147 lines
13 KiB
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147 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.929.49987" ID-GBIF-Dataset="97c79ed9-33fd-49b8-90c5-95a159839db2" ID-PMC="PMC7192954" ID-Pensoft-Pub="1313-2970-929-1" ID-Pensoft-UUID="EFB92B457B1D56A1996483C5EECFAAF0" ID-PubMed="32377146" ID-ZooBank="4FCA3A27–973C49F7–8264–6A34D279EC5C" ModsDocID="1313-2970-929-1" checkinTime="1587627147117" checkinUser="pensoft" docAuthor="Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki & Bjoern Stelbrink," docDate="2020" docId="17AF80062CC55BF8B55476DE58D31D70" docLanguage="en" docName="ZooKeys 929: 1-17" docOrigin="ZooKeys 929" docSource="http://dx.doi.org/10.3897/zookeys.929.49987" docTitle="Erhaia banepaensis Nesemann & S. Sharma 2007" docType="treatment" docVersion="4" id="EFB92B457B1D56A1996483C5EECFAAF0" lastPageNumber="8" masterDocId="EFB92B457B1D56A1996483C5EECFAAF0" masterDocTitle="The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya" masterLastPageNumber="17" masterPageNumber="1" pageNumber="8" updateTime="1668168753491" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gittenberger, Edmund</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, NL- 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:nameIdentifier type="email">egittenberger@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Leda, Pema</mods:namePart>
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<mods:affiliation>National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wangchuk, Jigme</mods:namePart>
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<mods:affiliation>Ugyen Wangchuck Institute for Conservation and Environmental Research, Bumthang, Bhutan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-3503-4369</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gyeltshen, Choki</mods:namePart>
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<mods:affiliation>National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6790-6278</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bjoern Stelbrink,</mods:namePart>
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<mods:affiliation>Department of Animal Ecology & Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany & Zoological Institute, Department of Environmental Sciences, University of Basel, Vesalgasse 1, 4051 Basel, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7471-4992</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>929</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>17</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.929.49987</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.929.49987</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-929-1</mods:identifier>
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<mods:identifier type="ZooBank">4FCA3A27-973C49F7-8264-6A34D279EC5C</mods:identifier>
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<treatment ID-GBIF-Taxon="163735481" LSID="urn:lsid:plazi:treatment:17AF80062CC55BF8B55476DE58D31D70" httpUri="http://treatment.plazi.org/id/17AF80062CC55BF8B55476DE58D31D70" lastPageNumber="8" pageId="7" pageNumber="8">
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<subSubSection pageId="7" pageNumber="8" type="nomenclature">
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<paragraph pageId="7" pageNumber="8">
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<taxonomicName LSID="17AF8006-2CC5-5BF8-B554-76DE58D31D70" authority="Nesemann & S. Sharma, 2007" authorityName="Nesemann & S. Sharma" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia banepaensis" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="banepaensis">Erhaia banepaensis Nesemann & S. Sharma, 2007</taxonomicName>
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<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="7" pageNumber="8">Fig. 8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="reference_group">
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<paragraph pageId="7" pageNumber="8">
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<taxonomicName authorityName="Nesemann & S. Sharma" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia banepaensis" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="banepaensis">Erhaia banepaensis</taxonomicName>
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Nesemann & S. Sharma, in
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<bibRefCitation author="Nesemann, H" journalOrPublisher="Molecular Biology and Evolution" pageId="14" pageNumber="15" refId="B13" refString="Nesemann, H, Sharma, S, Sharma, G, Khanal, SN, Pradhan, B, Shah, DN, Tachamo, RD, 2007. Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57-102." title="Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57 - 102." year="2007">Nesemann et al. 2007</bibRefCitation>
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: 64 ("Nepal, Central Zone, Kavre District, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa"; the altitude is not indicated).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="shell">
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<paragraph pageId="7" pageNumber="8">Shell.</paragraph>
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<paragraph pageId="7" pageNumber="8">
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The shells are described by Nesemann and Sharma (2007: 64) as 1.6-2.0 mm high, with 4-
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<normalizedToken originalValue="4½">41/2</normalizedToken>
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<normalizedToken originalValue="“convex”">"convex"</normalizedToken>
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whorls, an aperture that is "ovate but not widened and not enlarged", with an inner lip that is "thin and fused to the body whorl"; it can be distinguished from the other Nepalese
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<taxonomicName authorityName="Davis & Kuo" authorityYear="1985" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="7" pageNumber="8">Erhaia</emphasis>
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</taxonomicName>
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species by the "conical and compact shape" and
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<normalizedToken originalValue="“convex”">"convex"</normalizedToken>
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whorls (2007: 65).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="materials_examined">
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<paragraph pageId="7" pageNumber="8">Material examined</paragraph>
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<paragraph pageId="7" pageNumber="8">
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(photo only).
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<emphasis bold="true" pageId="7" pageNumber="8">
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<emphasis italics="true" pageId="7" pageNumber="8">Holotype</emphasis>
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.
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</emphasis>
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(NHMW 1033159).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="notes">
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<paragraph pageId="7" pageNumber="8">Notes.</paragraph>
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<paragraph pageId="7" pageNumber="8">
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Nesemann and S. Sharma are mentioned as authors for this species, without specifying for what part of the text in
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<bibRefCitation author="Nesemann, H" journalOrPublisher="Molecular Biology and Evolution" pageId="14" pageNumber="15" refId="B13" refString="Nesemann, H, Sharma, S, Sharma, G, Khanal, SN, Pradhan, B, Shah, DN, Tachamo, RD, 2007. Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57-102." title="Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57 - 102." year="2007">Nesemann et al (2007)</bibRefCitation>
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they have responsibility.
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</paragraph>
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<paragraph pageId="7" pageNumber="8">
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According to
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<bibRefCitation author="Nesemann, H" journalOrPublisher="Molecular Biology and Evolution" pageId="14" pageNumber="15" refId="B13" refString="Nesemann, H, Sharma, S, Sharma, G, Khanal, SN, Pradhan, B, Shah, DN, Tachamo, RD, 2007. Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57-102." title="Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57 - 102." year="2007">Nesemann et al. (2007</bibRefCitation>
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: 64)
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<taxonomicName lsidName="E. banepaensis" pageId="7" pageNumber="8" rank="species" species="banepaensis">
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<emphasis italics="true" pageId="7" pageNumber="8">E. banepaensis</emphasis>
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</taxonomicName>
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occurs sympatrically with
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<taxonomicName lsidName="E. chandeshwariensis" pageId="7" pageNumber="8" rank="species" species="chandeshwariensis">
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<emphasis italics="true" pageId="7" pageNumber="8">E. chandeshwariensis</emphasis>
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</taxonomicName>
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at the shared type locality of these species.
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<taxonomicName authorityName="Nesemann & S. Sharma" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia banepaensis" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="banepaensis">
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<emphasis italics="true" pageId="7" pageNumber="8">Erhaia banepaensis</emphasis>
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</taxonomicName>
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is supposed to be more widely distributed in Nepal between 1400 and 2085 m a.s.l., but the type series is restricted to the holotype (NHMW 103319 [not 1033159]) and a paratype (NHMW 103320). The location of the additional material is not indicated.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/402720" pageId="7" pageNumber="8" start="Figure 13" startId="F5">
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<paragraph pageId="7" pageNumber="8">
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<emphasis bold="true" pageId="7" pageNumber="8">Figure 13.</emphasis>
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Habitat of
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<taxonomicName baseAuthorityName="Benson" baseAuthorityYear="1843" class="Gastropoda" family="Pomatiopsidae" genus="Tricula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tricula montana" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="montana">
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<emphasis italics="true" pageId="7" pageNumber="8">Tricula montana</emphasis>
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</taxonomicName>
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; Bhutan, district Mongar, Chhuburee, 818 m a.s.l. Photo by JW.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |