305 lines
42 KiB
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305 lines
42 KiB
XML
<document id="1A97C144362F6F6F12BF468DB9FB7767" ID-CLB-Dataset="107843" ID-DOI="10.17109/AZH.65.1.9.2019" ID-GBIF-Dataset="f91eecb6-ca67-4cab-bc26-e5f2c0b2f2f6" ID-ISSN="2064-2474" ID-Zenodo-Dep="5734697" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638188792573" checkinUser="felipe" docAuthor="Kondorosy, Előd, Kovács, Szilvia & Zsalakovics, László" docDate="2019" docId="174F2C559101F007139D14F8FCBBFD6C" docLanguage="en" docName="ActaZoolAcadSciH.65.1.9-18.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 65 (1)" docSource="http://dx.doi.org/10.17109/azh.65.1.9.2019" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.4:ActaZoolAcadSciH.2015-.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-.journal_article" docStyleVersion="4" docTitle="Aristaenetus (Kmentaenetus) Kondorosy 2019, subgen. n." docType="treatment" docUuid="7FE778F2-B5F7-4A8F-B06B-2ABE289EF2F7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="13" masterDocId="EB76542D9100F003120A1058FFB0FFE0" masterDocTitle="Kmentaenetus, A New Subgenus Of Aristaenetus (Hemiptera: Rhyparochromidae: Lethaeini) From The Papuan Subregion" masterLastPageNumber="18" masterPageNumber="9" pageNumber="10" updateTime="1699470959474" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="E837D8E074835C81EF90B590CFA5D9EA">Kmentaenetus, A New Subgenus Of Aristaenetus (Hemiptera: Rhyparochromidae: Lethaeini) From The Papuan Subregion</mods:title>
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<mods:namePart id="A417BDB809E8397F019684DA6A6C131A">Kondorosy, Előd</mods:namePart>
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<mods:namePart id="69A1C67950B260C76EE817E879DE2FA1">Kovács, Szilvia</mods:namePart>
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<mods:namePart id="42197AA57493C69B7F24C3737E48C8F3">Zsalakovics, László</mods:namePart>
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<treatment id="174F2C559101F007139D14F8FCBBFD6C" ID-DOI="http://doi.org/10.5281/zenodo.6929242" ID-GBIF-Taxon="197867521" ID-Zenodo-Dep="6929242" LSID="urn:lsid:zoobank.org:act:7FE778F2-B5F7-4A8F-B06B-2ABE289EF2F7" httpUri="http://treatment.plazi.org/id/174F2C559101F007139D14F8FCBBFD6C" lastPageId="4" lastPageNumber="13" pageId="1" pageNumber="10">
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<subSubSection id="D7FCCEC89101F002139D14F8FCCDFB5A" box="[407,893,1184,1210]" pageId="1" pageNumber="10" type="nomenclature">
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<paragraph id="9F599D439101F002139D14F8FCCDFB5A" blockId="1.[407,893,1184,1210]" box="[407,893,1184,1210]" pageId="1" pageNumber="10">
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<heading id="C4112A2F9101F002139D14F8FCCDFB5A" box="[407,893,1184,1210]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="10" reason="2">
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<taxonomicName id="58E6E6C09101F002139D14F8FD41FB5A" ID-CoL="8QLWX" authority="Kondorosy" authorityName="Kondorosy" authorityYear="2019" box="[407,753,1184,1210]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="10" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Kmentaenetus">
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<emphasis id="AD9241519101F002139D14F8FDE9FB5A" bold="true" box="[407,601,1184,1210]" pageId="1" pageNumber="10">Kmentaenetus</emphasis>
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Kondorosy
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</taxonomicName>
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<taxonomicNameLabel id="B6A1FC2A9101F00210F214F8FCCDFB5A" box="[760,893,1184,1210]" pageId="1" pageNumber="10" rank="subGenus">subgen. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D7FCCEC89101F00212A41480FC36FB0D" box="[174,902,1239,1261]" pageId="1" pageNumber="10" type="type_taxon">
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<paragraph id="9F599D439101F00212A41480FC36FB0D" blockId="1.[174,902,1239,1261]" box="[174,902,1239,1261]" pageId="1" pageNumber="10">
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<typeStatus id="405D23E19101F00212A41480FF54FB0D" box="[174,228,1240,1261]" pageId="1" pageNumber="10">Type</typeStatus>
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species:
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<taxonomicName id="58E6E6C09101F002134B1480FD75FB0C" ID-CoL="8QPH7" authorityName="Kondorosy" authorityYear="2019" box="[321,709,1239,1261]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="10" phylum="Arthropoda" rank="species" species="diabolicus" subGenus="Kmentaenetus">
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<emphasis id="AD9241519101F002134B1480FD75FB0C" box="[321,709,1239,1261]" italics="true" pageId="1" pageNumber="10">Aristaenetus (Kmentaenetus) diabolicus</emphasis>
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</taxonomicName>
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Kondorosy,
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<taxonomicNameLabel id="B6A1FC2A9101F00211471480FC36FB0D" box="[845,902,1240,1261]" pageId="1" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="D7FCCEC89101F00012E01549FD5CFEEE" lastPageId="3" lastPageNumber="12" pageId="1" pageNumber="10" type="description">
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<paragraph id="9F599D439101F00212E01549FD2EFAA4" blockId="1.[174,1127,1297,1578]" pageId="1" pageNumber="10">Description: Body moderately elongate, punctate except of head dorsally and two spots on pronotum. Body dull, almost hairless.</paragraph>
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<paragraph id="9F599D439101F00112E01513FECEFEA7" blockId="1.[174,1127,1297,1578]" lastBlockId="2.[136,1089,190,1566]" lastPageId="2" lastPageNumber="11" pageId="1" pageNumber="10">Head pentagonal, dorsally and laterally impunctate, ventrally strongly transversely rugose, submedially punctate (punctures lacking around eyes, base of antennae, and anteriad of antenniferous tubercles); much longer then wide; dorsal side transversely rugose (except around and anterior to ocelli), midline relatively strongly rugose, reaching clypeus. Head everywhere with hardly visible tiny decumbent setae. Apex of clypeus acute, toothlike. Compound eyes prominent, large, postocular part longitudinally rugose, ventrally with transverse wrinkles. Ocelli at posterior margin of eyes, equidistant from eye and median. Antenniferous tubercle divergent, prominent, almost as long as eye, head almost as wide here as base of head. Base of head with two iridescent spots. Maxillary plates convex, smooth and shining, partly visible in dorsal view. Labium long, reaching middle of abdomen. Scape extending with half beyond apex of head. Antennae with very fine and dense decumbent pubescence and few stronger setae on scape, other segments with some fine semierect setae as well.</paragraph>
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<paragraph id="9F599D439102F00112CF1116FC54FC1B" blockId="2.[136,1089,190,1566]" pageId="2" pageNumber="11">Lateral margin of pronotum noticeably concave, apical half narrower. Posterior part of pronotal collar densely punctate, separated from pronotum with punctate furrow, pronotum behind collar slightly elevated. Transverse furrow shallow, inconspicuous, lateral margin of pronotum hardly visible, not laminate. Midline elevated, impunctate, reaching from middle of anterior lobe almost to posterior margin. Anterior lobe of pronotum with partly impunctate but not elevated calli. Humeral angles of pronotum not rounded. Anterior margin of pronotum nearly straight, medially slightly protruding, posterior margin slightly convex. Pronotum without anterolateral trichobothria. Scutellum triangular, largely punctate, with blunt Y-carina. Clavus with four regular rows of punctures, both middle rows united anteriorly near base. Corium between veins Sc and R densely, between R and Cu shallower and sparser punctate. Costal margin slightly S-shaped, apical margin nearly straight. Membrane with two closed basal cells, 4 veins visible. Coxae finely transversely rugose, legs otherwise smooth. Profemur slightly thicker at base than in apical half, anterior margin with three small teeth apically and 5 long thin setae; other femora unarmed, with some short stronger setae only. Tibiae straight, with rows of strong stiff setae, inner side in apical half also with fine semi-decumbent pubescence (on metatibia negligible). Tarsi with very dense short fine semi-decumbent pubescence beneath. Pleura with impunctate or rugose areas between punctures, sternum impunctate and dull except ventral collar, mesothorax ventrally with medial furrow, metasternum with strong medial carina. Mesosternum limited anteriorly and posteriorly with strong arched transverse carina. Ostiolar peritreme on male directed laterad (reaching about 3/4 of metapleuron), then curving posteriorly, terminating in semi-circle directed anteriad or laterad; ostiolar peritreme significantly shorter in females, directed laterally, then slightly posteriad. Evaporatorium large, reaching lateral margin of mesopleuron, occupying majority of metapleuron, almost reaching metepimeroid.</paragraph>
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<paragraph id="9F599D439102F00112CF1459FD62FA91" blockId="2.[136,1089,190,1566]" pageId="2" pageNumber="11">
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Abdomen with extremely dense micropunctures and tiny setae on ventral side. Median third of sternites III–IV smooth, lateral thirds with long transverse rugae with their lateral extremities curved posteriad submarginally. Surface of abdominal venter uneven, sternite III with median keel and with flat submedial elevation bearing trichobothria; base of sternite VI impressed sublaterally. Posterior margins of sternites III to V narrowly produced posteriorly at midline, lateral portions broadly S-shaped, curved anteriad, abruptly curved posteriad submarginally. Suture between sterna 4 and 5 curving cephalad, not reaching lateral margin; trichobothria and spiracles as usual in
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<taxonomicName id="58E6E6C09102F00111371493FC13FB01" authorityName="Stal" authorityYear="1872" box="[829,931,1227,1249]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="11" phylum="Arthropoda" rank="tribe" tribe="Lethaeini">Lethaeini</taxonomicName>
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. Sternite VI of female strongly narrowed but not divided medially, together with sternite V medially keeled. Tergites shiny, finely rugose and extremely finely punctate transversely (wrinkles of tergite VIII stronger); posterior portions of tergite VIII and hypopleurite VIII connected by a broad, membranous cuticular area. Posterior margin of tergite VIII medially straight, laterally convex, hypopleurite VIII terminating acutely.
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</paragraph>
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<paragraph id="9F599D439102F00112CF1520FC32F9FE" blockId="2.[136,1089,190,1566]" pageId="2" pageNumber="11">
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Male genital capsule (
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<figureCitation id="07DD81C69102F00113BB1520FE4CFA6E" box="[433,508,1400,1422]" captionStart="Fig" captionStartId="6.[136,172,1526,1548]" captionTargetBox="[140,1071,878,1495]" captionTargetId="figure-200@6.[136,1077,876,1501]" captionTargetPageId="6" captionText="Fig. 4. Aristaenetus (Kmentaenetus) woodwardi Kondorosy et Zsalakovics holotype male, dor- sal view" figureDoi="http://doi.org/10.5281/zenodo.5734703" httpUri="https://zenodo.org/record/5734703/files/figure.png" pageId="2" pageNumber="11">Fig 4D</figureCitation>
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) subglobose, with fine decumbent pubescence, posterior aperture nearly rhomboidal, anterior half with two acute and a blunt teeth on each side, cuplike sclerite large, reaching about halfway of aperture, with a blunt apex medially and with sinuous arches on both sides. Paramere broad, subtriangular, inner projection very broad, terminating in a tooth orientated to base; outer projection elongate apically in a tooth (as
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<figureCitation id="07DD81C69102F00112F61650FEFAF9FE" box="[252,330,1544,1566]" captionStart="Fig" captionStartId="6.[136,172,1526,1548]" captionTargetBox="[140,1071,878,1495]" captionTargetId="figure-200@6.[136,1077,876,1501]" captionTargetPageId="6" captionText="Fig. 4. Aristaenetus (Kmentaenetus) woodwardi Kondorosy et Zsalakovics holotype male, dor- sal view" figureDoi="http://doi.org/10.5281/zenodo.5734703" httpUri="https://zenodo.org/record/5734703/files/figure.png" pageId="2" pageNumber="11">Fig. 4A</figureCitation>
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); base of blade dorsally with a strong hump (
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<figureCitation id="07DD81C69102F00111221650FCC6F9FE" box="[808,886,1544,1566]" captionStart="Fig" captionStartId="6.[136,172,1526,1548]" captionTargetBox="[140,1071,878,1495]" captionTargetId="figure-200@6.[136,1077,876,1501]" captionTargetPageId="6" captionText="Fig. 4. Aristaenetus (Kmentaenetus) woodwardi Kondorosy et Zsalakovics holotype male, dor- sal view" figureDoi="http://doi.org/10.5281/zenodo.5734703" httpUri="https://zenodo.org/record/5734703/files/figure.png" pageId="2" pageNumber="11">Fig. 4C</figureCitation>
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).
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</paragraph>
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<paragraph id="9F599D439103F00012E010E6FD5CFEEE" blockId="3.[174,1127,190,385]" pageId="3" pageNumber="12">
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Spermathecal duct of female extremely long and tortuous (several times longer than body length), apical part after an oblique ring suddenly broadened to subglobular apical receptacle, significantly impressed lengthwise (
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<figureCitation id="07DD81C69103F00010A910A0FD50FEEE" box="[675,736,248,270]" captionStart="Figs 2–3" captionStartId="5.[174,219,1496,1518]" captionTargetBox="[175,1123,1125,1468]" captionTargetId="figure-154@5.[174,1126,1124,1470]" captionTargetPageId="5" captionText="Figs 2–3. 2 = Spermatheca of A. diabolicus Kondorosy. 3 = Male genitalia of Aristaenetus (Kmentaenetus) woodwardi Kondorosy et Zsalakovics; A–C: paramere, A = dorsal view, B = ventral view, C = lateral view, D = pygophore, dorsal view" figureDoi="http://doi.org/10.5281/zenodo.5734701" httpUri="https://zenodo.org/record/5734701/files/figure.png" pageId="3" pageNumber="12">Fig. 2</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="D7FCCEC89103F00012E0114DFDE1FE60" pageId="3" pageNumber="12" type="etymology">
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<paragraph id="9F599D439103F00012E0114DFDE1FE60" blockId="3.[174,1127,190,385]" pageId="3" pageNumber="12">
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Etymology – We named the new subgenus after the excellent heteropterologist Dr. Petr Kment who brought our attention to these specimens, was kind enough to loan them to us, and helped also with sending much old and new literature. The ending of the name is in reference to the genus
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<taxonomicName id="58E6E6C09103F00013C51133FDFBFE60" authorityName="Distant" authorityYear="1901" box="[463,587,363,384]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="AD9241519103F00013C51133FDFBFE60" box="[463,587,363,384]" italics="true" pageId="3" pageNumber="12">Aristaenetus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="D7FCCEC89103F00712E011FEFCBBFD6C" lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12" type="discussion">
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<paragraph id="9F599D439103F00012E011FEFD65FC58" blockId="3.[174,1127,422,1557]" pageId="3" pageNumber="12">
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Remarks – This new subgenus is erected to include the two new species described in this paper. On the Eastern Hemisphere species of 9 genera of the
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<taxonomicName id="58E6E6C09103F000130911B1FEC7FDE3" authorityName="Stal" authorityYear="1872" box="[259,375,489,515]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Lethaeini">Lethaeini</taxonomicName>
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have closed cells on the base of the membrane, similar to
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<emphasis id="AD9241519103F00012A41253FEE0FDC4" box="[174,336,523,548]" italics="true" pageId="3" pageNumber="12">Kmentaenetus</emphasis>
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. The largest genus,
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<taxonomicName id="58E6E6C09103F000105E1253FCEAFDC5" authority="Dallas, 1852" authorityName="Dallas" authorityYear="1852" box="[596,858,522,549]" class="Insecta" family="Rhyparochromidae" genus="Lethaeus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="AD9241519103F000105E1253FD08FDC5" box="[596,696,523,549]" italics="true" pageId="3" pageNumber="12">Lethaeus</emphasis>
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Dallas, 1852
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</taxonomicName>
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lacks a collar on the pronotum. Five further genera, like
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<taxonomicName id="58E6E6C09103F000107F1274FC1AFDA6" authority="Distant, 1909" authorityName="Distant" authorityYear="1909" box="[629,938,556,582]" class="Insecta" family="Rhyparochromidae" genus="Neolethaeus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="AD9241519103F000107F1274FD4EFDA6" box="[629,766,556,582]" italics="true" pageId="3" pageNumber="12">Neolethaeus</emphasis>
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Distant, 1909
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</taxonomicName>
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have a convex or straight lateral margin on pronotum (lateral margin is mostly narrowly laminate). Species of two of the remaining genera with concave pronotum (
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<taxonomicName id="58E6E6C09103F00012BD12C9FDF6FD4B" authority="Kondorosy, 2006" authorityName="Kondorosy" authorityYear="2006" box="[183,582,657,683]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetoides" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="AD9241519103F00012BD12C9FED5FD4B" box="[183,357,657,683]" italics="true" pageId="3" pageNumber="12">Aristaenetoides</emphasis>
|
||
<bibRefCitation id="FB77E0B29103F000137B12C9FDF6FD4B" author="KONDOROSY, E." box="[369,582,657,683]" pageId="3" pageNumber="12" pagination="115 - 119" refId="ref4503" refString="KONDOROSY, E. (2006): A new genus of Lethaeini (Insecta: Heteroptera: Lygaeidae) from the Oriental Region. - Annalen des Naturhistorischen Museums in Wien 107 B: 115 - 119." type="journal article" year="2006">Kondorosy, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="58E6E6C09103F000108612C9FBA7FD4C" authority="Scudder, 1962" authorityName="Scudder" authorityYear="1962" box="[652,1047,657,684]" class="Insecta" family="Rhyparochromidae" genus="Lethaeograndellus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F000108612C9FCE9FD4B" box="[652,857,657,683]" italics="true" pageId="3" pageNumber="12">Lethaeograndellus</emphasis>
|
||
Scudder, 1962
|
||
</taxonomicName>
|
||
) have among others punctate head ventrally, only
|
||
<taxonomicName id="58E6E6C09103F00010E212EBFCC9FD2D" authorityName="Distant" authorityYear="1901" box="[744,889,691,717]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F00010E212EBFCC9FD2D" box="[744,889,691,717]" italics="true" pageId="3" pageNumber="12">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
has similar striate head ventrally as
|
||
<emphasis id="AD9241519103F0001386128DFD9EFD0E" box="[396,558,725,750]" italics="true" pageId="3" pageNumber="12">Kmentaenetus</emphasis>
|
||
. Furthermore,
|
||
<taxonomicName id="58E6E6C09103F00010E0128CFC28FD0E" authorityName="Kondorosy" authorityYear="2006" box="[746,920,724,750]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetoides" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F00010E0128CFC28FD0E" box="[746,920,724,750]" italics="true" pageId="3" pageNumber="12">Aristaenetoides</emphasis>
|
||
</taxonomicName>
|
||
has only 3 rows of punctures on clavus (contrary to 4 of
|
||
<taxonomicName id="58E6E6C09103F000109B12AEFC92FCF0" authorityName="Distant" authorityYear="1901" box="[657,802,758,784]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F000109B12AEFC92FCF0" box="[657,802,758,784]" italics="true" pageId="3" pageNumber="12">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="AD9241519103F000115112AFFC4DFCF0" box="[859,1021,759,784]" italics="true" pageId="3" pageNumber="12">Kmentaenetus</emphasis>
|
||
).
|
||
<taxonomicName id="58E6E6C09103F000161E12AEFE82FCD1" authorityName="Scudder" authorityYear="1962" class="Insecta" family="Rhyparochromidae" genus="Lethaeograndellus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F000161E12AEFE82FCD1" italics="true" pageId="3" pageNumber="12">Lethaeograndellus</emphasis>
|
||
</taxonomicName>
|
||
has lamellate lateral margin of pronotum, punctate pronotal collar forming a V, and rugose head, callar area of pronotum and abdomen, while
|
||
<taxonomicName id="58E6E6C09103F00012A41303FE8FFC95" authorityName="Distant" authorityYear="1901" box="[174,319,859,885]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F00012A41303FE8FFC95" box="[174,319,859,885]" italics="true" pageId="3" pageNumber="12">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="AD9241519103F000138A1303FD92FC94" box="[384,546,859,884]" italics="true" pageId="3" pageNumber="12">Kmentaenetus</emphasis>
|
||
have weakly carinate lateral margin of pronotum, impunctate and slightly arched pronotal collar, and smooth or punctate head, callar area of pronotum and abdomen.
|
||
</paragraph>
|
||
<paragraph id="9F599D439103F00012E013E7FD36FA6E" blockId="3.[174,1127,422,1557]" pageId="3" pageNumber="12">
|
||
The anterolateral trichobothria of the pronotum are often missing e.g. in
|
||
<taxonomicName id="58E6E6C09103F00012A413B9FE87FC1B" authorityName="Distant" authorityYear="1909" box="[174,311,993,1019]" class="Insecta" family="Rhyparochromidae" genus="Neolethaeus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F00012A413B9FE87FC1B" box="[174,311,993,1019]" italics="true" pageId="3" pageNumber="12">Neolethaeus</emphasis>
|
||
</taxonomicName>
|
||
, as in
|
||
<taxonomicName id="58E6E6C09103F000138413BAFD41FC1B" authority="Woodward, 1968" authorityName="Woodward" authorityYear="1968" box="[398,753,993,1019]" class="Insecta" family="Rhyparochromidae" genus="Neolethaeus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="cantrelli">
|
||
<emphasis id="AD9241519103F000138413BAFDA2FC1B" box="[398,530,993,1019]" italics="true" pageId="3" pageNumber="12">N. cantrelli</emphasis>
|
||
<bibRefCitation id="FB77E0B29103F000101613B9FD41FC1B" author="WOODWARD, T. E." box="[540,753,993,1019]" pageId="3" pageNumber="12" pagination="35 - 42" refId="ref5110" refString="WOODWARD, T. E. (1968): A new species-pair of Neolethaeus Distant (Hemiptera: Lygaei- dae) from Northern Queensland and New Guinea. - Proceedings of the Royal Society of Queensland 80 (4): 35 - 42." type="journal article" year="1968">Woodward, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="58E6E6C09103F000111513BAFF56FBFC" authority="Woodward, 1968" authorityName="Woodward" authorityYear="1968" class="Insecta" family="Rhyparochromidae" genus="Neolethaeus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="cheesmanae">
|
||
<emphasis id="AD9241519103F000111513BAFC79FC1B" box="[799,969,993,1019]" italics="true" pageId="3" pageNumber="12">N. cheesmanae</emphasis>
|
||
<bibRefCitation id="FB77E0B29103F00011D913B9FF56FBFC" author="WOODWARD, T. E." pageId="3" pageNumber="12" pagination="35 - 42" refId="ref5110" refString="WOODWARD, T. E. (1968): A new species-pair of Neolethaeus Distant (Hemiptera: Lygaei- dae) from Northern Queensland and New Guinea. - Proceedings of the Royal Society of Queensland 80 (4): 35 - 42." type="journal article" year="1968">Woodward, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
(discussed by
|
||
<bibRefCitation id="FB77E0B29103F00013AA145BFDC3FBFD" author="WOODWARD, T. E." box="[416,627,1026,1053]" pageId="3" pageNumber="12" pagination="35 - 42" refId="ref5110" refString="WOODWARD, T. E. (1968): A new species-pair of Neolethaeus Distant (Hemiptera: Lygaei- dae) from Northern Queensland and New Guinea. - Proceedings of the Royal Society of Queensland 80 (4): 35 - 42." type="journal article" year="1968">WOODWARD 1968</bibRefCitation>
|
||
), so presence or absence of the pronotal trichobothrium is not always a constant feature in a genus. One of the most unique features is the elongate and strongly curved ostiolar peritreme of the males, which is characteristic only for the species of
|
||
<taxonomicName id="58E6E6C09103F00011251430FBD6FB62" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[815,1126,1127,1154]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F00011251430FC70FB62" box="[815,960,1128,1154]" italics="true" pageId="3" pageNumber="12">Aristaenetus</emphasis>
|
||
Distant, 1901
|
||
</taxonomicName>
|
||
and three species of
|
||
<taxonomicName id="58E6E6C09103F00013B914D1FD8CFB43" authorityName="Distant" authorityYear="1909" box="[435,572,1161,1187]" class="Insecta" family="Rhyparochromidae" genus="Neolethaeus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F00013B914D1FD8CFB43" box="[435,572,1161,1187]" italics="true" pageId="3" pageNumber="12">Neolethaeus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="58E6E6C09103F000104614D2FD60FB43" authorityName="Woodward" authorityYear="1968" box="[588,720,1161,1187]" class="Insecta" family="Rhyparochromidae" genus="Neolethaeus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="cantrelli">
|
||
<emphasis id="AD9241519103F000104614D2FD60FB43" box="[588,720,1161,1187]" italics="true" pageId="3" pageNumber="12">N. cantrelli</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="58E6E6C09103F00010EB14D2FC3AFB43" authorityName="Woodward" authorityYear="1968" box="[737,906,1161,1187]" class="Insecta" family="Rhyparochromidae" genus="Neolethaeus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="cheesmanae">
|
||
<emphasis id="AD9241519103F00010EB14D2FC3AFB43" box="[737,906,1161,1187]" italics="true" pageId="3" pageNumber="12">N. cheesmanae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="58E6E6C09103F00011C614D2FEE0FB25" authority="Walker, 1872" authorityName="Walker" authorityYear="1872" class="Insecta" family="Rhyparochromidae" genus="Neolethaeus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="descriptus">
|
||
<emphasis id="AD9241519103F00011C614D2FBD7FB43" box="[972,1127,1161,1187]" italics="true" pageId="3" pageNumber="12">N. descriptus</emphasis>
|
||
Walker, 1872
|
||
</taxonomicName>
|
||
, and in somewhat less developed form for the American genus
|
||
<taxonomicName id="58E6E6C09103F00012A41494FE00FB06" authority="Distant, 1893" authorityName="Distant" authorityYear="1893" box="[174,432,1228,1254]" class="Insecta" family="Rhyparochromidae" genus="Bubaces" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F00012A41494FEB9FB06" box="[174,265,1228,1254]" italics="true" pageId="3" pageNumber="12">Bubaces</emphasis>
|
||
Distant, 1893
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="FB77E0B29103F00013E41494FD0BFB07" author="O'DONNELL, J. E." box="[494,699,1228,1255]" pageId="3" pageNumber="12" pagination="535 - 538" refId="ref4721" refString="O'DONNELL, J. E. (2006): A new species of Bubaces Distant (Heteroptera, Rhyparochromi- dae, Rhyparochrominae). - Denisia 19: 535 - 538." type="journal article" year="2006">O'DONNELL 2006</bibRefCitation>
|
||
).
|
||
<taxonomicName id="58E6E6C09103F00010DA1494FCE9FB06" authorityName="Distant" authorityYear="1909" box="[720,857,1228,1254]" class="Insecta" family="Rhyparochromidae" genus="Neolethaeus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F00010DA1494FCE9FB06" box="[720,857,1228,1254]" italics="true" pageId="3" pageNumber="12">Neolethaeus</emphasis>
|
||
</taxonomicName>
|
||
is not very closely related to the new species because of the different pronotum (they have lateral carina and at most slightly concave lateral margin) and shape of the paramere (
|
||
<figureCitation id="07DD81C69103F00012BC1569FEA5FAAB" box="[182,277,1329,1355]" captionStart="Figs 2–3" captionStartId="5.[174,219,1496,1518]" captionTargetBox="[175,1123,1125,1468]" captionTargetId="figure-154@5.[174,1126,1124,1470]" captionTargetPageId="5" captionText="Figs 2–3. 2 = Spermatheca of A. diabolicus Kondorosy. 3 = Male genitalia of Aristaenetus (Kmentaenetus) woodwardi Kondorosy et Zsalakovics; A–C: paramere, A = dorsal view, B = ventral view, C = lateral view, D = pygophore, dorsal view" figureDoi="http://doi.org/10.5281/zenodo.5734701" httpUri="https://zenodo.org/record/5734701/files/figure.png" pageId="3" pageNumber="12">Fig. 3A</figureCitation>
|
||
). So, we concluded that our species are most closely related to the genus
|
||
<taxonomicName id="58E6E6C09103F000130A150BFE21FA8D" authorityName="Distant" authorityYear="1901" box="[256,401,1363,1389]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD9241519103F000130A150BFE21FA8D" box="[256,401,1363,1389]" italics="true" pageId="3" pageNumber="12">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
but because of the important differences we suppose it is appropriate to create a new subgenus.
|
||
</paragraph>
|
||
<paragraph id="9F599D439103F00012E015CEFECAF9F5" blockId="3.[174,1127,422,1557]" pageId="3" pageNumber="12">
|
||
<bibRefCitation id="FB77E0B29103F00012E015CEFE79FA50" author="O'DONNELL, J. E." box="[234,457,1430,1456]" pageId="3" pageNumber="12" pagination="441 - 470" refId="ref4681" refString="O'DONNELL, J. E. (1991): A survey of male genitalia in Lethaeine genera (Heteroptera: Lygaeidae: Rhyparochrominae). - Journal of the New York Entomological Society 99: 441 - 470." type="journal article" year="1991">O'DONNELL (1991)</bibRefCitation>
|
||
investigated the male genitalia of most lethaeine genera, including both
|
||
<taxonomicName id="58E6E6C09103F00013AE15E0FD2CFA31" box="[420,668,1463,1490]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="AD9241519103F00013AE15E0FD85FA32" box="[420,565,1464,1490]" italics="true" pageId="3" pageNumber="12">Aristaenetus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
The paramere of
|
||
<taxonomicName id="58E6E6C09103F000117115E0FBABFA31" authority="Kondorosy & Kovács & Zsalakovics, 2019" authorityName="Kondorosy & Kovács & Zsalakovics" authorityYear="2019" box="[891,1051,1463,1490]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="woodwardi" status="sp. nov.">
|
||
<emphasis id="AD9241519103F000117115E0FBABFA31" box="[891,1051,1463,1490]" italics="true" pageId="3" pageNumber="12">A. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B6A1FC2A9103F000162915E0FBD6FA32" box="[1059,1126,1464,1490]" pageId="3" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
|
||
is unique (
|
||
<figureCitation id="07DD81C69103F00013321581FE24FA13" box="[312,404,1497,1523]" captionStart="Figs 2–3" captionStartId="5.[174,219,1496,1518]" captionTargetBox="[175,1123,1125,1468]" captionTargetId="figure-154@5.[174,1126,1124,1470]" captionTargetPageId="5" captionText="Figs 2–3. 2 = Spermatheca of A. diabolicus Kondorosy. 3 = Male genitalia of Aristaenetus (Kmentaenetus) woodwardi Kondorosy et Zsalakovics; A–C: paramere, A = dorsal view, B = ventral view, C = lateral view, D = pygophore, dorsal view" figureDoi="http://doi.org/10.5281/zenodo.5734701" httpUri="https://zenodo.org/record/5734701/files/figure.png" pageId="3" pageNumber="12">Fig. 3A</figureCitation>
|
||
) and similar to the paramere of
|
||
<taxonomicName id="58E6E6C09103F00011371581FEC6F9F5" authority="Woodward et O'Donnell, 1988" authorityName="Woodward et O'Donnell" authorityYear="1988" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="AD9241519103F00011371581FC1EFA13" box="[829,942,1497,1523]" italics="true" pageId="3" pageNumber="12">A. similis</emphasis>
|
||
<bibRefCitation id="FB77E0B29103F00011B21581FEC6F9F5" author="WOODWARD, T. E. & O'DONNELL, J. E." pageId="3" pageNumber="12" pagination="481 - 491" refId="ref5156" refString="WOODWARD, T. E. & O'DONNELL, J. E. (1988): The genus Aristaenetus Distant (Hemiptera: Lygaeidae: Rhyparochrominae) with the description of a new species. - Memoirs of the Queensland Museum 25 (2): 481 - 491." type="journal article" year="1988">Woodward et O'Donnell, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9F599D439104F00712CF10E5FCBBFD6C" blockId="4.[136,1089,189,653]" pageId="4" pageNumber="13">
|
||
The main differences between the two subgenera are that the species belonging to
|
||
<taxonomicName id="58E6E6C09104F00713001087FE2BFF19" authorityName="Distant" authorityYear="1901" box="[266,411,223,249]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="AD9241519104F00713001087FE2BFF19" box="[266,411,223,249]" italics="true" pageId="4" pageNumber="13">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B6A1FC2A9104F007139510B8FE6EFF1A" box="[415,478,224,250]" pageId="4" pageNumber="13" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
have anterolateral trichobothria on the pronotum, however, in
|
||
<emphasis id="AD9241519104F0071329115AFE75FEFB" box="[291,453,258,283]" italics="true" pageId="4" pageNumber="13">Kmentaenetus</emphasis>
|
||
these are missing; also, the shape of the pronotum is different, the anterior half of the pronotum is wider in
|
||
<taxonomicName id="58E6E6C09104F0071157117BFC5EFEDD" authorityName="Distant" authorityYear="1901" box="[861,1006,291,317]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="AD9241519104F0071157117BFC5EFEDD" box="[861,1006,291,317]" italics="true" pageId="4" pageNumber="13">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B6A1FC2A9104F00711FD117BFB8BFEDD" box="[1015,1083,291,317]" pageId="4" pageNumber="13" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
, while it is much more slender in
|
||
<emphasis id="AD9241519104F0071011111DFD0DFEBE" box="[539,701,325,350]" italics="true" pageId="4" pageNumber="13">Kmentaenetus</emphasis>
|
||
species. Furthermore,
|
||
<emphasis id="AD9241519104F00711DE111DFF76FE9F" italics="true" pageId="4" pageNumber="13">Kmentaenetus</emphasis>
|
||
species have transversely rugose heads while
|
||
<taxonomicName id="58E6E6C09104F007110D113EFC28FE60" authorityName="Distant" authorityYear="1901" box="[775,920,358,384]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="AD9241519104F007110D113EFC28FE60" box="[775,920,358,384]" italics="true" pageId="4" pageNumber="13">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B6A1FC2A9104F00711AA113EFC51FE60" box="[928,993,358,384]" pageId="4" pageNumber="13" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
species have punctate heads. The eye-antenna distance is in
|
||
<emphasis id="AD9241519104F007112911D0FC75FE41" box="[803,965,392,417]" italics="true" pageId="4" pageNumber="13">Kmentaenetus</emphasis>
|
||
nearly as long as eye length, species, however, in
|
||
<taxonomicName id="58E6E6C09104F007108811F1FCA3FE23" authorityName="Distant" authorityYear="1901" box="[642,787,425,451]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="AD9241519104F007108811F1FCA3FE23" box="[642,787,425,451]" italics="true" pageId="4" pageNumber="13">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B6A1FC2A9104F007111111F1FCEEFE23" box="[795,862,425,451]" pageId="4" pageNumber="13" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
it is about half as long as eye length. In
|
||
<emphasis id="AD9241519104F00713AE1193FDF6FE04" box="[420,582,459,484]" italics="true" pageId="4" pageNumber="13">Kmentaenetus</emphasis>
|
||
the eye is delimited by radial wrinkles, while in
|
||
<taxonomicName id="58E6E6C09104F00712F911B4FE34FDE6" authorityName="Distant" authorityYear="1901" box="[243,388,492,518]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="AD9241519104F00712F911B4FE34FDE6" box="[243,388,492,518]" italics="true" pageId="4" pageNumber="13">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B6A1FC2A9104F007138111B4FE7BFDE6" box="[395,459,492,518]" pageId="4" pageNumber="13" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
the surface delimiting the eyes is smooth. Pronotal collar is impunctate in
|
||
<taxonomicName id="58E6E6C09104F00713AB1256FD82FDC8" authorityName="Distant" authorityYear="1901" box="[417,562,526,552]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="AD9241519104F00713AB1256FD82FDC8" box="[417,562,526,552]" italics="true" pageId="4" pageNumber="13">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B6A1FC2A9104F00710321256FDC9FDC8" box="[568,633,526,552]" pageId="4" pageNumber="13" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
and with punctures in
|
||
<emphasis id="AD9241519104F00711921256FB8AFDC7" box="[920,1082,526,551]" italics="true" pageId="4" pageNumber="13">Kmentaenetus</emphasis>
|
||
. The pronotal humeral angles are angular or tooth-like in
|
||
<emphasis id="AD9241519104F007116F1268FBB7FDA9" box="[869,1031,560,585]" italics="true" pageId="4" pageNumber="13">Kmentaenetus</emphasis>
|
||
, not rounded as in
|
||
<taxonomicName id="58E6E6C09104F007133D1209FE78FD8B" authorityName="Distant" authorityYear="1901" box="[311,456,593,619]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="AD9241519104F007133D1209FE78FD8B" box="[311,456,593,619]" italics="true" pageId="4" pageNumber="13">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B6A1FC2A9104F00713C71209FDBDFD8B" box="[461,525,593,619]" pageId="4" pageNumber="13" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
The abdomen is ventrally only sparsely punctate in
|
||
<taxonomicName id="58E6E6C09104F00712D6122BFEDDFD6D" authorityName="Distant" authorityYear="1901" box="[220,365,627,653]" class="Insecta" family="Rhyparochromidae" genus="Aristaenetus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="AD9241519104F00712D6122BFEDDFD6D" box="[220,365,627,653]" italics="true" pageId="4" pageNumber="13">Aristaenetus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B6A1FC2A9104F007137E122BFE05FD6D" box="[372,437,627,653]" pageId="4" pageNumber="13" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
and not extremely densely.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |