207 lines
23 KiB
XML
207 lines
23 KiB
XML
<document id="A4BB748E63F47AA31635FDDD28425B59" ID-CLB-Dataset="7002" ID-DOI="10.11646/zootaxa.4965.2.9" ID-GBIF-Dataset="73b8e644-f46f-4526-9b4e-b323f2205c8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4751849" ID-ZooBank="C0F65518-6EA9-4099-AEA2-51C5EECFFC78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619710709549" checkinUser="plazi" docAuthor="Allsopp, Peter G." docDate="2021" docId="0C628795FF82FF8AFF1B11D2FC6CDBC4" docLanguage="en" docName="zootaxa.4965.2.9.pdf" docOrigin="Zootaxa 4965 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesystoechus costatus Carne 1958" docType="treatment" docVersion="7" lastPageNumber="368" masterDocId="F05BFFEDFF87FF8FFF8C1258FFB1DD73" masterDocTitle="The Australian endemic genera Mesystoechus Waterhouse, 1878, Amblochilus Blanchard, 1851, and Bilobatus Machatschke, 1970 revisited (Coleoptera Scarabaeidae: Rutelinae: Anoplognathini)" masterLastPageNumber="374" masterPageNumber="363" pageNumber="368" updateTime="1698938014406" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4DA15C6392A1AC0B1FBF778F1671221A">The Australian endemic genera Mesystoechus Waterhouse, 1878, Amblochilus Blanchard, 1851, and Bilobatus Machatschke, 1970 revisited (Coleoptera Scarabaeidae: Rutelinae: Anoplognathini)</mods:title>
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<mods:namePart id="F84253C1D033E1D42C5EDBF4E7D48C9B">Allsopp, Peter G.</mods:namePart>
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<mods:title id="07D6A97FC060CEFA7FB5BA2A96F2C416">Zootaxa</mods:title>
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<mods:part id="D4307C8B331F52796375C49A1E300471">
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<mods:date id="C0FDAE721573C914FA0A6309032063B8">2021</mods:date>
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<treatment id="0C628795FF82FF8AFF1B11D2FC6CDBC4" ID-DOI="http://doi.org/10.5281/zenodo.4752426" ID-GBIF-Taxon="181983392" ID-Zenodo-Dep="4752426" LSID="urn:lsid:plazi:treatment:0C628795FF82FF8AFF1B11D2FC6CDBC4" httpUri="http://treatment.plazi.org/id/0C628795FF82FF8AFF1B11D2FC6CDBC4" lastPageNumber="368" pageId="5" pageNumber="368">
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<heading id="DF3C81EFFF82FF8AFF1B11D2FDF6DED7" bold="true" box="[151,583,906,933]" fontSize="11" level="1" pageId="5" pageNumber="368" reason="1">
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<taxonomicName id="43CB4D00FF82FF8AFF1B11D2FDF6DED7" authority="Carne, 1958" authorityName="Carne" authorityYear="1958" box="[151,583,906,933]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="costatus">
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<emphasis id="B6BFEA91FF82FF8AFF1B11D2FDF6DED7" bold="true" box="[151,583,906,933]" pageId="5" pageNumber="368">
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<emphasis id="B6BFEA91FF82FF8AFF1B11D2FE19DED6" bold="true" box="[151,424,906,933]" italics="true" pageId="5" pageNumber="368">Mesystoechus costatus</emphasis>
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<bibRefCitation id="E05A4B72FF82FF8AFE2311D2FDF6DED7" author="Carne, P. B." box="[431,583,906,932]" pageId="5" pageNumber="368" pagination="162 - 240" refId="ref7741" refString="Carne, P. B. (1958) A review of the Australian Rutelinae (Coleoptera: Scarabaeidae). Australian Journal of Zoology, 6, 162 - 240. https: // doi. org / 10.1071 / ZO 9580162" type="journal article" year="1958">Carne, 1958</bibRefCitation>
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</emphasis>
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<taxonomicName id="43CB4D00FF82FF8AFF1B1197FD87DE94" authority="Carne, 1958: 216" authorityName="Carne" authorityPageNumber="216" authorityYear="1958" box="[151,566,975,999]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="costatus">
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<emphasis id="B6BFEA91FF82FF8AFF1B1197FECCDE94" box="[151,381,975,999]" italics="true" pageId="5" pageNumber="368">Mesystoechus costatus</emphasis>
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<bibRefCitation id="E05A4B72FF82FF8AFE081197FD87DE94" author="Carne, P. B." box="[388,566,975,999]" pageId="5" pageNumber="368" pagination="162 - 240" refId="ref7741" refString="Carne, P. B. (1958) A review of the Australian Rutelinae (Coleoptera: Scarabaeidae). Australian Journal of Zoology, 6, 162 - 240. https: // doi. org / 10.1071 / ZO 9580162" type="journal article" year="1958">Carne, 1958: 216</bibRefCitation>
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.
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<emphasis id="B6BFEA91FF82FF8AFF1B164DFE92D95C" bold="true" box="[151,291,1045,1071]" pageId="5" pageNumber="368">Type series.</emphasis>
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<specimenCount id="92CDFD0AFF82FF8AFE15164EFE00D943" box="[409,433,1046,1072]" pageId="5" pageNumber="368" type="male">♂</specimenCount>
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:
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<collectingCountry id="FCDC7613FF82FF8AFE37164DFDECD95C" box="[443,605,1045,1071]" name="Australia" pageId="5" pageNumber="368">AUSTRALIA</collectingCountry>
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.
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<collectingRegion id="460FF861FF82FF8AFDE4164DFC98D95C" box="[616,809,1045,1071]" country="Australia" name="Queensland" pageId="5" pageNumber="368">QUEENSLAND</collectingRegion>
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:
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<location id="81146058FF82FF8AFCBE164DFC30D95C" LSID="urn:lsid:plazi:treatment:0C628795FF82FF8AFF1B11D2FC6CDBC4:81146058FF82FF8AFCBE164DFC30D95C" box="[818,897,1045,1071]" country="Australia" name="Too'ba" pageId="5" pageNumber="368" stateProvince="Queensland">Too’ba</location>
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[Toowoomba], 3.i.[19]22 [
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<collectionCode id="E2DAAE46FF82FF8AFB3D164DFB57D943" box="[1201,1254,1045,1072]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="5" pageNumber="368" type="Museum">QM</collectionCode>
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].
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<paragraph id="84743683FF82FF8AFF1B1662FF5DD90B" blockId="5.[151,1437,1045,1719]" pageId="5" pageNumber="368">
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<typeStatus id="5B708821FF82FF8AFF1B1662FEB9D927" box="[151,264,1082,1108]" pageId="5" pageNumber="368" type="paratype">Paratypes</typeStatus>
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:
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<collectingCountry id="FCDC7613FF82FF8AFE981661FE07D920" box="[276,438,1081,1107]" name="Australia" pageId="5" pageNumber="368">AUSTRALIA</collectingCountry>
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.
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:
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<specimenCount id="92CDFD0AFF82FF8AFD1E1661FD06D920" box="[658,695,1081,1107]" pageId="5" pageNumber="368" type="male">3♂</specimenCount>
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, same data as holotype, one with
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<location id="81146058FF82FF8AFBC91661FB60D920" LSID="urn:lsid:plazi:treatment:0C628795FF82FF8AFF1B11D2FC6CDBC4:81146058FF82FF8AFBC91661FB60D920" box="[1093,1233,1081,1107]" country="Australia" name="Toowoomba" pageId="5" pageNumber="368" stateProvince="Queensland">Toowoomba</location>
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spelt out [
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<collectionCode id="E2DAAE46FF82FF8AFAC21661FA29D920" box="[1358,1432,1081,1107]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="5" pageNumber="368">ANIC</collectionCode>
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,
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<collectionCode id="E2DAAE46FF82FF8AFF1B1605FF53D90B" box="[151,226,1117,1144]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34515" lsid="urn:lsid:biocol.org:col:34515" name="University of Queensland Insect Collection" pageId="5" pageNumber="368">UQIC</collectionCode>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="84743683FF82FF8AFF4B16D9FC36D997" blockId="5.[151,1437,1045,1719]" pageId="5" pageNumber="368">
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<emphasis id="B6BFEA91FF82FF8AFF4B16D9FE4CD9E8" bold="true" box="[199,509,1153,1179]" pageId="5" pageNumber="368">Other material examined.</emphasis>
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<materialsCitation id="34A33CDEFF82FF8AFD8916D9FD33D9B3" ID-GBIF-Occurrence="3110759323" collectingDate="1970-02-16" collectingDateMax="1970-02-20" collectingDateMin="1970-02-16" collectionCode="NHML, UQIC" collectorName="G. B. Monteith" country="Australia" location="Emu Vale" municipality="Bald Mtn. area" pageId="5" pageNumber="368" specimenCount="2" specimenCount-male="2" stateProvince="Queensland">
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<collectingCountry id="FCDC7613FF82FF8AFD8916D9FD16D9E8" box="[517,679,1153,1179]" name="Australia" pageId="5" pageNumber="368">AUSTRALIA</collectingCountry>
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.
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<collectingRegion id="460FF861FF82FF8AFD3F16D9FCC5D9E8" box="[691,884,1153,1179]" country="Australia" name="Queensland" pageId="5" pageNumber="368">QUEENSLAND</collectingRegion>
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:
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<specimenCount id="92CDFD0AFF82FF8AFCF316D9FC15D9E8" box="[895,932,1153,1179]" pageId="5" pageNumber="368" type="male">2♂</specimenCount>
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,
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<collectingMunicipality id="6410ACF9FF82FF8AFC3C16D9FBEBD9EF" box="[944,1114,1153,1180]" pageId="5" pageNumber="368">Bald Mtn. area</collectingMunicipality>
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3-4,000’, via
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<location id="81146058FF82FF8AFB7616DAFADAD9E8" LSID="urn:lsid:plazi:treatment:0C628795FF82FF8AFF1B11D2FC6CDBC4:81146058FF82FF8AFB7616DAFADAD9E8" box="[1274,1387,1153,1180]" country="Australia" municipality="Bald Mtn. area" name="Emu Vale" pageId="5" pageNumber="368" stateProvince="Queensland">Emu Vale</location>
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,
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<date id="F0751043FF82FF8AFAFB16D9FEBAD9CC" pageId="5" pageNumber="368" value="1970-02-16" valueMax="1970-02-20" valueMin="1970-02-16">
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<collectingDate id="E031E9ABFF82FF8AFAFB16D9FEBAD9CC" pageId="5" pageNumber="368" value="1970-02-16" valueMax="1970-02-20" valueMin="1970-02-16">16- 20.ii.1970</collectingDate>
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,
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<collectorName id="293E5355FF82FF8AFE9416FDFE0CD9CC" box="[280,445,1189,1215]" pageId="5" pageNumber="368">G.B. Monteith</collectorName>
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[
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<collectionCode id="E2DAAE46FF82FF8AFE4716FEFD92D9B3" box="[459,547,1190,1216]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="5" pageNumber="368">NHML</collectionCode>
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,
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<collectionCode id="E2DAAE46FF82FF8AFDBC16FDFDCAD9B3" box="[560,635,1189,1216]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34515" lsid="urn:lsid:biocol.org:col:34515" name="University of Queensland Insect Collection" pageId="5" pageNumber="368">UQIC</collectionCode>
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]
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</materialsCitation>
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;
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<materialsCitation id="34A33CDEFF82FF8AFD0216FDFAA5D9B3" ID-GBIF-Occurrence="3110759332" box="[654,1300,1189,1216]" collectingDate="1971-01-17" collectionCode="UQIC" collectorName="G. Monteith" country="Australia" location="Bald Mt." pageId="5" pageNumber="368" specimenCount="1" specimenCount-female="1" stateProvince="Queensland">
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<specimenCount id="92CDFD0AFF82FF8AFD0216FDFD00D9CC" box="[654,689,1189,1215]" pageId="5" pageNumber="368" type="female">1♀</specimenCount>
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,
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<location id="81146058FF82FF8AFD4D16FDFC96D9B3" LSID="urn:lsid:plazi:treatment:0C628795FF82FF8AFF1B11D2FC6CDBC4:81146058FF82FF8AFD4D16FDFC96D9B3" box="[705,807,1189,1216]" country="Australia" name="Bald Mt." pageId="5" pageNumber="368" stateProvince="Queensland">Bald Mt.</location>
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Emu Vale,
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<date id="F0751043FF82FF8AFC2116FDFBA9D9CC" box="[941,1048,1189,1215]" pageId="5" pageNumber="368" value="1971-01-17">
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<collectingDate id="E031E9ABFF82FF8AFC2116FDFBA9D9CC" box="[941,1048,1189,1215]" pageId="5" pageNumber="368" value="1971-01-17">17.i.1971</collectingDate>
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</date>
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,
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<collectorName id="293E5355FF82FF8AFBAA16FDFB00D9CC" box="[1062,1201,1189,1215]" pageId="5" pageNumber="368">G. Monteith</collectorName>
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[
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<collectionCode id="E2DAAE46FF82FF8AFB4C16FDFABCD9B3" box="[1216,1293,1189,1216]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34515" lsid="urn:lsid:biocol.org:col:34515" name="University of Queensland Insect Collection" pageId="5" pageNumber="368">UQIC</collectionCode>
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]
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</materialsCitation>
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;
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<materialsCitation id="34A33CDEFF82FF8AFAAD16FDFDB4D997" ID-GBIF-Occurrence="3110759309" collectingDate="1926-01" collectionCode="QM" collectorName="E. Dumigan" country="Australia" location="Bunya Mtns" pageId="5" pageNumber="368" specimenCount="1" specimenCount-male="1" stateProvince="Queensland">
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<specimenCount id="92CDFD0AFF82FF8AFAAD16FDFAF4D9CC" box="[1313,1349,1189,1215]" pageId="5" pageNumber="368" type="male">1♂</specimenCount>
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,
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<location id="81146058FF82FF8AFADF16FEFF62D997" LSID="urn:lsid:plazi:treatment:0C628795FF82FF8AFF1B11D2FC6CDBC4:81146058FF82FF8AFADF16FEFF62D997" country="Australia" name="Bunya Mtns" pageId="5" pageNumber="368" stateProvince="Queensland">Bunya Mtns</location>
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,
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<date id="F0751043FF82FF8AFF531691FE99D990" box="[223,296,1225,1251]" pageId="5" pageNumber="368" value="1926-01">
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<collectingDate id="E031E9ABFF82FF8AFF531691FE99D990" box="[223,296,1225,1251]" pageId="5" pageNumber="368" value="1926-01">i.1926</collectingDate>
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</date>
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,
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<collectorName id="293E5355FF82FF8AFEB81692FE0AD990" box="[308,443,1225,1252]" pageId="5" pageNumber="368">E. Dumigan</collectorName>
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[
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;
|
||
<materialsCitation id="34A33CDEFF82FF8AFD9C1691FC32D997" ID-GBIF-Occurrence="3110759324" box="[528,899,1225,1252]" collectionCode="QM" country="Australia" location="Bunya Mts" pageId="5" pageNumber="368" specimenCount="1" specimenCount-male="1" stateProvince="Queensland">
|
||
<specimenCount id="92CDFD0AFF82FF8AFD9C1691FD84D990" box="[528,565,1225,1251]" pageId="5" pageNumber="368" type="male">1♂</specimenCount>
|
||
,
|
||
<location id="81146058FF82FF8AFDCD1692FD71D997" LSID="urn:lsid:plazi:treatment:0C628795FF82FF8AFF1B11D2FC6CDBC4:81146058FF82FF8AFDCD1692FD71D997" box="[577,704,1226,1252]" country="Australia" name="Bunya Mts" pageId="5" pageNumber="368" stateProvince="Queensland">Bunya Mts</location>
|
||
, 7.i.[19]26 [
|
||
<collectionCode id="E2DAAE46FF82FF8AFCC41691FCCCD997" box="[840,893,1225,1252]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="5" pageNumber="368" type="Museum">QM</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CCD16508FF82FF8AFF4B16B5FD78DB38" pageId="5" pageNumber="368" type="diagnosis">
|
||
<paragraph id="84743683FF82FF8AFF4B16B5FC19D8AC" blockId="5.[151,1437,1045,1719]" pageId="5" pageNumber="368">
|
||
<emphasis id="B6BFEA91FF82FF8AFF4B16B5FDABD874" bold="true" box="[199,538,1261,1287]" pageId="5" pageNumber="368">
|
||
Diagnosis. Male (
|
||
<figureCitation id="1CF02A06FF82FF8AFE1E16B5FDBCD874" box="[402,525,1261,1287]" captionStart="FIGURES 7–15" captionStartId="6.[151,264,1869,1893]" captionTargetBox="[198,1389,181,1845]" captionTargetId="figure-20@6.[198,1389,181,1845]" captionTargetPageId="6" captionText="FIGURES 7–15. Mesystoechus species. 7–8, Mesystoechus ciliatus Waterhouse, 1878, male: 7, dorsal view; 8, lateral view; 9–10, Mesystoechus costatus Carne, 1958, male: 9, dorsal view; 10, lateral view; 11–14, Mesystoechus lithgowae, new species, male holotype: 11, dorsal view; 12, lateral view; 13, parameres, frontal view; 14, aedeagus, lateral view; 15, known distribution of Mesystoechus spp. in southeastern Queensland and northeastern New South Wales: yellow squares—M. ciliatus; blue triangles—M. costatus; red star—M. lithgowae. Figs. 7–14 photographed by Greg Daniels." figureDoi="http://doi.org/10.5281/zenodo.4751863" httpUri="https://zenodo.org/record/4751863/files/figure.png" pageId="5" pageNumber="368">Figs. 9–10</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Body
|
||
<quantity id="43339B66FF82FF8AFDEF16B5FD6AD87B" box="[611,731,1261,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="5" pageNumber="368" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
|
||
long; head, pronotum, and scutellum black with a green or coppery-green sheen; elytra with the striae pale yellowish brown, costate intervals deep brown; venter and pygidium reddish brown with white setae; legs and prescutellum with yellow setae. Mentum with anterior margin slightly concave. Anterior margin of upper section of clypeus weakly recurved. Anterior margin of pronotum with a few setae near angles. Elytral intervals smooth and strongly costate. Larger claws simple, without a minute tooth. Description by
|
||
<bibRefCitation id="E05A4B72FF82FF8AFF3617F9FE30D8C8" author="Carne, P. B." box="[186,385,1441,1468]" pageId="5" pageNumber="368" pagination="162 - 240" refId="ref7741" refString="Carne, P. B. (1958) A review of the Australian Rutelinae (Coleoptera: Scarabaeidae). Australian Journal of Zoology, 6, 162 - 240. https: // doi. org / 10.1071 / ZO 9580162" type="journal article" year="1958">Carne (1958: 216</bibRefCitation>
|
||
; head of male Fig. 120; right maxilla of male Fig. 116; mentum of male Fig. 117; mandible of male Fig. 118; protibia of male Fig. 119; parameres of male Fig. 121).
|
||
</paragraph>
|
||
<paragraph id="84743683FF82FF8AFF4B17B1FD78DB38" blockId="5.[151,1437,1045,1719]" pageId="5" pageNumber="368">
|
||
<emphasis id="B6BFEA91FF82FF8AFF4B17B1FE94DB70" bold="true" box="[199,293,1513,1539]" pageId="5" pageNumber="368">Female.</emphasis>
|
||
Body
|
||
<quantity id="43339B66FF82FF8AFEF817B1FE6BDB77" box="[372,474,1513,1540]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="5" pageNumber="368" unit="mm" value="14.5">14.5 mm</quantity>
|
||
long; head dark brown to black, pronotum dark brown, scutellum lighter brown, all without a green sheen. Antennal club shorter than shaft. Upper surface of clypeus with median swelling. Tarsi reduced, shorter than in males. Otherwise like male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CCD16508FF82FF8AFF4B140DFC6CDBC4" pageId="5" pageNumber="368" type="distribution">
|
||
<paragraph id="84743683FF82FF8AFF4B140DFB82DBE0" blockId="5.[151,1437,1045,1719]" pageId="5" pageNumber="368">
|
||
<emphasis id="B6BFEA91FF82FF8AFF4B140DFE7CDB1C" bold="true" box="[199,461,1621,1647]" pageId="5" pageNumber="368">
|
||
Distribution (
|
||
<figureCitation id="1CF02A06FF82FF8AFEE5140DFE0FDB1C" box="[361,446,1621,1647]" captionStart="FIGURES 7–15" captionStartId="6.[151,264,1869,1893]" captionTargetBox="[198,1389,181,1845]" captionTargetId="figure-20@6.[198,1389,181,1845]" captionTargetPageId="6" captionText="FIGURES 7–15. Mesystoechus species. 7–8, Mesystoechus ciliatus Waterhouse, 1878, male: 7, dorsal view; 8, lateral view; 9–10, Mesystoechus costatus Carne, 1958, male: 9, dorsal view; 10, lateral view; 11–14, Mesystoechus lithgowae, new species, male holotype: 11, dorsal view; 12, lateral view; 13, parameres, frontal view; 14, aedeagus, lateral view; 15, known distribution of Mesystoechus spp. in southeastern Queensland and northeastern New South Wales: yellow squares—M. ciliatus; blue triangles—M. costatus; red star—M. lithgowae. Figs. 7–14 photographed by Greg Daniels." figureDoi="http://doi.org/10.5281/zenodo.4751863" httpUri="https://zenodo.org/record/4751863/files/figure.png" pageId="5" pageNumber="368">Fig. 15</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Known only from east of Warwick, through Toowoomba and to the Bunya Mountains in southeastern
|
||
<collectingRegion id="460FF861FF82FF8AFECB1421FE7EDBE0" box="[327,463,1657,1683]" country="Australia" name="Queensland" pageId="5" pageNumber="368">Queensland</collectingRegion>
|
||
. All areas are elevated and have deep kraznosem soils.
|
||
</paragraph>
|
||
<paragraph id="84743683FF82FF8AFF4B14C5FC6CDBC4" blockId="5.[151,1437,1045,1719]" box="[199,989,1693,1719]" pageId="5" pageNumber="368">
|
||
<emphasis id="B6BFEA91FF82FF8AFF4B14C5FE34DBC4" bold="true" box="[199,389,1693,1719]" pageId="5" pageNumber="368">Natural history.</emphasis>
|
||
Adults have been collected during January-February.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |