treatments-xml/data/06/77/7E/06777E90C84D5B858F5AA98785A5D09A.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2022" docId="06777E90C84D5B858F5AA98785A5D09A" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina exclamationis Mabille 1887" docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-25</mods:number>
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<mods:detail type="volume">
<mods:number>1119</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>115</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<treatment ID-GBIF-Taxon="199839032" LSID="urn:lsid:plazi:treatment:06777E90C84D5B858F5AA98785A5D09A" httpUri="http://treatment.plazi.org/id/06777E90C84D5B858F5AA98785A5D09A" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="06777E90-C84D-5B85-8F5A-A98785A5D09A" authority="Mabille, 1887" authorityName="Mabille" authorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina exclamationis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="exclamationis">Pupina exclamationis Mabille, 1887</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 &quot; forma minor &quot;. Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J)." figureDoi="10.3897/zookeys.1119.85400.figure29" httpUri="https://binary.pensoft.net/fig/735390" pageId="0" pageNumber="1">Figs 29I-K</figureCitation>
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 &quot; forma minor &quot; from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B)." figureDoi="10.3897/zookeys.1119.85400.figure34" httpUri="https://binary.pensoft.net/fig/735395" pageId="0" pageNumber="1">, 34B, C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Mabille" authorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina exclamationis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="exclamationis">Pupina exclamationis</taxonomicName>
Mabille, 1887: 137, 138, pl. 4, figs 11, 12. Type locality: Tonkin.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.14809" author="Fischer, P" journalOrPublisher="Imprimerie Dejussieu Pere et Fils, Autun" pageId="0" pageNumber="1" refId="B41" refString="Fischer, P, 1891. Catalogue et distribution geographique des Mollusques terrestres, fluviatilies &amp; marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Imprimerie Dejussieu Pere et Fils, Autun, DOI: https://doi.org/10.5962/bhl.title.14809" title="Catalogue et distribution geographique des Mollusques terrestres, fluviatilies &amp; marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." url="https://doi.org/10.5962/bhl.title.14809" year="1891">Fischer 1891</bibRefCitation>
: 108.
<bibRefCitation author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
: 431, Bac-Kan, Tonkin; Monts Mauson, Tonkin [Mount Mau Son, Lang Son Province, Vietnam].
<bibRefCitation author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. 2015</bibRefCitation>
: 126, fig. 6a, Son La Province, Vietnam.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Mabille" authorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) exclamationis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="exclamationis" subGenus="Tylotoechus">Pupina (Tylotoechus) exclamationis</taxonomicName>
-
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
: 312.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
MNHN-IM-2000-35840 (4 shells; Figs
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 &quot; forma minor &quot;. Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J)." figureDoi="10.3897/zookeys.1119.85400.figure29" httpUri="https://binary.pensoft.net/fig/735390" pageId="0" pageNumber="1">29I, J</figureCitation>
,
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 &quot; forma minor &quot; from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B)." figureDoi="10.3897/zookeys.1119.85400.figure34" httpUri="https://binary.pensoft.net/fig/735395" pageId="0" pageNumber="1">34B</figureCitation>
) from Tonkin.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
NHMUK 1901.12.23.205-210 &quot;forma minor&quot; ex. H. Fruhstorfer coll. (5 shells; Figs
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 &quot; forma minor &quot;. Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J)." figureDoi="10.3897/zookeys.1119.85400.figure29" httpUri="https://binary.pensoft.net/fig/735390" pageId="0" pageNumber="1">29K</figureCitation>
,
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 &quot; forma minor &quot; from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B)." figureDoi="10.3897/zookeys.1119.85400.figure34" httpUri="https://binary.pensoft.net/fig/735395" pageId="0" pageNumber="1">34C</figureCitation>
) from Than-Moi,
<collectingCountry name="Vietnam">Tonkin</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell ovate-fusiform to fusiform; last whorl ca. three quarters of shell height. Apertural lip somewhat thickened but not expanded; apertural lip slightly curved when observed from lateral view. Columellar tooth fin-shaped.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Mabille" authorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina exclamationis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="exclamationis">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina exclamationis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. peguensis" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="peguensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. peguensis</emphasis>
</taxonomicName>
in having a glossy surface and a curved apertural lip when observed from lateral view, but differs in having a more fusiform shell shape and a less distinct parietal callus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Northern Vietnam (
<bibRefCitation author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>