on each side, the labial margin flexing upwards at the rictus (approx. mid-orbital position), with 1-2 elongate scales proximal to the flexure and several minute scales along the flexure proximal to these. Seven infralabials on either side. At the lip, mental scale slightly narrower than adjacent infralabial, mental only two thirds the width of rostral (1.1 mm versus 1.8 mm respectively) and in contact with three postmental scales; mental similar in size and shape to the surrounding gular scales, the central one of which is distinctly smaller. Scales on throat much smaller than those on belly, scales touching infralabials larger. Fourteen scales across the crown at level of front of eyes; 10 scales between nostril and front of eye; 12 scales from ear to eye; 67 scales around mid-body. Ear opening deep, oblique and roughly oval, less than half as high as wide (0.42
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
0.95 mm respectively). Scales on dorsum smooth, non-overlapping, largest at mid-body, smaller on nape and tail base. Scales on ventrum flattened, not overlapping, roughly twice the size of lateral granules and 1.4 times the size of scales along the dorsal mid-line. Regenerated tail dorsoventrally flattened, roughly as broad as the neck, with ventral scales larger than those on the dorsal surface. Limbs well-developed, hindlimbs slightly longer than forelimbs; all limbs without obvious mite pockets at posterior or anterior margin of limb insertions. All digits with a large pair of distal scansors, separated by a curved claw, notably smaller on the fingers than toes, and followed after a gap (about the width of terminal scansor) by a smaller pair of scansors; infero-median row of digital scales slightly enlarged transversely, the distal two rows being paired in both digits and toes, where the terminal scale adjoining the first pair of scansors may be swollen and scansor-like; 6 enlarged central and two paired distal scale rows under 3rd toe, while other toes have paired scale rows, 8 under the 4th toe.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="4AA3093F14F3331B02665C8753C139FF" httpUri="http://table.plazi.org/id/4AA3093F14F3331B02665C8753C139FF" pageId="0" pageNumber="435" start="Table 7" startId="T7">
|
||
<paragraph pageId="0" pageNumber="435">
|
||
<emphasis bold="true" pageId="0" pageNumber="435">Table 7.</emphasis>
|
||
Measurements (in mm) and scale counts for the type series of
|
||
<taxonomicName authorityName="Conradie & Schmitz & Lobón-Rovira & Becker & Vaz Pinto & Hauptfleisch" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura otjihipa" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="otjihipa">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">Afroedura otjihipa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="435">
|
||
<table pageId="0" pageNumber="435">
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="435">Catalogue Number</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="435">NMNW R11253</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="435">NMNW R11245</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Type Status</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Holotype</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Paratype</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Sex</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Female</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Male</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Snout-vent length</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">57.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">59.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Tail length</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Tail condition</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Truncated</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Regenerated</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Head length</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">13.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">15.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Head width</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">13.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">13.3</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Snout length</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">5.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">6.0</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Eye distance</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">3.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">3.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Eye-Ear distance</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">4.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">5.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Precloacal pores (males)</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">12</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Dorsal rows per tail verticil</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">5</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Ventral rows per tail verticil</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">6</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">6</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Scales below 4th toe</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">8</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Midbody scale rows</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">67</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">65</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Scales between eyes</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">14</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">14</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Scales: nostril to eye</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">10</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">11</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Scales: ear to eye</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">12</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">13</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Supranasals in contact</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Yes</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Yes</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Supralabials</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">8</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="435">
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">Infralabials</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="435" rowspan="1">8</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="435" type="paratype variation">
|
||
<paragraph pageId="0" pageNumber="435">Paratype variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="435">
|
||
SVL 59.9 mm adult male, tail truncated, precloacal pores 12. Measurements and meristic characters of paratype are presented in Table
|
||
<tableCitation captionStart="Table 7" captionStartId="T7" captionText="Table 7. Measurements (in mm) and scale counts for the type series of Afroedura otjihipa sp. nov." httpUri="http://table.plazi.org/id/4AA3093F14F3331B02665C8753C139FF" pageId="0" pageNumber="435" tableUuid="4AA3093F14F3331B02665C8753C139FF">7</tableCitation>
|
||
. The paratype is very similar to the holotype with regard to scalation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="435" type="colouration">
|
||
<paragraph pageId="0" pageNumber="435">Colouration.</paragraph>
|
||
<paragraph pageId="0" pageNumber="435">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="435">In life</emphasis>
|
||
(holotype NMNW R11253, Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Afroedura otjihipa sp. nov. (NMNW R 11253) from Otjihipa Middleberg, Kunene Region, Namibia. Scale bar: 1 cm. Photos: Francois Becker." figureDoi="10.3897/zse.98.86299.figure7" httpUri="https://binary.pensoft.net/fig/772235" pageId="0" pageNumber="435">7C-D</figureCitation>
|
||
): dark brown with yellowish patterns, fading to whitish on limbs and top of head; yellow patterns are irregular, asymmetrical patches and spots along the body, symmetrical paired spots around the nape and near the tail base; there is a thin, irregular, broken or continuous yellow bar on the nape; another broken, irregular yellow bar across the scapular region to the shoulders; three asymmetrical yellow double-bars which may present as pairs of medially-angled triangles posteriorly, across the back, each with an irregular dark brown core; another broken yellow bar or collection of symmetrical spots around the sacrum; head dark brown with yellow blotches on the crown with intervening pale yellow colouration; dark brown bar from nostril across the upper margins of the ear opening, connecting with dark brown lateral bar on the neck; a thin pale yellow canthal stripe extends on both sides from the nasal region to anterior margins of eye, continuing posteriorly from the eye onto the nape; skin above eyes copper blue with dark brown spots; upper and lower labials light grey with dense brown speckling, denser anteriorly and on supralabials; lateral sides of the body with a mix of dark brown and yellowish blotches, as a continuation of the dorsal patterns; limbs dark brown above with scattered light grey markings; tail (regenerated) with an asymmetrical chequered pattern of dark brown and light grey; iris copper in colour with a narrow black elliptic pupil with crenulated edge, and black reticulation; venter uniform beige with scattered brown specks mostly on lateral edges; ventrally, limbs with scattered brown spots, mostly near lateral surfaces. In preservative: yellowish patterns faded to light grey, dark brown to grey-brown, and eyes faded to bluish grey, with original colouration of pupils and iris no longer evident.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="435">Paratype colouration</emphasis>
|
||
: Similar colouration and patterning as to the holotype, but the yellow bands and patterns are more clearly defined: the bar on the nape is nearly continuous, that on the scapular region has a clear dark brown core, and three pairs of asymmetrical, medially-pointing, dark brown, triangular blobs are clearly outlined by irregular yellow lines; no clear bar near the tail base, but a collection of symmetrical spots. The original tails are not present on the preserved specimen, but were observed briefly in life before capture. The original tails of another pair of individuals in a nearby rock crack (not caught) were also observed. Tail bars could not be counted, but bold black-and-white banding was clearly visible.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.98.86299.figure7" httpUri="https://binary.pensoft.net/fig/772235" pageId="0" pageNumber="435" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="435">
|
||
<emphasis bold="true" pageId="0" pageNumber="435">Figure 7.</emphasis>
|
||
Holotype of
|
||
<taxonomicName authorityName="Conradie & Schmitz & Lobón-Rovira & Becker & Vaz Pinto & Hauptfleisch" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura otjihipa" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="otjihipa">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">Afroedura otjihipa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (NMNW R11253) from Otjihipa Middleberg, Kunene Region, Namibia. Scale bar: 1 cm. Photos: Francois Becker.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="435" type="natural history">
|
||
<paragraph pageId="0" pageNumber="435">Natural history and habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="435">
|
||
A rupicolous species living in narrow rock crevices in relatively small sandstone outcrops in arid woodland savannah (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitat photos of: A-B. Afroedura pundomontana sp. nov.: Morro do Pundo, 25 km west of Bocoio, Benguela Province, Angola; C-D. Afroedura otjihipa sp. nov.: Otjihipa Middleberg, Kunene Region, Namibia. Photos: A-B. Pedro Vaz Pinto; C-D. Francois Becker." figureDoi="10.3897/zse.98.86299.figure5" httpUri="https://binary.pensoft.net/fig/772233" pageId="0" pageNumber="435">5C-D</figureCitation>
|
||
), at elevations of 1,800-1,900 m a.s.l. in the Otjihipa Mountains. It was not found in the dolomite formations near the type locality, despite greater search time dedicated to those areas. The rock cracks where they were found were smaller than is typical for this group and were similar throughout this surface formation. Congeners in the
|
||
<taxonomicName lsidName="A. bogerti" pageId="0" pageNumber="435" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
group are normally found only in deep rock cracks in and amongst large boulders. Such habitat features were present in the surveyed area, but only in dolomite formations. The much less crevice-rich sandstone formation, with thin, straight cracks formed between the sandstone strata, appeared to be favoured syntopically by
|
||
<taxonomicName lsidName="A. otjihipa" pageId="0" pageNumber="435" rank="species" species="otjihipa">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">A. otjihipa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" lsidName="Cordylus namakuiyus" order="Squamata" pageId="0" pageNumber="435" phylum="Chordata" rank="species" species="namakuiyus">
|
||
<emphasis italics="true" pageId="0" pageNumber="435">Cordylus namakuiyus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="435" type="distribution">
|
||
<paragraph pageId="0" pageNumber="435">Distribution and conservation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="435">
|
||
Currently known from a single sandstone ridge on Otjihipa Middleberg in the extreme north-west of the Kunene Region, Namibia (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic tree topology based on the combined mitochondrial (12 S, 16 S, Cyt-b, ND 2) and nuclear (c-mos, RAG 1) genes, using Afroedura karroica as outgroup. Support values for Bayesian posterior probabilities (above nodes) and maximum likelihood bootstraps (below nodes) are indicated in the tree (shown values: ML: ≥ 70 % / PP: ≥ 0.75)." figureDoi="10.3897/zse.98.86299.figure2" httpUri="https://binary.pensoft.net/fig/772230" pageId="0" pageNumber="435">2</figureCitation>
|
||
). The species remains poorly known, but it is probably stable in numbers as the local habitat is currently not threatened and is topographically unsuitable for human habitation. It likely occurs more broadly across the Otjihipa Mountain range. In accordance with IUCN Red List Guidelines (
|
||
<bibRefCitation author="Herrman, H-W" journalOrPublisher="Journal of Arid Environments" pageId="0" pageNumber="435" publicationUrl="https://www.iucnredlist.org" refId="B23" refString="2022. . https://www.iucnredlist.org" url="https://www.iucnredlist.org" year="2022">IUCN 2022</bibRefCitation>
|
||
) we propose this species to be classified as Data Deficient (DD) at this stage, but due to the remoteness of the locality and because no notifiable threats exist, it could be listed as Least Concern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |