279 lines
23 KiB
XML
279 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.339.4903" ID-GBIF-Dataset="f5e2c4d6-742c-4235-bf0e-b97e1ec678d2" ID-PMC="PMC3800827" ID-Pensoft-Pub="1313-2970-339-1" ID-PubMed="24146585" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-339-1" ModsDocOrigin="ZooKeys 339" ModsDocTitle="Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae)" checkinTime="1451246844427" checkinUser="pensoft" docAuthor="Borkenhagen, Kai & Krupp, Friedhelm" docDate="2013" docId="172EB0233D61E16171EFC009AC764B55" docLanguage="en" docName="ZooKeys 339: 1-53" docOrigin="ZooKeys 339" docSource="http://dx.doi.org/10.3897/zookeys.339.4903" docTitle="Carasobarbus kosswigi Ladiges 1960" docType="treatment" docVersion="3" lastPageNumber="29" masterDocId="FF8B0A687873FFFAFFC0C22FFFF3FFD2" masterDocTitle="Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="27" updateTime="1668156576871" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Borkenhagen, Kai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Krupp, Friedhelm</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>339</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>53</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.339.4903</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.339.4903</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-339-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152048583" LSID="urn:lsid:plazi:treatment:172EB0233D61E16171EFC009AC764B55" httpUri="http://treatment.plazi.org/id/172EB0233D61E16171EFC009AC764B55" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
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<subSubSection pageId="26" pageNumber="27" type="nomenclature">
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName LSID="http://species-id.net/wiki/Carasobarbus_kosswigi" authority="Ladiges, 1960" authorityName="Ladiges" authorityYear="1960" class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi (Ladiges, 1960)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="reference_group">
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Cyclocheilichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocheilichthys kosswigi" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="kosswigi">Cyclocheilichthys kosswigi</taxonomicName>
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<bibRefCitation author="Ladiges, W" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="45" pageNumber="46" pagination="105 - 150" title="Suesswasserfische der Tuerkei. I. Teil Cyprinidae." volume="58" year="1960">Ladiges 1960</bibRefCitation>
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: 135.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="material">
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<paragraph pageId="26" pageNumber="27">Material.</paragraph>
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<paragraph pageId="26" pageNumber="27">
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Type material. Holotype of
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Cyclocheilichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocheilichthys kosswigi" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="kosswigi">Cyclocheilichthys kosswigi</taxonomicName>
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: ZMH H 1148, Turkey, Batman
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<normalizedToken originalValue="Çayı">Cayi</normalizedToken>
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[
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="37.787777">37°47'16"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="41.014168">41°0'51"E</geoCoordinate>
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], C. Kosswig, IV.1939.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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Non-type material. Tigris-Euphrates system. NMW 90369, 1, Turkey, Batman
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<normalizedToken originalValue="Çayı">Cayi</normalizedToken>
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||
near Baschkaja [
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||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="37.8875">37°53'15"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="41.13222">41°7'56"E</geoCoordinate>
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||
], V. Pietschmann, 15 Jul 1914. - NMW 90805, 1, Turkey,
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||
<normalizedToken originalValue="Gökçesu">Goekcesu</normalizedToken>
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||
<normalizedToken originalValue="Çayı">Cayi</normalizedToken>
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||
(
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||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="37.75">37°45'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="41.75">41°45'E</geoCoordinate>
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), 26 Sep 1985. - ZMH 9548, 2, Turkey,
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<normalizedToken originalValue="Ceylanpınar">Ceylanpinar</normalizedToken>
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[
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="36.84722">36°50'50"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="40.05">40°3'0"E</geoCoordinate>
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]. - SMF 33119, 1, Syria, Nahr al
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<normalizedToken originalValue="Khābūr">Khabūr</normalizedToken>
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at Al
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<normalizedToken originalValue="Ḩasakah">Ḩasakah</normalizedToken>
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[
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="36.5025">36°30'9"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="40.74778">40°44'52"E</geoCoordinate>
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], F. Krupp. - SMF 30172, 1, Syria, Nahr al
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<normalizedToken originalValue="Khābūr">Khabūr</normalizedToken>
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near Tall
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<normalizedToken originalValue="Budayrī">Budayrī</normalizedToken>
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(
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="36.4">36°24'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="40.866665">40°52'E</geoCoordinate>
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), F. Krupp, 2-4 Nov 1986. - SMF 30173, 1, Syria, Nahr al
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<normalizedToken originalValue="Khābūr">Khabūr</normalizedToken>
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near
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<normalizedToken originalValue="Nahāb">Nahab</normalizedToken>
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(
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="36.383335">36°23'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="40.833332">40°50'E</geoCoordinate>
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), F. Krupp, 23-27 May 1989. - SMF 30174, 1, Syria, Nahr al
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<normalizedToken originalValue="Khābūr">Khabūr</normalizedToken>
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near
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<normalizedToken originalValue="Nahāb">Nahab</normalizedToken>
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(
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="36.383335">36°23'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="40.833332">40°50'E</geoCoordinate>
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), F. Krupp, 28
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<normalizedToken originalValue="Sep–">Sep-</normalizedToken>
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8 Oct 1988. - CMNFI 79-0290, 2, Iran,
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<normalizedToken originalValue="Qaşr-e">Qasr-e</normalizedToken>
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<normalizedToken originalValue="Shīrīn">Shīrīn</normalizedToken>
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(
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="34.516666">34°31'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="45.583332">45°35'E</geoCoordinate>
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). - CMNFI 79-0289, 1, Iran, 25-30 km from
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<normalizedToken originalValue="Qaşr-e">Qasr-e</normalizedToken>
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<normalizedToken originalValue="Shīrīn">Shīrīn</normalizedToken>
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(
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="34.466667">34°28'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="45.866665">45°52'E</geoCoordinate>
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). - BMNH 1974.2.22:1292-1296, 4; BMNH 1974.2.22:1281, 1, Iraq, Euphrates at
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<normalizedToken originalValue="Ḩadīthah">Ḩadīthah</normalizedToken>
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[
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||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="34.139725">34°8'23"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="42.378056">42°22'41"E</geoCoordinate>
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], 19 Oct 1953. - CMNFI 79-0275, 1, Iran,
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<normalizedToken originalValue="Rūdkhāneh-ye">Rūdkhaneh-ye</normalizedToken>
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<normalizedToken originalValue="Kashgān">Kashgan</normalizedToken>
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, 2 km from
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<normalizedToken originalValue="Ma‘mūlān">Ma'mūlan</normalizedToken>
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(
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="33.416668">33°25'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="47.966667">47°58'E</geoCoordinate>
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). - SMF 33129, 3, Iran,
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<normalizedToken originalValue="Rūdkhāneh-ye">Rūdkhaneh-ye</normalizedToken>
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Karkheh at Pol-e Dokhtar (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.16">33°9'36"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="47.72">47°43'12"E</geoCoordinate>
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), N. Alwan et al., 3 Mar 2008. - ZM-CBSU 4153, 1; ZM-CBSU 4154, 1, Iran,
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<normalizedToken originalValue="Rūdkhāneh-ye">Rūdkhaneh-ye</normalizedToken>
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||
Dez at
|
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<normalizedToken originalValue="Dezfūl">Dezfūl</normalizedToken>
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[
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="32.3825">32°22'57"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="48.401947">48°24'7"E</geoCoordinate>
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], F. Bossaghzadeh, 8 Jun 2005.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="diagnosis">
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<paragraph pageId="26" pageNumber="27">Diagnosis.</paragraph>
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<paragraph pageId="26" pageNumber="27">Two pairs of barbels; 32 to 38 scales in the lateral line, usually 14 to 16 scales around the least circumference of the caudal peduncle; last unbranched dorsal-fin ray markedly longer than head; mouth narrow, lower lip spatulate and median lobe present.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
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<paragraph pageId="26" pageNumber="27">Description.</paragraph>
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<paragraph lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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Body moderately high, laterally compressed and without a nuchal hump. The greatest body depth is at the point of the origin of the dorsal fin. The ventral profile of the head is straight, its dorsal profile has a slight to pronounced hump
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<pageBreakToken pageId="27" pageNumber="28" start="start">near</pageBreakToken>
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the nostrils (Figs 21, 22). The head is short and narrow. The mouth is inferior. The maximum body depth is bigger than the head length (Fig. 12). The lips are comparatively thick and the lower jaw is narrow with a sharp horny sheath and a median lobe. The two pairs of barbels (Table 2) are stout and the anterior pair is quite long. The eyes are rather high in the middle of the head and rather small. The morphometric characters are summarised in Table 1.
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</paragraph>
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<caption pageId="27" pageNumber="28">
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<paragraph pageId="27" pageNumber="28">
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Figure 21.
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
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, holotype (ZMH 1148) from Batman
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<normalizedToken originalValue="Çayı">Cayi</normalizedToken>
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.
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</paragraph>
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</caption>
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<caption pageId="27" pageNumber="28">
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<paragraph pageId="27" pageNumber="28">
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Figure 22.
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
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, live specimen from
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<normalizedToken originalValue="Rūdkhāneh-ye">Rūdkhaneh-ye</normalizedToken>
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Karkheh.
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</paragraph>
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</caption>
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<paragraph pageId="27" pageNumber="28">The dorsal fin is long and usually has four unbranched and nine or 10 branched rays (Table 3). The last unbranched ray is long and well ossified; only the tip is flexible. It is considerably longer than the head (Fig. 4). The anal fin usually has three unbranched rays and six branched rays (Table 4). Its base is long. The bases of the dorsal and anal fin have a sheath of scales.</paragraph>
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<paragraph pageId="27" pageNumber="28">There are 32 to 38 scales in the lateral line (Table 5), 5.5 to seven scales above the lateral line (Table 6), 4.5 to 6.5 scales below the lateral line (Table 7) and (12) 14 to 16 scales around the least circumference of the caudal peduncle (Table 8). The scales are shown in Fig. 5.</paragraph>
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<paragraph pageId="28" pageNumber="29">
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<pageBreakToken pageId="28" pageNumber="29" start="start">The</pageBreakToken>
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pharyngeal teeth count is 2.3.5-5.3.2 in seven specimens, 2.3.5- in one specimen and -4.3.2 in one specimen. The pharyngeal teeth are hooked at their tips (Fig. 6).
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</paragraph>
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<paragraph pageId="28" pageNumber="29">Live specimens are silvery. The back is darker than the belly, which is almost white (Fig. 22). Fixed specimens are yellow-brown and some have a darker back.</paragraph>
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<paragraph pageId="28" pageNumber="29">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
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differs from all congeners, except
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus sublimus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="sublimus">Carasobarbus sublimus</taxonomicName>
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, by having a spatulate lower jaw with a median lobe on the lower lip vs. a crescent-shaped lower jaw and a lower lip without median lobe. It differs from
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus sublimus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="sublimus">Carasobarbus sublimus</taxonomicName>
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by having 32 to 38 scales in the lateral line vs. 27 to 29 and modally 14 scales around the least circumference of the caudal peduncle vs. 12 and by having a longer and more ossified last unbranched ray in the dorsal fin.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="29" type="distribution">
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<paragraph pageId="28" pageNumber="29">Distribution.</paragraph>
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<paragraph pageId="28" pageNumber="29">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
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occurs in the Euphrates-Tigris system (Fig. 7).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="29" type="habitats and biology">
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<paragraph pageId="28" pageNumber="29">Habitats and biology.</paragraph>
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<paragraph pageId="28" pageNumber="29">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
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is rare, inhabits fast-flowing reaches of rivers and feeds on small animals (
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<bibRefCitation author="Krupp, F" editor="Kuehne, H" journalOrPublisher="Harrassowitz-Verlag, Wiesbaden" pageId="45" pageNumber="46" pagination="41 - 51" title="Die Fischfauna des Nahr al-Ḫabūr, Nordost-Syrien." year="2008">Krupp and Schneider 2008</bibRefCitation>
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). The maximum length is about 150 mm SL and this species has no economic importance (
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<bibRefCitation author="Krupp, F" editor="Kuehne, H" journalOrPublisher="Harrassowitz-Verlag, Wiesbaden" pageId="45" pageNumber="46" pagination="41 - 51" title="Die Fischfauna des Nahr al-Ḫabūr, Nordost-Syrien." year="2008">Krupp and Schneider 2008</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="29" type="conservation status">
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<paragraph pageId="28" pageNumber="29">Conservation status.</paragraph>
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<paragraph pageId="28" pageNumber="29">Little information is available, but because this species is dependent on fast-flowing water, it is probably impacted by the construction of dams.</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="29" type="discussion">
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<paragraph pageId="28" pageNumber="29">Discussion.</paragraph>
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<paragraph pageId="28" pageNumber="29">
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
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was described from the Batman
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<normalizedToken originalValue="Çayı">Cayi</normalizedToken>
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and placed in the genus
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Cyclocheilichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocheilichthys" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Cyclocheilichthys</taxonomicName>
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(
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<bibRefCitation author="Ladiges, W" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="45" pageNumber="46" pagination="105 - 150" title="Suesswasserfische der Tuerkei. I. Teil Cyprinidae." volume="58" year="1960">Ladiges 1960</bibRefCitation>
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). Karaman erected the new genus
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<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Kosswigobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kosswigobarbus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Kosswigobarbus</taxonomicName>
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for this species (
|
||
<bibRefCitation author="Karaman, MS" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="44" pageNumber="45" pagination="175 - 254" title="Suesswasserfische der Tuerkei. 8. Teil: Revision der Barben Europas, Vorderasiens und Nordafrikas." volume="67" year="1971">Karaman 1971</bibRefCitation>
|
||
). Coad gave a detailed re-description of this species and transferred it to the genus
|
||
<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Barbus</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Coad, BW" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="42" pageNumber="43" pagination="263 - 265" title="A re-description and generic re-assignment of Kosswigobarbus kosswigi (Ladiges, 1960), a cyprinid fish from Turkey and Iran." volume="79" year="1982">Coad 1982</bibRefCitation>
|
||
).
|
||
<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Kosswigobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kosswigobarbus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Kosswigobarbus</taxonomicName>
|
||
was revalidated (
|
||
<bibRefCitation author="Ekmekci, FG" journalOrPublisher="Folia Zoologica" pageId="43" pageNumber="44" pagination="87 - 96" title="A revision of the generic position of Barynotus (Systomus) verhoeffi, and the validity of the genera Carasobarbus, Kosswigobarbus and Mesopotamichthys (Pisces, Cyprinidae)." volume="47" year="1998">
|
||
<normalizedToken originalValue="Ekmekçi">Ekmekci</normalizedToken>
|
||
and Banarescu 1998
|
||
</bibRefCitation>
|
||
) and sometimes used as a subgenus of
|
||
<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Barbus</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Tsigenopoulos, CS" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="46" pageNumber="47" pagination="851 - 856" title="Phylogenetic relationships of hexaploid large-sized barbs (genus Labeobarbus, Cyprinidae) based on mtDNA data." url="10.1016/j.ympev.2010.02.006" volume="56" year="2010">Tsigenopoulos et al. 2010</bibRefCitation>
|
||
). Later the species was placed in
|
||
<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Carasobarbus</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Borkenhagen, K" journalOrPublisher="Environmental Biology of Fishes" pageId="42" pageNumber="43" pagination="327 - 335" title="The molecular systematics of the Carasobarbus species from Iran and adjacent areas, with comments on Carasobarbus albus (Heckel, 1843)." url="10.1007/s10641-011-9787-1" volume="91" year="2011">Borkenhagen et al. 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="28" pageNumber="29">
|
||
<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
|
||
is paraphyletic with respect to
|
||
<taxonomicName class="Actinopterygii" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus sublimus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="sublimus">Carasobarbus sublimus</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Borkenhagen, K" journalOrPublisher="Environmental Biology of Fishes" pageId="42" pageNumber="43" pagination="327 - 335" title="The molecular systematics of the Carasobarbus species from Iran and adjacent areas, with comments on Carasobarbus albus (Heckel, 1843)." url="10.1007/s10641-011-9787-1" volume="91" year="2011">Borkenhagen et al. 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |