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<document id="B7525964DEE06CD73ED74F41DAF00671" ID-CLB-Dataset="2511" ID-DOI="10.5281/zenodo.5339720" ID-GBIF-Dataset="b94435d4-ba95-49a5-9656-fdaaf1eb8ae3" ID-ISSN="0374-1036" ID-Zenodo-Dep="5339720" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630184679885" checkinUser="felipe" docAuthor="Bezděk, Jan" docDate="2012" docId="17698A63FFF1E86AFE36E2D1FE30A052" docLanguage="en" docName="ActaEntMusNatPra.2012.52.suppl.403-428.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Beenenia Bezděk 2012, gen. nov." docType="treatment" docVersion="8" lastPageNumber="406" masterDocId="EB50F21BFFF0E869FFC6E611FFEEA461" masterDocTitle="Galerucinae (Coleoptera: Chrysomelidae) of Socotra Island, with a review of taxa recorded from Yemen" masterLastPageNumber="428" masterPageNumber="403" pageNumber="404" updateTime="1699196791282" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F73D34DF10A0BC1E6CCBA602D96F6CE1">Galerucinae (Coleoptera: Chrysomelidae) of Socotra Island, with a review of taxa recorded from Yemen</mods:title>
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<mods:name id="6E1AA34EA8DF8231CEAC30C2906175DA" type="personal">
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<mods:namePart id="F3895AE922B94DAC1B077475148FBF0E">Bezděk, Jan</mods:namePart>
<mods:affiliation id="9F00082A7B9C7FB5C2936EE11808961B">Mendel University, Department of Zoology, Zemědělská 1, CZ 613 00 Brno, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="460547BDBA2CABD91E1D9C62348A20D6" type="email">bezdek@mendelu.cz</mods:nameIdentifier>
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<mods:title id="E4005008B53FBC2A6A0D7F77E52D837C">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="3BFC8A1DD99A56B96DDB6BD5BA0332D7">2012</mods:date>
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<mods:number id="418AB074A40DF35BC599F0D099199248">2012-12-17</mods:number>
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<treatment id="17698A63FFF1E86AFE36E2D1FE30A052" ID-DOI="http://doi.org/10.5281/zenodo.5415750" ID-GBIF-Taxon="192438073" ID-Zenodo-Dep="5415750" LSID="urn:lsid:plazi:treatment:17698A63FFF1E86AFE36E2D1FE30A052" httpUri="http://treatment.plazi.org/id/17698A63FFF1E86AFE36E2D1FE30A052" lastPageId="3" lastPageNumber="406" pageId="1" pageNumber="404">
<subSubSection id="D7DA68FEFFF1E868FE36E2D1FD2CA0B9" box="[496,706,1216,1240]" pageId="1" pageNumber="404" type="nomenclature">
<paragraph id="9F7F3B75FFF1E868FE36E2D1FD2CA0B9" blockId="1.[496,706,1216,1240]" box="[496,706,1216,1240]" pageId="1" pageNumber="404">
<heading id="C4378C19FFF1E868FE36E2D1FD2CA0B9" bold="true" box="[496,706,1216,1240]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="404" reason="3">
<emphasis id="ADB4E767FFF1E868FE36E2D1FD2CA0B9" bold="true" box="[496,706,1216,1240]" pageId="1" pageNumber="404">
<taxonomicName id="58C040F6FFF1E868FE36E2D1FDB6A0B9" authority="Bezděk, 2012" authorityName="Bezděk" authorityYear="2012" box="[496,600,1216,1240]" class="Insecta" family="Chrysomelidae" genus="Beenenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="404" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="ADB4E767FFF1E868FE36E2D1FDB6A0B9" bold="true" box="[496,600,1216,1240]" italics="true" pageId="1" pageNumber="404">Beenenia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6875A1CFFF1E868FD98E2D1FD2CA0B9" box="[606,706,1216,1240]" pageId="1" pageNumber="404" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D7DA68FEFFF1E868FFB7E2FEFCAFA166" box="[113,833,1263,1288]" pageId="1" pageNumber="404" type="type_taxon">
<paragraph id="9F7F3B75FFF1E868FFB7E2FEFCAFA166" blockId="1.[113,833,1263,1288]" box="[113,833,1263,1288]" pageId="1" pageNumber="404">
<emphasis id="ADB4E767FFF1E868FFB7E2FEFEEBA166" bold="true" box="[113,261,1263,1287]" pageId="1" pageNumber="404">
<typeStatus id="407B85D7FFF1E868FFB7E2FEFF47A166" box="[113,169,1263,1287]" pageId="1" pageNumber="404">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="58C040F6FFF1E868FECAE2E0FE0AA169" authority="Bezděk, 2012" authorityName="Bezděk" authorityYear="2012" box="[268,484,1265,1288]" class="Insecta" family="Chrysomelidae" genus="Beenenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="scanticola" status="sp. nov.">
<emphasis id="ADB4E767FFF1E868FECAE2E0FE0AA169" box="[268,484,1265,1288]" italics="true" pageId="1" pageNumber="404">Beenenia scanticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6875A1CFFF1E868FE2CE2E1FDD1A166" box="[490,575,1264,1287]" pageId="1" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="D7DA68FEFFF1E86AFFB7E30FFF3EA652" lastPageId="3" lastPageNumber="406" pageId="1" pageNumber="404" type="description">
<paragraph id="9F7F3B75FFF1E868FFB7E30FFBD0A117" blockId="1.[113,1089,1310,1622]" pageId="1" pageNumber="404">
<emphasis id="ADB4E767FFF1E868FFB7E30FFF15A157" bold="true" box="[113,251,1310,1334]" pageId="1" pageNumber="404">Description.</emphasis>
Body elongate, dorsum metallic dark greenish to black or brownish with distinct metallic tint, antennae black, legs black with metallic tint, with pale coxae and trochanters (in
<taxonomicName id="58C040F6FFF1E868FFB7E371FF07A116" authority="Bezděk, 2012" authorityName="Bezděk" authorityYear="2012" box="[113,233,1376,1399]" class="Insecta" family="Chrysomelidae" genus="Beenenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="kabateki" status="sp. nov.">
<emphasis id="ADB4E767FFF1E868FFB7E371FF07A116" box="[113,233,1376,1399]" italics="true" pageId="1" pageNumber="404">B. kabateki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6875A1CFFF1E868FF36E34EFEADA117" box="[240,323,1375,1398]" pageId="1" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
also with pale knees), dorsum densely pubescent, semiopaque to shiny.
</paragraph>
<paragraph id="9F7F3B75FFF1E86BFF56E36FFBA8A0D7" blockId="1.[113,1089,1310,1622]" lastBlockId="2.[151,1127,1118,1622]" lastPageId="2" lastPageNumber="405" pageId="1" pageNumber="404">Head large, measured with eyes slightly narrower or as wide as anterior margin of pronotum, eyes small, moderately projecting beyond head outline. Labrum transverse, with anterior margin shallowly incised, covered with several long setae. Anterior part of head very short, genae 0.30 times as long as diameter of eye. Nasal keel poorly developed but visible. Frontal tubercles large, convex, distinctly elevated, impunctate, with apices elongate, separated from each other by thin sulcus which continues through vertex. Frons shallowly depressed behind frontal tubercles but not grooved. Vertex large, moderately convex, densely covered with large punctures and setae. Antennae relatively robust, short, reaching slightly behind midlenght of body, antennomeres IIVI flatenned and slightly impressed in basal part, apically extended, antennomeres VIIXI robust, gradually less flattened, last two antennomeres subtubular.</paragraph>
<caption id="CBBF6BFDFFF2E86BFF51E5C5FEF4A063" ID-DOI="http://doi.org/10.5281/zenodo.5339722" ID-Zenodo-Dep="5339722" httpUri="https://zenodo.org/record/5339722/files/figure.png" pageId="2" pageNumber="405" startId="2.[151,190,980,999]" targetBox="[236,1057,235,962]" targetPageId="2">
<paragraph id="9F7F3B75FFF2E86BFF51E5C5FEF4A063" blockId="2.[151,1127,980,1026]" pageId="2" pageNumber="405">
Figs. 12. Habitus. 1
<taxonomicName id="58C040F6FFF2E86BFEA6E5C4FDE3A786" authority="Bezděk, 2012" authorityName="Bezděk" authorityYear="2012" box="[352,525,981,999]" class="Insecta" family="Chrysomelidae" genus="Beenenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="scanticola" status="sp. nov.">
<emphasis id="ADB4E767FFF2E86BFEA6E5C4FDE3A786" box="[352,525,981,999]" italics="true" pageId="2" pageNumber="405">Beenenia scanticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6875A1CFFF2E86BFDD5E5C4FDB8A789" box="[531,598,981,1000]" pageId="2" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
(holotype, male, 6.0 mm); 2
<taxonomicName id="58C040F6FFF2E86BFCA7E5C4FC2FA786" authority="Bezděk, 2012" authorityName="Bezděk" authorityYear="2012" box="[865,961,981,999]" class="Insecta" family="Chrysomelidae" genus="Beenenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="405" phylum="Arthropoda" rank="species" species="kabateki" status="sp. nov.">
<emphasis id="ADB4E767FFF2E86BFCA7E5C4FC2FA786" box="[865,961,981,999]" italics="true" pageId="2" pageNumber="405">B. kabateki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6875A1CFFF2E86BFC01E5C4FBE5A789" box="[967,1035,981,1000]" pageId="2" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
(holotype, male, 5.4 mm).
</paragraph>
</caption>
<paragraph id="9F7F3B75FFF2E86BFF73E2AFFCD9A1D4" blockId="2.[151,1127,1118,1622]" pageId="2" pageNumber="405">Pronotum transverse, 1.752.05 times as wide as long, narrower than base of elytra. Anterior margin thinly bordered only at sides or unbordered, lateral margin deeply and thinly bordered, posterior margin shallowly and thinly bordered. Surface of pronotum moderately convex, with shallow depressions laterally and in middle of both anterior and posterior part of pronotum, densely covered with fine to large punctures. All angles prominent, triangular, projecting laterally, anterior angles bearing 4 to 6 long pale setae, posterior angles with two or three long pale setae. Scutellum subtriangular, with widely rounded tip, subopaque, covered with microsculpture or indistinct fine punctures, and fine setae.</paragraph>
<paragraph id="9F7F3B75FFF2E86BFF73E3AFFD95A234" blockId="2.[151,1127,1118,1622]" pageId="2" pageNumber="405">
Elytra with well developed humeral calli. Surface very densely covered with small confused punctures and with one or
<specimenCount id="89C6F0FCFFF2E86BFE73E3CFFDF2A197" box="[437,540,1502,1526]" pageId="2" pageNumber="405" type="generic" typeStatus="types">two types</specimenCount>
of setae. Interspaces between punctures narrower than diameter of each puncture. Postscutellar area slightly depressed. Epipleura not developed. Basal part of elytral lateral slope inflexed down inwards, towards apex more or less vertical. Humeral calli well developed. Macropterous.
</paragraph>
<paragraph id="9F7F3B75FFF3E86AFF56E6CDFCA9A535" blockId="3.[113,1089,219,1075]" pageId="3" pageNumber="406">
Legs moderately long and narrow, middle and posterior tibiae unarmed at apex. Tibiae even, without longitudinal ridges or furrows. Metatarsomere I as long as tarsomeres IIIII combined. Anterior coxal cavities open posteriorly, prosternum very thinly visible between coxae. Claws bifid with inner branch shorter than outer one (
<figureCitation id="07FB27F0FFF3E86AFD3EE72AFCD7A535" box="[760,825,315,340]" captionStart="Figs" captionStartId="4.[151,190,1535,1554]" captionTargetBox="[179,1095,226,1474]" captionTargetId="figure-12@4.[151,1125,219,1478]" captionTargetPageId="4" captionText="Figs. 38. Beenenia scanticola sp. nov. 3 aedeagus (a dorsal view, b lateral view); 4 last male ventrite; 5 sternite VIII and tignum; 6 vaginal palpi; 7 spermatheca; 8 claws. Scale bars: 1 mm for Figs. 35, 0.5 mm for Figs. 67, 0.25 mm for Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.5339724" httpUri="https://zenodo.org/record/5339724/files/figure.png" pageId="3" pageNumber="406">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="9F7F3B75FFF3E86AFF56E74AFC69A5B5" blockId="3.[113,1089,219,1075]" pageId="3" pageNumber="406">
Basal two third of aedeagus robust and more or less tubular, apical third slightly asymmetrical, flat and bent downwards, apex widely rounded with distinct small process (
<figureCitation id="07FB27F0FFF3E86AFC37E76AFBD2A5F5" box="[1009,1084,379,404]" captionStart="Figs" captionStartId="4.[151,190,1535,1554]" captionTargetBox="[179,1095,226,1474]" captionTargetId="figure-12@4.[151,1125,219,1478]" captionTargetPageId="4" captionText="Figs. 38. Beenenia scanticola sp. nov. 3 aedeagus (a dorsal view, b lateral view); 4 last male ventrite; 5 sternite VIII and tignum; 6 vaginal palpi; 7 spermatheca; 8 claws. Scale bars: 1 mm for Figs. 35, 0.5 mm for Figs. 67, 0.25 mm for Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.5339724" httpUri="https://zenodo.org/record/5339724/files/figure.png" pageId="3" pageNumber="406">Figs. 3</figureCitation>
,
<figureCitation id="07FB27F0FFF3E86AFFB7E78DFF6FA5D5" box="[113,129,412,436]" captionStart="Figs" captionStartId="7.[113,152,1546,1565]" captionTargetBox="[128,1076,222,1442]" captionTargetId="figure-12@7.[113,1087,219,1449]" captionTargetPageId="7" captionText="Figs. 912. Beenenia kabateki sp. nov.9 aedeagus (a dorsal view, b lateral view); 10 sternite VIII and tignum; 11 vaginal palpi; 12 spermatheca. Scale bars: 1 mm for Figs. 910, 0.5 mm for Figs. 1112." figureDoi="http://doi.org/10.5281/zenodo.5339726" httpUri="https://zenodo.org/record/5339726/files/figure.png" pageId="3" pageNumber="406">9</figureCitation>
). Spermatheca with C-shaped cornu, nodulus small, narrower than base of cornu, ductus receptaculi short (
<figureCitation id="07FB27F0FFF3E86AFEF7E7AAFE92A5B2" box="[305,380,443,467]" captionStart="Figs" captionStartId="4.[151,190,1535,1554]" captionTargetBox="[179,1095,226,1474]" captionTargetId="figure-12@4.[151,1125,219,1478]" captionTargetPageId="4" captionText="Figs. 38. Beenenia scanticola sp. nov. 3 aedeagus (a dorsal view, b lateral view); 4 last male ventrite; 5 sternite VIII and tignum; 6 vaginal palpi; 7 spermatheca; 8 claws. Scale bars: 1 mm for Figs. 35, 0.5 mm for Figs. 67, 0.25 mm for Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.5339724" httpUri="https://zenodo.org/record/5339724/files/figure.png" pageId="3" pageNumber="406">Figs. 7</figureCitation>
,
<figureCitation id="07FB27F0FFF3E86AFE4EE7ADFE48A5B5" box="[392,422,444,468]" captionStart="Figs" captionStartId="7.[113,152,1546,1565]" captionTargetBox="[128,1076,222,1442]" captionTargetId="figure-12@7.[113,1087,219,1449]" captionTargetPageId="7" captionText="Figs. 912. Beenenia kabateki sp. nov.9 aedeagus (a dorsal view, b lateral view); 10 sternite VIII and tignum; 11 vaginal palpi; 12 spermatheca. Scale bars: 1 mm for Figs. 910, 0.5 mm for Figs. 1112." figureDoi="http://doi.org/10.5281/zenodo.5339726" httpUri="https://zenodo.org/record/5339726/files/figure.png" pageId="3" pageNumber="406">12</figureCitation>
). Vaginal palpi short and robust (
<figureCitation id="07FB27F0FFF3E86AFCC1E7AAFCBCA5B2" box="[775,850,443,467]" captionStart="Figs" captionStartId="4.[151,190,1535,1554]" captionTargetBox="[179,1095,226,1474]" captionTargetId="figure-12@4.[151,1125,219,1478]" captionTargetPageId="4" captionText="Figs. 38. Beenenia scanticola sp. nov. 3 aedeagus (a dorsal view, b lateral view); 4 last male ventrite; 5 sternite VIII and tignum; 6 vaginal palpi; 7 spermatheca; 8 claws. Scale bars: 1 mm for Figs. 35, 0.5 mm for Figs. 67, 0.25 mm for Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.5339724" httpUri="https://zenodo.org/record/5339724/files/figure.png" pageId="3" pageNumber="406">Figs. 6</figureCitation>
,
<figureCitation id="07FB27F0FFF3E86AFC98E7ADFC94A5B5" box="[862,890,444,468]" captionStart="Figs" captionStartId="7.[113,152,1546,1565]" captionTargetBox="[128,1076,222,1442]" captionTargetId="figure-12@7.[113,1087,219,1449]" captionTargetPageId="7" captionText="Figs. 912. Beenenia kabateki sp. nov.9 aedeagus (a dorsal view, b lateral view); 10 sternite VIII and tignum; 11 vaginal palpi; 12 spermatheca. Scale bars: 1 mm for Figs. 910, 0.5 mm for Figs. 1112." figureDoi="http://doi.org/10.5281/zenodo.5339726" httpUri="https://zenodo.org/record/5339726/files/figure.png" pageId="3" pageNumber="406">11</figureCitation>
).
</paragraph>
<paragraph id="9F7F3B75FFF3E86AFF56E7CAFF3EA652" blockId="3.[113,1089,219,1075]" pageId="3" pageNumber="406">
Sexual dimorphism. Weakly indicated. Males on average smaller. Last ventrite in male with posterior margin widely triangularly impressed but not incised (
<figureCitation id="07FB27F0FFF3E86AFC86E7EAFC6EA675" box="[832,896,507,532]" captionStart="Figs" captionStartId="4.[151,190,1535,1554]" captionTargetBox="[179,1095,226,1474]" captionTargetId="figure-12@4.[151,1125,219,1478]" captionTargetPageId="4" captionText="Figs. 38. Beenenia scanticola sp. nov. 3 aedeagus (a dorsal view, b lateral view); 4 last male ventrite; 5 sternite VIII and tignum; 6 vaginal palpi; 7 spermatheca; 8 claws. Scale bars: 1 mm for Figs. 35, 0.5 mm for Figs. 67, 0.25 mm for Fig. 8." figureDoi="http://doi.org/10.5281/zenodo.5339724" httpUri="https://zenodo.org/record/5339724/files/figure.png" pageId="3" pageNumber="406">Fig. 4</figureCitation>
), in female evenly rounded.
</paragraph>
</subSubSection>
<subSubSection id="D7DA68FEFFF3E86AFFB7E42AFC5DA795" pageId="3" pageNumber="406" type="diagnosis">
<paragraph id="9F7F3B75FFF3E86AFFB7E42AFE90A772" blockId="3.[113,1089,219,1075]" pageId="3" pageNumber="406">
<emphasis id="ADB4E767FFF3E86AFFB7E42AFE85A632" bold="true" box="[113,363,571,595]" pageId="3" pageNumber="406">Differential diagnosis.</emphasis>
<taxonomicName id="58C040F6FFF3E86AFEB7E42CFE3AA635" authority="Bezděk, 2012" authorityName="Bezděk" authorityYear="2012" box="[369,468,573,596]" class="Insecta" family="Chrysomelidae" genus="Beenenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="ADB4E767FFF3E86AFEB7E42CFE3AA635" box="[369,468,573,596]" italics="true" pageId="3" pageNumber="406">Beenenia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6875A1CFFF3E86AFE1DE42DFDD2A632" box="[475,572,572,596]" pageId="3" pageNumber="406" rank="genus">gen. nov.</taxonomicNameLabel>
belongs to the tribe
<taxonomicName id="58C040F6FFF3E86AFCD3E42AFC67A632" authorityName="Latreille" authorityYear="1802" box="[789,905,571,595]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="tribe" tribe="Galerucini">Galerucini</taxonomicName>
. Because of even tibiae without any longitudinal ridges or furrows,
<taxonomicName id="58C040F6FFF3E86AFD4EE44CFD05A615" authority="Bezděk, 2012" authorityName="Bezděk" authorityYear="2012" box="[648,747,605,628]" class="Insecta" family="Chrysomelidae" genus="Beenenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="ADB4E767FFF3E86AFD4EE44CFD05A615" box="[648,747,605,628]" italics="true" pageId="3" pageNumber="406">Beenenia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6875A1CFFF3E86AFD34E44DFCBAA612" box="[754,852,604,628]" pageId="3" pageNumber="406" rank="genus">gen. nov.</taxonomicNameLabel>
has to be classified in the section Atysites (
<bibRefCitation id="FB514684FFF3E86AFE88E46DFE0DA6F2" author="CHAPUIS F." box="[334,483,635,660]" pageId="3" pageNumber="406" refId="ref13606" refString="CHAPUIS F. 1875: Famille des Phytophages. In: LACORDAIRE T. &amp; CHAPUIS F. (eds.): Histoire naturelle des insectes. Genera des coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome onzieme. Roret, Paris, 420 pp." type="book" year="1875">CHAPUIS 1875</bibRefCitation>
,
<bibRefCitation id="FB514684FFF3E86AFE2BE46DFD0FA6F5" author="SEENO T. N. &amp; WILCOX J. A." box="[493,737,636,660]" pageId="3" pageNumber="406" pagination="1 - 221" refId="ref14153" refString="SEENO T. N. &amp; WILCOX J. A. 1982: Leaf beetle genera (Coleoptera: Chrysomelidae). Entomography 1: 1 - 221." type="journal article" year="1982">SEENO &amp; WILCOX 1982</bibRefCitation>
). From all genera of the Atysites,
<taxonomicName id="58C040F6FFF3E86AFFB7E48CFF3AA6D5" authority="Bezděk, 2012" authorityName="Bezděk" authorityYear="2012" box="[113,212,669,692]" class="Insecta" family="Chrysomelidae" genus="Beenenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="ADB4E767FFF3E86AFFB7E48CFF3AA6D5" box="[113,212,669,692]" italics="true" pageId="3" pageNumber="406">Beenenia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6875A1CFFF3E86AFF1BE48DFEAEA6D2" box="[221,320,668,692]" pageId="3" pageNumber="406" rank="genus">gen. nov.</taxonomicNameLabel>
can be distinguished by the complete absence of epipleura. Externally similar genera
<taxonomicName id="58C040F6FFF3E86AFECAE4ACFDF5A6B5" authority="Jacoby, 1900" authorityName="Jacoby" authorityYear="1900" box="[268,539,699,724]" class="Insecta" family="Chrysomelidae" genus="Luperocella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="ADB4E767FFF3E86AFECAE4ACFE65A6B5" box="[268,395,701,724]" italics="true" pageId="3" pageNumber="406">Luperocella</emphasis>
Jacoby, 1900
</taxonomicName>
;
<taxonomicName id="58C040F6FFF3E86AFDE3E4ACFCDAA6B5" authority="Crotch, 1873" authorityName="Crotch" authorityYear="1873" box="[549,820,699,724]" class="Insecta" family="Chrysomelidae" genus="Galerucella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="ADB4E767FFF3E86AFDE3E4ACFD4AA6B5" box="[549,676,701,724]" italics="true" pageId="3" pageNumber="406">Galerucella</emphasis>
Crotch, 1873
</taxonomicName>
;
<taxonomicName id="58C040F6FFF3E86AFCFBE4ACFBD2A6B2" authority="Joannis, 1865" authorityName="Joannis" authorityYear="1865" box="[829,1084,699,724]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="ADB4E767FFF3E86AFCFBE4ACFC4AA6B5" box="[829,932,701,724]" italics="true" pageId="3" pageNumber="406">Pyrrhalta</emphasis>
Joannis, 1865
</taxonomicName>
;
<taxonomicName id="58C040F6FFF3E86AFFB7E4CCFE32A695" authority="Laboissiere, 1934" authorityName="Laboissiere" authorityYear="1934" box="[113,476,731,756]" class="Insecta" family="Chrysomelidae" genus="Xanthogaleruca" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="ADB4E767FFF3E86AFFB7E4CCFEF2A695" box="[113,284,733,756]" italics="true" pageId="3" pageNumber="406">Xanthogaleruca</emphasis>
Laboissière, 1934
</taxonomicName>
and
<taxonomicName id="58C040F6FFF3E86AFDCBE4CCFCE5A695" authority="Weise, 1902" authorityName="Weise" authorityYear="1902" box="[525,779,731,756]" class="Insecta" family="Chrysomelidae" genus="Tarachodia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="ADB4E767FFF3E86AFDCBE4CCFD69A695" box="[525,647,733,756]" italics="true" pageId="3" pageNumber="406">Tarachodia</emphasis>
Weise, 1902
</taxonomicName>
can be separated also by thin, almost filiform antennae.
</paragraph>
<paragraph id="9F7F3B75FFF3E86AFF56E50DFC5DA795" blockId="3.[113,1089,219,1075]" pageId="3" pageNumber="406">
In habitus,
<taxonomicName id="58C040F6FFF3E86AFEC6E50CFE8FA755" authority="Bezděk, 2012" authorityName="Bezděk" authorityYear="2012" box="[256,353,797,820]" class="Insecta" family="Chrysomelidae" genus="Beenenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="ADB4E767FFF3E86AFEC6E50CFE8FA755" box="[256,353,797,820]" italics="true" pageId="3" pageNumber="406">Beenenia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6875A1CFFF3E86AFEA0E50DFE2AA752" box="[358,452,796,820]" pageId="3" pageNumber="406" rank="genus">gen. nov.</taxonomicNameLabel>
is similar also to several genera from the section Coelomerites (characterized by tibiae with longitudinal ridges). However, only in the genus
<taxonomicName id="58C040F6FFF3E86AFC6AE52CFF14A715" authority="Jacoby, 1888" authorityName="Jacoby" authorityYear="1888" class="Insecta" family="Chrysomelidae" genus="Schematizella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="ADB4E767FFF3E86AFC6AE52CFBAEA735" box="[940,1088,829,852]" italics="true" pageId="3" pageNumber="406">Schematizella</emphasis>
Jacoby, 1888
</taxonomicName>
epipleura are significantly reduced visible in the basal fifth and than extremely narrow or vanishing (Jacoby 1888; R. Beenen, pers. comm. 2011). Antennomeres IIVI are not flattened in
<taxonomicName id="58C040F6FFF3E86AFED3E58CFE47A7D5" authorityName="Jacoby" authorityYear="1888" box="[277,425,925,948]" class="Insecta" family="Chrysomelidae" genus="Schematizella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="ADB4E767FFF3E86AFED3E58CFE47A7D5" box="[277,425,925,948]" italics="true" pageId="3" pageNumber="406">Schematizella</emphasis>
</taxonomicName>
. Other similar genera of Coelomerites differ in well developed epipleura and absence of pubescence on dorsal side of the body (
<taxonomicName id="58C040F6FFF3E86AFCEEE5ACFBF2A7B5" authority="Jacoby, 1903" authorityName="Jacoby" authorityYear="1903" box="[808,1052,955,980]" class="Insecta" family="Chrysomelidae" genus="Arimetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="ADB4E767FFF3E86AFCEEE5ACFC66A7B5" box="[808,904,957,980]" italics="true" pageId="3" pageNumber="406">Arimetus</emphasis>
Jacoby, 1903
</taxonomicName>
) or in mucronate mid tibiae (
<taxonomicName id="58C040F6FFF3E86AFEB9E5CCFD6DA795" authority="Weise, 1902" authorityName="Weise" authorityYear="1902" box="[383,643,987,1012]" class="Insecta" family="Chrysomelidae" genus="Dircemella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="ADB4E767FFF3E86AFEB9E5CCFE18A795" box="[383,502,989,1012]" italics="true" pageId="3" pageNumber="406">Dircemella</emphasis>
Weise, 1902
</taxonomicName>
;
<taxonomicName id="58C040F6FFF3E86AFD4BE5CCFC46A795" authority="Weise, 1902" authorityName="Weise" authorityYear="1902" box="[653,936,987,1012]" class="Insecta" family="Chrysomelidae" genus="Hemiphracta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="ADB4E767FFF3E86AFD4BE5CCFCF4A795" box="[653,794,989,1012]" italics="true" pageId="3" pageNumber="406">Hemiphracta</emphasis>
Weise, 1902
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="D7DA68FEFFF3E86AFFB7E5EDFE30A052" pageId="3" pageNumber="406" type="etymology">
<paragraph id="9F7F3B75FFF3E86AFFB7E5EDFE30A052" blockId="3.[113,1089,219,1075]" pageId="3" pageNumber="406">
<emphasis id="ADB4E767FFF3E86AFFB7E5EDFF1FA075" bold="true" box="[113,241,1020,1044]" pageId="3" pageNumber="406">Etymology.</emphasis>
Dedicated to Ron Beenen (Nieuwegein,
<collectingCountry id="E7D77BE5FFF3E86AFD5FE5EAFCA5A072" box="[665,843,1019,1043]" name="Netherlands" pageId="3" pageNumber="406">The Netherlands</collectingCountry>
), an excellent specialist in
<taxonomicName id="58C040F6FFF3E86AFF4BE20AFEFFA052" authorityName="Latreille" authorityYear="1802" box="[141,273,1051,1075]" pageId="3" pageNumber="404" rank="subFamily" subFamily="Galerucinae">Galerucinae</taxonomicName>
. Gender: feminine.
</paragraph>
</subSubSection>
</treatment>
</document>