treatments-xml/data/03/DA/CA/03DACA42450EE578FF3AFDE8FBD1C2C1.xml

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<mods:title id="047B0165AA74E5CD7CB82443A77D4BEF">A new species of Rhabdops Boulenger, 1893 (Serpentes: Natricinae) from the northern Western Ghats region of India</mods:title>
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<paragraph id="8BCC7B54450EE56EFF3AFDE8FC5BC226" blockId="4.[151,976,583,679]" box="[151,928,583,610]" pageId="4" pageNumber="31">
<heading id="D084CC38450EE56EFF3AFDE8FC5BC226" bold="true" box="[151,928,583,610]" fontSize="11" level="1" pageId="4" pageNumber="31" reason="1">
<emphasis id="B907A746450EE56EFF3AFDE8FC5BC226" bold="true" box="[151,928,583,610]" pageId="4" pageNumber="31">
<taxonomicName id="4C7300D7450EE56EFF3AFDE8FE77C225" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[151,396,583,609]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="31" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A746450EE56EFF3AFDE8FE77C225" bold="true" box="[151,396,583,609]" italics="true" pageId="4" pageNumber="31">Rhabdops aquaticus</emphasis>
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<taxonomicNameLabel id="A2341A3D450EE56EFE3EFDE7FE0AC226" box="[403,497,584,610]" pageId="4" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
Giri, Deepak, Captain and Gower
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</paragraph>
<paragraph id="8BCC7B54450EE56EFF3AFDC5FD9CC2E3" blockId="4.[151,976,583,679]" pageId="4" pageNumber="31">
urn:lsid:zoobank.org:act:C6DF899F-9513-4C14-B3B8-2EBBEF5EA62B (
<figureCitation id="134867D1450EE56EFF32FD21FEF0C2E3" box="[159,267,654,679]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1468,1490]" captionStartId-1="3.[151,250,1917,1939]" captionStartId-2="6.[151,250,1983,2005]" captionStartId-3="7.[151,250,1970,1992]" captionStartId-4="8.[151,250,1088,1110]" captionStartId-5="10.[151,250,1970,1992]" captionTargetBox-0="[182,1403,275,1428]" captionTargetBox-1="[359,1225,236,1841]" captionTargetBox-2="[198,1364,220,1894]" captionTargetBox-3="[205,1382,224,1914]" captionTargetBox-4="[157,1430,161,1062]" captionTargetBox-5="[174,1413,194,1947]" captionTargetId-0="figure@2.[151,1436,253,1446]" captionTargetId-1="figure@3.[312,1276,193,1878]" captionTargetId-2="figure@6.[168,1393,193,1927]" captionTargetId-3="figure@7.[173,1414,193,1949]" captionTargetId-4="figure@8.[151,1436,157,1066]" captionTargetId-5="figure@10.[173,1414,193,1949]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="10" captionText-0="FIGURE 1. Map showing the distribution of the genus Rhabdops in the Western Ghats region of peninsular India. Circles denote records of Rhabdops olivaceus and squares denote records of Rhabdops aquaticus sp. nov. Green symbols indicates vouchered records and white symbols indicate non-vouchered reports. Arrow indicates the type locality of the respective species. See Appendix 3 for locality details." captionText-1="FIGURE 2. (a) Maximum likelihood tree showing phylogenetic relationships of the genus Rhabdops and other natricine snakes. Bootstrap support shown at each internal branch. (b) Bayesian tree showing phylogenetic relationships of the genus Rhabdops and other natricine snakes. Posterior probability shown at each internal branch." captionText-2="FIGURE 3. Holotype of Rhabdops aquaticus sp. nov. (NCBS-AU 163) whole preserved specimen in A) dorsal and B) ventral views. Scale bar 10 mm." captionText-3="FIGURE 4. Holotype of Rhabdops aquaticus sp. nov. (NCBS-AU 163) head of preserved specimen in A) dorsal, B) ventral, C) right lateral and D) left lateral views. Scale bar 10 mm." captionText-4="FIGURE 5. Paratypes of Rhabdops aquaticus sp. nov. (specimen numbers given in first row). Heads of preserved specimen in A) dorsal, B) ventral, C) right lateral and D) left lateral views. Scale bar 10 mm." captionText-5="FIGURE 6. Rhabdops aquaticus sp. nov. showing colouration in life of: A) larger (presumably adult) specimen from Baraki, and B) smaller (possibly juvenile) specimen from Amboli. Specimens not collected." httpUri-0="https://zenodo.org/record/1051967/files/figure.png" httpUri-1="https://zenodo.org/record/1051969/files/figure.png" httpUri-2="https://zenodo.org/record/1051971/files/figure.png" httpUri-3="https://zenodo.org/record/1051973/files/figure.png" httpUri-4="https://zenodo.org/record/1051975/files/figure.png" httpUri-5="https://zenodo.org/record/1051977/files/figure.png" pageId="4" pageNumber="31">Figs. 1–6</figureCitation>
, 
<figureCitation id="134867D1450EE56EFEBBFD20FEDFC2E3" box="[278,292,655,679]" captionStart="FIGURE 9" captionStartId="15.[151,250,1970,1992]" captionTargetBox="[174,1413,194,1948]" captionTargetId="figure@15.[173,1414,193,1949]" captionTargetPageId="15" captionText="FIGURE 9. Habitat of Rhabdops aquaticus sp. nov. during the monsoon. A) Stream in Koyna showing (right of image) more lentic water where snakes were seen. B) Summit plateau at Baraki showing rocks and ephemeral pools." httpUri="https://zenodo.org/record/1051983/files/figure.png" pageId="4" pageNumber="31">9</figureCitation>
–10; 
<tableCitation id="C6F14EEF450EE56EFEF0FD21FE48C2E3" box="[349,435,654,679]" captionStart="TABLE 2" captionStartId="5.[125,202,585,606]" captionText="TABLE 2. Meristic anđ morphometric (in millimetres) đata for examineđ specimens of Rhabdops aquaticus sp. nov. All specimens are paratypes except for the holotype (*) anđ a referređ specimen (**). Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because" pageId="4" pageNumber="31">Table 2</tableCitation>
; Appendix 1–3)
</paragraph>
</subSubSection>
<subSubSection id="C36928DF450EE56EFF3AFD79FBB4C36E" pageId="4" pageNumber="31" type="reference_group">
<paragraph id="8BCC7B54450EE56EFF3AFD79FBB4C36E" blockId="4.[151,1436,726,810]" pageId="4" pageNumber="31">
<taxonomicName id="4C7300D7450EE56EFF3AFD79FDE4C2A8" authority="(Beddome, 1863)" authorityName="Beddome" authorityYear="1863" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[151,543,726,748]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="31" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A746450EE56EFF3AFD79FE98C2A8" box="[151,355,726,748]" italics="true" pageId="4" pageNumber="31">Rhabdops olivaceus</emphasis>
(
<bibRefCitation id="EFE206A5450EE56EFEDDFD79FDECC2A8" author="Beddome" box="[368,535,726,748]" pageId="4" pageNumber="31" refString="Beddome, R. H. (1863) Further notes upon the snakes of the Madras Presidency; with descriptions of new species. Madras Quarterly Journal of Medical Science, 6, 41 - 48." type="journal article" year="1863">Beddome, 1863</bibRefCitation>
)
</taxonomicName>
: 
<bibRefCitation id="EFE206A5450EE56EFD84FD79FCBDC2A8" author="Soman" box="[553,838,726,748]" pageId="4" pageNumber="31" refString="Soman, P. W. (1962) Extension of range of olivaceous smooth snake [Rhabdops olivaceus (Beddome)]. Journal of Bombay Natural History Society, 59, 966." type="journal article" year="1962">Soman (1962: 966, in part)</bibRefCitation>
, 
<bibRefCitation id="EFE206A5450EE56EFCFCFD79FAD9C2A8" author="Whitaker" box="[849,1314,726,748]" pageId="4" pageNumber="31" refString="Whitaker, R. &amp; Captain, A. (2004) Snakes of India: The Field Guide. Draco Books, Chennai, 479 pp." type="book" year="2004">Whitaker &amp; Captain (2004: 264–267, in part)</bibRefCitation>
, 
<bibRefCitation id="EFE206A5450EE56EFA80FD79FE23C34F" author="Chikane" pageId="4" pageNumber="31" refString="Chikane, S. &amp; Bhosale, H. (2012) Reptiles of Kaas, Northern Western Ghats, Maharashtra, India, with comments on their habitat and abundance. Sauria, 34, 3 - 15." type="journal article" year="2012">Chikane &amp; Bhosale (2012: 14, in part)</bibRefCitation>
, 
<bibRefCitation id="EFE206A5450EE56EFE4FFD5AFD51C34F" author="Ganesh" box="[482,682,757,779]" pageId="4" pageNumber="31" refString="Ganesh, S. R., Chandramouli, S. R. &amp; Gowrishankar, P. (2012) Record lengths of two endemic caenophidian snakes from the Western Ghats Mountains, India. Hamadryad, 36, 44 - 46." type="journal article" year="2012">Ganesh et al. (2012</bibRefCitation>
:45, in part), 
<bibRefCitation id="EFE206A5450EE56EFC85FD5AFBEDC34F" author="Srinivasulu" box="[808,1046,757,779]" pageId="4" pageNumber="31" refString="Srinivasulu, B., Srinivasulu, C., Das, A., Achyuthan, N. S. &amp; Kulkarni, N. U. (2013) Rhabdops olivaceus. The IUCN Red List of Threatened Species 2013: e. T 172649 A 1358911." type="journal article" year="2013">Srinivasulu et al. (2013</bibRefCitation>
, in part), 
<bibRefCitation id="EFE206A5450EE56EFBDBFD5AFEBDC36E" author="Bhosale" pageId="4" pageNumber="31" refString="Bhosale, H. &amp; Joshi, D. (2014) Notes on distribution, natural history and habitat use of a Colubridae snake, Rhabdops olivaceus (Beddome, 1863). Russian Journal of Herpetology, 21, 166 - 168." type="journal article" year="2014">Bhosale &amp; Joshi (2014: 166– 168, in part)</bibRefCitation>
, 
<bibRefCitation id="EFE206A5450EE56EFEFDFCBBFDBBC36E" author="Srinivasulu" box="[336,576,788,810]" pageId="4" pageNumber="31" refString="Srinivasulu, C., Srinivasulu, B. &amp; Molur, S. (2014) The status and distribution of reptiles in the Western Ghats, India. Conservation Assessment and Management Plan (CAMP). Wildlife Information Liaison Development Society, Coimbatore, Tamil Nadu, 148 pp." type="book" year="2014">Srinivasulu et al. (2014</bibRefCitation>
: 18, 29, 55–56, in part), 
<bibRefCitation id="EFE206A5450EE56EFC95FCBBFBB4C36E" author="Murthy" box="[824,1103,788,810]" pageId="4" pageNumber="31" refString="Murthy, T. S. N. (2016) A pictorial handbook on some lizards and snakes of Western Ghats. Zoological Survey of India, Kolkata, India, 132 pp." type="book" year="2016">Murthy (2016: 100, in part)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36928DF450EE56EFF3AFCFEFA94C392" pageId="4" pageNumber="31" type="diagnosis">
<paragraph id="8BCC7B54450EE56EFF3AFCFEFA94C392" blockId="4.[151,1437,849,1990]" pageId="4" pageNumber="31">
<emphasis id="B907A746450EE56EFF3AFCFEFEEAC32E" bold="true" box="[151,273,849,874]" pageId="4" pageNumber="31">Diagnosis.</emphasis>
A 
<taxonomicName id="4C7300D7450EE56EFE95FCFEFE53C32E" authorityName="Boulenger" authorityYear="1893" box="[312,424,849,874]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="B907A746450EE56EFE95FCFEFE53C32E" box="[312,424,849,874]" italics="true" pageId="4" pageNumber="31">Rhabdops</emphasis>
</taxonomicName>
with paired internasals and prefrontals, a mostly pale (yellow or whitish) venter with a dark, clearly demarcated irregular midventral stripe and more than 222 ventral scales. Beyond number of ventrals, pattern and colour, 
<taxonomicName id="4C7300D7450EE56EFED3FC36FD9DC3F5" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[382,614,921,946]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="31" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A746450EE56EFED3FC36FD9DC3F5" box="[382,614,921,946]" italics="true" pageId="4" pageNumber="31">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D450EE56EFDDFFC36FD2BC3F6" box="[626,720,921,946]" pageId="4" pageNumber="31" rank="species">
<emphasis id="B907A746450EE56EFDDFFC36FD2BC3F6" bold="true" box="[626,720,921,946]" pageId="4" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from its most similar congener, 
<taxonomicName id="4C7300D7450EE56EFB00FC35FAC0C3F6" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[1197,1339,921,946]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="31" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A746450EE56EFB00FC35FAC0C3F6" box="[1197,1339,921,946]" italics="true" pageId="4" pageNumber="31">R. olivaceus</emphasis>
</taxonomicName>
, also in nuclear and mitochondrial DNA sequences. See below for a detailed account of colour and pattern differences.
</paragraph>
</subSubSection>
<subSubSection id="C36928DF450EE56EFF6AFC4EFAEAC4B2" pageId="4" pageNumber="31" type="materials_examined">
<paragraph id="8BCC7B54450EE56EFF6AFC4EFA66C45A" blockId="4.[151,1437,849,1990]" pageId="4" pageNumber="31">
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<emphasis id="B907A746450EE56EFF6AFC4EFEC0C3BE" bold="true" box="[199,315,993,1018]" pageId="4" pageNumber="31">
<typeStatus id="54C8C5F6450EE56EFF6AFC4EFECDC3BE" box="[199,310,993,1018]" pageId="4" pageNumber="31" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBD5D32F450EE56EFEEFFC4EFE1CC3BE" box="[322,487,993,1018]" collectionCode="NCBS-AU" pageId="4" pageNumber="31">NCBS-AU163</specimenCode>
(
<figureCitation id="134867D1450EE56EFE5BFC4EFDBFC3BE" box="[502,580,993,1018]" captionStart="FIGURE 3" captionStartId="6.[151,250,1983,2005]" captionTargetBox="[198,1364,220,1894]" captionTargetId="figure@6.[168,1393,193,1927]" captionTargetPageId="6" captionText="FIGURE 3. Holotype of Rhabdops aquaticus sp. nov. (NCBS-AU 163) whole preserved specimen in A) dorsal and B) ventral views. Scale bar 10 mm." httpUri="https://zenodo.org/record/1051971/files/figure.png" pageId="4" pageNumber="31">Figs. 3</figureCitation>
, 
<figureCitation id="134867D1450EE56EFDFCFC4DFD9AC3BE" box="[593,609,994,1018]" captionStart="FIGURE 4" captionStartId="7.[151,250,1970,1992]" captionTargetBox="[205,1382,224,1914]" captionTargetId="figure@7.[173,1414,193,1949]" captionTargetPageId="7" captionText="FIGURE 4. Holotype of Rhabdops aquaticus sp. nov. (NCBS-AU 163) head of preserved specimen in A) dorsal, B) ventral, C) right lateral and D) left lateral views. Scale bar 10 mm." httpUri="https://zenodo.org/record/1051973/files/figure.png" pageId="4" pageNumber="31">4</figureCitation>
), mature female, collected from 
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, 
<collectingMunicipality id="6BA8E12E450EE56EFB9FFC4EFB05C3BE" box="[1074,1278,993,1018]" pageId="4" pageNumber="31">Sindudurg district</collectingMunicipality>
, 
<collectingRegion id="49B7B5B6450EE56EFAA4FC4EFA63C3BE" box="[1289,1432,993,1018]" country="India" name="Maharashtra" pageId="4" pageNumber="31">Maharashtra</collectingRegion>
, 
<collectingCountry id="F3643BC4450EE56EFF3AFBAAFF2AC45A" box="[151,209,1029,1054]" name="India" pageId="4" pageNumber="31">India</collectingCountry>
(
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, 
<geoCoordinate id="EE471D93450EE56EFED2FBA9FDE9C45A" box="[383,530,1029,1054]" direction="east" orientation="longitude" pageId="4" pageNumber="31" precision="1" value="73.99752">E 73.997517</geoCoordinate>
; 
<elevation id="005E9C67450EE56EFDB0FBAAFD9FC45A" box="[541,612,1029,1054]" metricMagnitude="2" metricUnit="m" metricValue="7.45" pageId="4" pageNumber="31" unit="m" value="745.0">
<quantity id="4C8BD6B1450EE56EFDB0FBAAFD9FC45A" box="[541,612,1029,1054]" metricMagnitude="2" metricUnit="m" metricValue="7.45" pageId="4" pageNumber="31" unit="m" value="745.0">745 m</quantity>
</elevation>
) by 
<collectorName id="26861E82450EE56EFD38FBAAFD0DC459" box="[661,758,1029,1054]" pageId="4" pageNumber="31">Varad B.</collectorName>
Giri, 
<collectorName id="26861E82450EE56EFC95FBAAFC1BC459" box="[824,992,1029,1054]" pageId="4" pageNumber="31">Swapnil Pawar</collectorName>
and 
<collectorName id="26861E82450EE56EFBBBFBAAFB16C45A" box="[1046,1261,1029,1054]" pageId="4" pageNumber="31">Akshay Khandekar</collectorName>
on 
<collectingDate id="EF89A47C450EE56EFABBFBAAFA63C45A" box="[1302,1432,1029,1054]" pageId="4" pageNumber="31" value="2015-07-15">
<date id="FFCD5D94450EE56EFABBFBAAFA63C45A" box="[1302,1432,1029,1054]" pageId="4" pageNumber="31" value="2015-07-15">15.vii.2015</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCC7B54450EE56EFF6AFB86FAEAC4B2" blockId="4.[151,1437,849,1990]" pageId="4" pageNumber="31">
<materialsCitation id="3B1B7109450EE56EFF6AFB86FE9BC422" ID-GBIF-Occurrence="1632929378" collectingDate="1962-11-01" collectionCode="BNHS" collectorName="All &amp; Heads &amp; Koyna &amp; P. W. Soman" location="Satara District" pageId="4" pageNumber="31" specimenCode="BNHS 1724" specimenCount="1" stateProvince="Maharashtra" typeStatus="paratype">
<emphasis id="B907A746450EE56EFF6AFB86FE67C406" bold="true" box="[199,412,1065,1090]" pageId="4" pageNumber="31">
<typeStatus id="54C8C5F6450EE56EFF6AFB86FEC4C406" box="[199,319,1065,1090]" pageId="4" pageNumber="31" type="paratype">Paratypes</typeStatus>
(n = 7).
</emphasis>
<collectorName id="26861E82450EE56EFE0EFB86FE3CC406" box="[419,455,1065,1090]" pageId="4" pageNumber="31">All</collectorName>
males. 
<collectorName id="26861E82450EE56EFDB0FB86FD98C406" box="[541,611,1065,1090]" pageId="4" pageNumber="31">Heads</collectorName>
figured in 
<figureCitation id="134867D1450EE56EFD72FB86FCDFC406" box="[735,804,1065,1090]" captionStart="FIGURE 5" captionStartId="8.[151,250,1088,1110]" captionTargetBox="[157,1430,161,1062]" captionTargetId="figure@8.[151,1436,157,1066]" captionTargetPageId="8" captionText="FIGURE 5. Paratypes of Rhabdops aquaticus sp. nov. (specimen numbers given in first row). Heads of preserved specimen in A) dorsal, B) ventral, C) right lateral and D) left lateral views. Scale bar 10 mm." httpUri="https://zenodo.org/record/1051975/files/figure.png" pageId="4" pageNumber="31">Fig. 5</figureCitation>
. 
<specimenCode id="DBD5D32F450EE56EFC82FB85FC47C406" box="[815,956,1066,1090]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="31">BNHS 1724</specimenCode>
, 
<collectorName id="26861E82450EE56EFC6AFB85FBEEC406" box="[967,1045,1066,1090]" pageId="4" pageNumber="31">Koyna</collectorName>
, 
<location id="8EAC2D8F450EE56EFB8DFB85FB3FC406" LSID="urn:lsid:plazi:treatment:03DACA42450EE578FF3AFDE8FBD1C2C1:8EAC2D8F450EE56EFB8DFB85FB3FC406" box="[1056,1220,1065,1090]" name="Satara District" pageId="4" pageNumber="31" stateProvince="Maharashtra">Satara District</location>
, 
<collectingRegion id="49B7B5B6450EE56EFB7DFB86FAA4C406" box="[1232,1375,1065,1090]" country="India" name="Maharashtra" pageId="4" pageNumber="31">Maharashtra</collectingRegion>
, 
<collectorName id="26861E82450EE56EFAC7FB85FF13C422" pageId="4" pageNumber="31">P.W. Soman</collectorName>
, 
<collectingDate id="EF89A47C450EE56EFF5EFBE2FE9BC422" box="[243,352,1101,1126]" pageId="4" pageNumber="31" value="1962-11-01">
<date id="FFCD5D94450EE56EFF5EFBE2FE9BC422" box="[243,352,1101,1126]" pageId="4" pageNumber="31" value="1962-11-01">1.xi.1962</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="3B1B7109450EE56EFEC6FBE1FE42C4CE" ID-GBIF-Occurrence="1632929377" collectingDate="2003-09-18" collectionCode="BNHS" collectorName="V. B. Giri" county="Humbarli" elevation="940" latitude="17.410446" location="Satara District" longLatPrecision="1" longitude="73.72673" municipality="Koyna" pageId="4" pageNumber="31" specimenCode="BNHS 3234" specimenCount="1" stateProvince="Maharashtra" typeStatus="paratype">
<specimenCode id="DBD5D32F450EE56EFEC6FBE1FE03C422" box="[363,504,1102,1126]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="31">BNHS 3234</specimenCode>
, 
<collectingCounty id="62AD03D8450EE56EFDAEFBE2FD8AC422" box="[515,625,1101,1126]" pageId="4" pageNumber="31">Humbarli</collectingCounty>
, 
<collectingMunicipality id="6BA8E12E450EE56EFDD1FBE1FD32C422" box="[636,713,1102,1126]" pageId="4" pageNumber="31">Koyna</collectingMunicipality>
, 
<location id="8EAC2D8F450EE56EFD78FBE1FC82C422" LSID="urn:lsid:plazi:treatment:03DACA42450EE578FF3AFDE8FBD1C2C1:8EAC2D8F450EE56EFD78FBE1FC82C422" box="[725,889,1101,1126]" county="Humbarli" latitude="17.410446" longLatPrecision="1" longitude="73.72673" municipality="Koyna" name="Satara District" pageId="4" pageNumber="31" stateProvince="Maharashtra">Satara District</location>
, 
<collectingRegion id="49B7B5B6450EE56EFC29FBE2FBEBC422" box="[900,1040,1101,1126]" country="India" name="Maharashtra" pageId="4" pageNumber="31">Maharashtra</collectingRegion>
(
<geoCoordinate id="EE471D93450EE56EFBB2FBE1FB4EC422" box="[1055,1205,1101,1126]" direction="north" orientation="latitude" pageId="4" pageNumber="31" precision="1" value="17.410446">N 17.410446</geoCoordinate>
, 
<geoCoordinate id="EE471D93450EE56EFB6DFBE2FAB0C422" box="[1216,1355,1101,1126]" direction="east" orientation="longitude" pageId="4" pageNumber="31" precision="1" value="73.72673">E73.726732</geoCoordinate>
, 
<elevation id="005E9C67450EE56EFADFFBE1FF4BC4CE" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="4" pageNumber="31" unit="m" value="940.0">
<quantity id="4C8BD6B1450EE56EFADFFBE1FF4BC4CE" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="4" pageNumber="31" unit="m" value="940.0">940 m</quantity>
</elevation>
), 
<collectorName id="26861E82450EE56EFF6BFBDDFECAC4CE" box="[198,305,1137,1162]" pageId="4" pageNumber="31">V.B. Giri</collectorName>
, 
<collectingDate id="EF89A47C450EE56EFE93FBDEFE42C4CE" box="[318,441,1137,1162]" pageId="4" pageNumber="31" value="2003-09-18">
<date id="FFCD5D94450EE56EFE93FBDEFE42C4CE" box="[318,441,1137,1162]" pageId="4" pageNumber="31" value="2003-09-18">18.ix.2003</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="3B1B7109450EE56EFE6AFBDDFE62C4EA" ID-GBIF-Occurrence="1632929374" collectingDate="2004-07-13" collectionCode="BNHS" location="Satara District" pageId="4" pageNumber="31" specimenCode="BNHS 3235" specimenCount="1" stateProvince="Maharashtra" typeStatus="paratype">
<specimenCode id="DBD5D32F450EE56EFE6AFBDDFDADC4CE" box="[455,598,1137,1162]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="31">BNHS 3235</specimenCode>
, Humbarli, Koyna, 
<location id="8EAC2D8F450EE56EFC97FBDDFC1BC4CE" LSID="urn:lsid:plazi:treatment:03DACA42450EE578FF3AFDE8FBD1C2C1:8EAC2D8F450EE56EFC97FBDDFC1BC4CE" box="[826,992,1137,1162]" name="Satara District" pageId="4" pageNumber="31" stateProvince="Maharashtra">Satara District</location>
, 
<collectingRegion id="49B7B5B6450EE56EFC40FBDEFB87C4CE" box="[1005,1148,1137,1162]" country="India" name="Maharashtra" pageId="4" pageNumber="31">Maharashtra</collectingRegion>
, I. Agarwal, V.B. Giri, I. Kehimkar, 
<collectingDate id="EF89A47C450EE56EFEBBFB3AFE62C4EA" box="[278,409,1173,1198]" pageId="4" pageNumber="31" value="2004-07-13">
<date id="FFCD5D94450EE56EFEBBFB3AFE62C4EA" box="[278,409,1173,1198]" pageId="4" pageNumber="31" value="2004-07-13">13.vii.2004</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="3B1B7109450EE56EFE0AFB39FAB8C4EA" ID-GBIF-Occurrence="1632929380" box="[423,1347,1173,1198]" collectingDate="1986-02" collectionCode="BNHS" collectorName="S. A. Chougule" location="Sindhudurg District" pageId="4" pageNumber="31" specimenCode="BNHS 3347" specimenCount="1" stateProvince="Maharashtra" typeStatus="paratype">
<specimenCode id="DBD5D32F450EE56EFE0AFB39FDCCC4EA" box="[423,567,1174,1198]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="31">BNHS 3347</specimenCode>
, Amboli, 
<location id="8EAC2D8F450EE56EFD06FB3AFC75C4EA" LSID="urn:lsid:plazi:treatment:03DACA42450EE578FF3AFDE8FBD1C2C1:8EAC2D8F450EE56EFD06FB3AFC75C4EA" box="[683,910,1173,1198]" name="Sindhudurg District" pageId="4" pageNumber="31" stateProvince="Maharashtra">Sindhudurg District</location>
, 
<collectingRegion id="49B7B5B6450EE56EFC31FB3AFBD0C4EA" box="[924,1067,1173,1198]" country="India" name="Maharashtra" pageId="4" pageNumber="31">Maharashtra</collectingRegion>
, 
<collectorName id="26861E82450EE56EFB95FB39FB1FC4EA" box="[1080,1252,1173,1198]" pageId="4" pageNumber="31">S.A. Chougule</collectorName>
, 
<collectingDate id="EF89A47C450EE56EFB5FFB3AFAB8C4EA" box="[1266,1347,1173,1198]" pageId="4" pageNumber="31" value="1986-02">
<date id="FFCD5D94450EE56EFB5FFB3AFAB8C4EA" box="[1266,1347,1173,1198]" pageId="4" pageNumber="31" value="1986-02">ii.1986</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="3B1B7109450EE56EFAFFFB39FBEFC496" ID-GBIF-Occurrence="1632929376" collectingDate="2009-09" collectionCode="BNHS" collectorName="V. B. Giri" location="Sindhudurg District" pageId="4" pageNumber="31" specimenCode="BNHS 3509" specimenCount="1" stateProvince="Maharashtra" typeStatus="paratype">
<specimenCode id="DBD5D32F450EE56EFAFFFB39FF34C496" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="31">BNHS 3509</specimenCode>
and 3510, Amboli, 
<location id="8EAC2D8F450EE56EFE15FB16FD60C496" LSID="urn:lsid:plazi:treatment:03DACA42450EE578FF3AFDE8FBD1C2C1:8EAC2D8F450EE56EFE15FB16FD60C496" box="[440,667,1209,1234]" name="Sindhudurg District" pageId="4" pageNumber="31" stateProvince="Maharashtra">Sindhudurg District</location>
, 
<collectingRegion id="49B7B5B6450EE56EFD05FB16FCCCC496" box="[680,823,1209,1234]" country="India" name="Maharashtra" pageId="4" pageNumber="31">Maharashtra</collectingRegion>
, 
<collectorName id="26861E82450EE56EFCE9FB15FC54C496" box="[836,943,1209,1234]" pageId="4" pageNumber="31">V.B. Giri</collectorName>
, 
<collectingDate id="EF89A47C450EE56EFC11FB16FBEFC496" box="[956,1044,1209,1234]" pageId="4" pageNumber="31" value="2009-09">
<date id="FFCD5D94450EE56EFC11FB16FBEFC496" box="[956,1044,1209,1234]" pageId="4" pageNumber="31" value="2009-09">ix.2009</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="3B1B7109450EE56EFB8FFB16FAF7C4B2" ID-GBIF-Occurrence="1632929379" collectingDate="2014-08-20" collectionCode="NCBS-AU" collectorName="V. B. Giri &amp; Swapnil Pawar" elevation="975" latitude="16.748432" location="Kolhapur District" longLatPrecision="1" longitude="73.85016" municipality="Baraki" pageId="4" pageNumber="31" specimenCode="NCBS-AU162" specimenCount="1" stateProvince="Maharashtra" typeStatus="paratype">
<specimenCode id="DBD5D32F450EE56EFB8FFB16FB32C496" box="[1058,1225,1209,1234]" collectionCode="NCBS-AU" pageId="4" pageNumber="31">NCBS-AU162</specimenCode>
, 
<collectingMunicipality id="6BA8E12E450EE56EFB7AFB16FADEC496" box="[1239,1317,1209,1234]" pageId="4" pageNumber="31">Baraki</collectingMunicipality>
, 
<location id="8EAC2D8F450EE56EFA9EFB16FF15C4B2" LSID="urn:lsid:plazi:treatment:03DACA42450EE578FF3AFDE8FBD1C2C1:8EAC2D8F450EE56EFA9EFB16FF15C4B2" latitude="16.748432" longLatPrecision="1" longitude="73.85016" municipality="Baraki" name="Kolhapur District" pageId="4" pageNumber="31" stateProvince="Maharashtra">Kolhapur District</location>
, 
<collectingRegion id="49B7B5B6450EE56EFF54FB72FE7EC4B2" box="[249,389,1245,1270]" country="India" name="Maharashtra" pageId="4" pageNumber="31">Maharashtra</collectingRegion>
(
<geoCoordinate id="EE471D93450EE56EFE39FB71FDD1C4B2" box="[404,554,1245,1270]" direction="north" orientation="latitude" pageId="4" pageNumber="31" precision="1" value="16.748432">N 16.748433</geoCoordinate>
, 
<geoCoordinate id="EE471D93450EE56EFD98FB71FD33C4B2" box="[565,712,1245,1270]" direction="east" orientation="longitude" pageId="4" pageNumber="31" precision="1" value="73.85016">E 73.850155</geoCoordinate>
, 
<elevation id="005E9C67450EE56EFD7FFB72FCE7C4B2" box="[722,796,1245,1270]" metricMagnitude="2" metricUnit="m" metricValue="9.75" pageId="4" pageNumber="31" unit="m" value="975.0">
<quantity id="4C8BD6B1450EE56EFD7FFB72FCE7C4B2" box="[722,796,1245,1270]" metricMagnitude="2" metricUnit="m" metricValue="9.75" pageId="4" pageNumber="31" unit="m" value="975.0">975 m</quantity>
</elevation>
), 
<collectorName id="26861E82450EE56EFC9DFB71FC6DC4B2" box="[816,918,1245,1270]" pageId="4" pageNumber="31">V.B. Giri</collectorName>
and 
<collectorName id="26861E82450EE56EFC61FB72FB8CC4B1" box="[972,1143,1245,1270]" pageId="4" pageNumber="31">Swapnil Pawar</collectorName>
, 
<collectingDate id="EF89A47C450EE56EFB2FFB72FAF7C4B2" box="[1154,1292,1245,1270]" pageId="4" pageNumber="31" value="2014-08-20">
<date id="FFCD5D94450EE56EFB2FFB72FAF7C4B2" box="[1154,1292,1245,1270]" pageId="4" pageNumber="31" value="2014-08-20">20.viii.2014</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36928DF450EE567FF6AFAAEFE59C120" lastPageId="13" lastPageNumber="40" pageId="4" pageNumber="31" type="description">
<paragraph id="8BCC7B54450EE56EFF6AFAAEFB65C5C1" blockId="4.[151,1437,849,1990]" pageId="4" pageNumber="31">
<emphasis id="B907A746450EE56EFF6AFAAEFDFAC55E" bold="true" box="[199,513,1281,1306]" pageId="4" pageNumber="31">Referred specimen (n = 1).</emphasis>
BNHS 3194, male. The BNHS catalogue reports the locality as “near Mula-Mutha River, Manjari Budruk, Pune District” but we doubt that this is correct because, unlike all of the other known localities for the species, Pune is ca. 
<quantity id="4C8BD6B1450EE56EFDE3FAE6FD62C526" box="[590,665,1353,1378]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="31" unit="km" value="35.0">35 km</quantity>
east of the Western Ghats and instead part of the Deccan region. Without verification, we consider the precise locality unknown, probably Maharashtra state.
</paragraph>
<paragraph id="8BCC7B54450EE56EFF6AFA3EFAE4C5B6" blockId="4.[151,1437,849,1990]" pageId="4" pageNumber="31">
<emphasis id="B907A746450EE56EFF6AFA3EFE1DC5EE" bold="true" box="[199,486,1425,1450]" pageId="4" pageNumber="31">
Description of 
<typeStatus id="54C8C5F6450EE56EFEDAFA3EFE1BC5EE" box="[375,480,1425,1450]" pageId="4" pageNumber="31" type="holotype">holotype</typeStatus>
.
</emphasis>
See 
<tableCitation id="C6F14EEF450EE56EFDB1FA3EFD8BC5EE" box="[540,624,1425,1450]" captionStart="TABLE 2" captionStartId="5.[125,202,585,606]" captionText="TABLE 2. Meristic anđ morphometric (in millimetres) đata for examineđ specimens of Rhabdops aquaticus sp. nov. All specimens are paratypes except for the holotype (*) anđ a referređ specimen (**). Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because" pageId="4" pageNumber="31">Table 2</tableCitation>
for morphometric and meristic data. Female. Good condition; ca. 
<quantity id="4C8BD6B1450EE56EFAF6FA3DFA66C5ED" box="[1371,1437,1426,1449]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="31" unit="mm" value="7.0">7 mm</quantity>
longitudinal ventral incision into coelom ca. 
<quantity id="4C8BD6B1450EE56EFD27FA19FD1CC589" box="[650,743,1462,1486]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="31" unit="mm" value="200.0">200 mm</quantity>
from snout tip. Some body scales, especially posteriorly, with slightly blistered surface, possibly caused by artefactual separation of parts of oberhautchen scale layer.
</paragraph>
<paragraph id="8BCC7B54450EE56EFF6AFA52FB74C68E" blockId="4.[151,1437,849,1990]" pageId="4" pageNumber="31">
Body subcylindrical, slightly flatter on venter, widest at midbody, gently tapering anteriorly and posteriorly, though with artefactual ca. 
<quantity id="4C8BD6B1450EE56EFE70F98EFDCAC67D" box="[477,561,1569,1594]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="31" unit="mm" value="25.0">25 mm</quantity>
constriction ca. 
<quantity id="4C8BD6B1450EE56EFD54F98EFCB6C67D" box="[761,845,1569,1594]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="31" unit="mm" value="35.0">35 mm</quantity>
from vent. Tail much more strongly tapered than posterior of body. Head broader than tall, slightly wider than anterior of body. In dorsal view head ovate, sides gently converging anteriorly, slightly convex, apex of convexity approximately level with back of frontal. Front of snout not pointed, truncated. In lateral view head tapers gently from back to eye, more strongly tapered in front of eye. Paired scales on top of head abutting along midline rather than imbricate/overlapping.
</paragraph>
<paragraph id="8BCC7B54450EE56EFF6AF979FE4FC782" blockId="4.[151,1437,849,1990]" pageId="4" pageNumber="31">In dorsal view rostral approximately three times broader than long, shorter than distance between it and frontal; projects beyond tip of lower jaw; ventrally with transverse concavity, notched (U-shaped) at margin of mouth. Frontal hexagonal, lateral edges anteriorly diverging; shorter and smaller than each parietal, longer than distance between it and snout tip. Paired internasals smaller than paired prefrontals. Nasals squarish (though 5-sided), ventral edge notched centrally, marginally larger (difference less on left) than slightly elongate loreal. Single supraocular each side, slightly smaller than prefrontals. Two subequal preoculars smaller than two postoculars (upper postocular larger).</paragraph>
<caption id="DF0C2BDC450FE56FFFD0FDE6F8A2C23D" pageId="5" pageNumber="32">
<paragraph id="8BCC7B54450FE56FFFD0FDE6F8A2C23D" blockId="5.[125,1884,584,714]" pageId="5" pageNumber="32">
<emphasis id="B907A746450FE56FFFD0FDE6FF18C21A" bold="true" box="[125,227,585,606]" pageId="5" pageNumber="32">TABLE 2.</emphasis>
Meristic anđ morphometric (in millimetres) đata for examineđ specimens of 
<taxonomicName id="4C7300D7450FE56FFC1BFDE7FB8BC21A" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[950,1136,584,606]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="32" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A746450FE56FFC1BFDE7FB8BC21A" box="[950,1136,584,606]" italics="true" pageId="5" pageNumber="32">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D450FE56FFBD5FDE6FB3FC21A" box="[1144,1220,585,606]" pageId="5" pageNumber="32" rank="species">
<emphasis id="B907A746450FE56FFBD5FDE6FB3FC21A" bold="true" box="[1144,1220,585,606]" pageId="5" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
All specimens are paratypes except for the holotype (*) anđ a referređ specimen (**). Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because
</paragraph>
</caption>
<paragraph id="8BCC7B54450FE56FFF3AFDD0FF2EC28E" blockId="5.[125,1884,584,714]" pageId="5" pageNumber="32">slight đamage to specimen. Invariant characters not recorđeđ here incluđe: number of preoculars (2,2), postoculars (2,2), total scales surrounđing eye (6,6), prefrontals (2), internasals (2), supralabials (5,5); largest infralabial (5,5), supralabial in contact with eye (3,3), supralabials in contact with loreal (1&amp;2, 1&amp;2), anal shielđs (paiređ, right overlapping left). See Appenđix 2 for localities.</paragraph>
<paragraph id="8BCC7B54450FE56FFFD0FC8CF8F1C604" pageId="5" pageNumber="32">
<table id="F97389F4450F1A95FFD0FD43F8B8C604" box="[125,1859,748,1600]" gridcols="10" gridrows="20" pageId="5" pageNumber="32">
<tr id="35437916450F1A95FFD0FD43F8B8C37C" box="[125,1859,748,824]" gridrow="0" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FD43FD93C37C" box="[125,616,748,824]" gridcol="0" gridrow="0" pageId="5" pageNumber="32">Sex</th>
<th id="7692106A450F1A95FCF9FD43FC5AC37C" box="[852,929,748,824]" gridcol="1" gridrow="0" pageId="5" pageNumber="32">NCBS- AU163* F</th>
<th id="7692106A450F1A95FC64FD43FBFEC37C" box="[969,1029,748,824]" gridcol="2" gridrow="0" pageId="5" pageNumber="32">BNHS 3510 M</th>
<th id="7692106A450F1A95FB93FD43FB80C37C" box="[1086,1147,748,824]" gridcol="3" gridrow="0" pageId="5" pageNumber="32">BNHS 3509 M</th>
<th id="7692106A450F1A95FB19FD43FB0CC37C" box="[1204,1271,748,824]" gridcol="4" gridrow="0" pageId="5" pageNumber="32">NCBS- AU162 M</th>
<th id="7692106A450F1A95FA84FD43FA9DC37C" box="[1321,1382,748,824]" gridcol="5" gridrow="0" pageId="5" pageNumber="32">BNHS 3234 M</th>
<th id="7692106A450F1A95FA32FD43FA20C37C" box="[1439,1499,748,824]" gridcol="6" gridrow="0" pageId="5" pageNumber="32">BNHS 1724 M</th>
<th id="7692106A450F1A95F9B9FD43F9ABC37C" box="[1556,1616,748,824]" gridcol="7" gridrow="0" pageId="5" pageNumber="32">BNHS 3347 M</th>
<th id="7692106A450F1A95F927FD43F93DC37C" box="[1674,1734,748,824]" gridcol="8" gridrow="0" pageId="5" pageNumber="32">BNHS 3255 M</th>
<th id="7692106A450F1A95F952FD43F8B8C37C" box="[1791,1859,748,824]" gridcol="9" gridrow="0" pageId="5" pageNumber="32">BNHS 3194** M</th>
</tr>
<tr id="35437916450F1A95FFD0FCE7F8B8C319" box="[125,1859,840,861]" gridrow="1" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FCE7FD93C319" box="[125,616,840,861]" gridcol="0" gridrow="1" pageId="5" pageNumber="32">Snout tip-vent length length</th>
<td id="7692106A450F1A95FCF9FCE7FC5AC319" box="[852,929,840,861]" gridcol="1" gridrow="1" pageId="5" pageNumber="32">650</td>
<td id="7692106A450F1A95FC64FCE7FBFEC319" box="[969,1029,840,861]" gridcol="2" gridrow="1" pageId="5" pageNumber="32">745</td>
<td id="7692106A450F1A95FB93FCE7FB80C319" box="[1086,1147,840,861]" gridcol="3" gridrow="1" pageId="5" pageNumber="32">720</td>
<td id="7692106A450F1A95FB19FCE7FB0CC319" box="[1204,1271,840,861]" gridcol="4" gridrow="1" pageId="5" pageNumber="32">580</td>
<td id="7692106A450F1A95FA84FCE7FA9DC319" box="[1321,1382,840,861]" gridcol="5" gridrow="1" pageId="5" pageNumber="32">505</td>
<td id="7692106A450F1A95FA32FCE7FA20C319" box="[1439,1499,840,861]" gridcol="6" gridrow="1" pageId="5" pageNumber="32">379</td>
<td id="7692106A450F1A95F9B9FCE7F9ABC319" box="[1556,1616,840,861]" gridcol="7" gridrow="1" pageId="5" pageNumber="32">350</td>
<td id="7692106A450F1A95F927FCE7F93DC319" box="[1674,1734,840,861]" gridcol="8" gridrow="1" pageId="5" pageNumber="32">337</td>
<td id="7692106A450F1A95F952FCE7F8B8C319" box="[1791,1859,840,861]" gridcol="9" gridrow="1" pageId="5" pageNumber="32">540</td>
</tr>
<tr id="35437916450F1A95FFD0FCC2F8B8C3C6" box="[125,1859,877,898]" gridrow="2" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FCC2FD93C3C6" box="[125,616,877,898]" gridcol="0" gridrow="2" pageId="5" pageNumber="32">Tail length</th>
<td id="7692106A450F1A95FCF9FCC2FC5AC3C6" box="[852,929,877,898]" gridcol="1" gridrow="2" pageId="5" pageNumber="32">120</td>
<td id="7692106A450F1A95FC64FCC2FBFEC3C6" box="[969,1029,877,898]" gridcol="2" gridrow="2" pageId="5" pageNumber="32">200</td>
<td id="7692106A450F1A95FB93FCC2FB80C3C6" box="[1086,1147,877,898]" gridcol="3" gridrow="2" pageId="5" pageNumber="32">180</td>
<td id="7692106A450F1A95FB19FCC2FB0CC3C6" box="[1204,1271,877,898]" gridcol="4" gridrow="2" pageId="5" pageNumber="32">155</td>
<td id="7692106A450F1A95FA84FCC2FA9DC3C6" box="[1321,1382,877,898]" gridcol="5" gridrow="2" pageId="5" pageNumber="32">140</td>
<td id="7692106A450F1A95FA32FCC2FA20C3C6" box="[1439,1499,877,898]" gridcol="6" gridrow="2" pageId="5" pageNumber="32">89</td>
<td id="7692106A450F1A95F9B9FCC2F9ABC3C6" box="[1556,1616,877,898]" gridcol="7" gridrow="2" pageId="5" pageNumber="32">75</td>
<td id="7692106A450F1A95F927FCC2F93DC3C6" box="[1674,1734,877,898]" gridcol="8" gridrow="2" pageId="5" pageNumber="32">65</td>
<td id="7692106A450F1A95F952FCC2F8B8C3C6" box="[1791,1859,877,898]" gridcol="9" gridrow="2" pageId="5" pageNumber="32">140</td>
</tr>
<tr id="35437916450F1A95FFD0FC3DF8B8C3E3" box="[125,1859,914,935]" gridrow="3" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FC3DFD93C3E3" box="[125,616,914,935]" gridcol="0" gridrow="3" pageId="5" pageNumber="32">Total ventral scales (Dowling methođ)</th>
<td id="7692106A450F1A95FCF9FC3DFC5AC3E3" box="[852,929,914,935]" gridcol="1" gridrow="3" pageId="5" pageNumber="32">230</td>
<td id="7692106A450F1A95FC64FC3DFBFEC3E3" box="[969,1029,914,935]" gridcol="2" gridrow="3" pageId="5" pageNumber="32">228</td>
<td id="7692106A450F1A95FB93FC3DFB80C3E3" box="[1086,1147,914,935]" gridcol="3" gridrow="3" pageId="5" pageNumber="32">227</td>
<td id="7692106A450F1A95FB19FC3DFB0CC3E3" box="[1204,1271,914,935]" gridcol="4" gridrow="3" pageId="5" pageNumber="32">227</td>
<td id="7692106A450F1A95FA84FC3DFA9DC3E3" box="[1321,1382,914,935]" gridcol="5" gridrow="3" pageId="5" pageNumber="32">229</td>
<td id="7692106A450F1A95FA32FC3DFA20C3E3" box="[1439,1499,914,935]" gridcol="6" gridrow="3" pageId="5" pageNumber="32">225</td>
<td id="7692106A450F1A95F9B9FC3DF9ABC3E3" box="[1556,1616,914,935]" gridcol="7" gridrow="3" pageId="5" pageNumber="32">229</td>
<td id="7692106A450F1A95F927FC3DF93DC3E3" box="[1674,1734,914,935]" gridcol="8" gridrow="3" pageId="5" pageNumber="32">226</td>
<td id="7692106A450F1A95F952FC3DF8B8C3E3" box="[1791,1859,914,935]" gridcol="9" gridrow="3" pageId="5" pageNumber="32">224</td>
</tr>
<tr id="35437916450F1A95FFD0FC18F8B8C388" box="[125,1859,951,972]" gridrow="4" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FC18FD93C388" box="[125,616,951,972]" gridcol="0" gridrow="4" pageId="5" pageNumber="32">Total subcauđal scales</th>
<td id="7692106A450F1A95FCF9FC18FC5AC388" box="[852,929,951,972]" gridcol="1" gridrow="4" pageId="5" pageNumber="32">66,66</td>
<td id="7692106A450F1A95FC64FC18FBFEC388" box="[969,1029,951,972]" gridcol="2" gridrow="4" pageId="5" pageNumber="32">77,77</td>
<td id="7692106A450F1A95FB93FC18FB80C388" box="[1086,1147,951,972]" gridcol="3" gridrow="4" pageId="5" pageNumber="32">79,79</td>
<td id="7692106A450F1A95FB19FC18FB0CC388" box="[1204,1271,951,972]" gridcol="4" gridrow="4" pageId="5" pageNumber="32">79,79</td>
<td id="7692106A450F1A95FA84FC18FA9DC388" box="[1321,1382,951,972]" gridcol="5" gridrow="4" pageId="5" pageNumber="32">77,76</td>
<td id="7692106A450F1A95FA32FC18FA20C388" box="[1439,1499,951,972]" gridcol="6" gridrow="4" pageId="5" pageNumber="32">75,75</td>
<td id="7692106A450F1A95F9B9FC18F9ABC388" box="[1556,1616,951,972]" gridcol="7" gridrow="4" pageId="5" pageNumber="32">81,81</td>
<td id="7692106A450F1A95F927FC18F93DC388" box="[1674,1734,951,972]" gridcol="8" gridrow="4" pageId="5" pageNumber="32">77,76</td>
<td id="7692106A450F1A95F952FC18F8B8C388" box="[1791,1859,951,972]" gridcol="9" gridrow="4" pageId="5" pageNumber="32">78,78</td>
</tr>
<tr id="35437916450F1A95FFD0FC73F8B8C3B5" box="[125,1859,988,1009]" gridrow="5" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FC73FD93C3B5" box="[125,616,988,1009]" gridcol="0" gridrow="5" pageId="5" pageNumber="32">Heađ length (snout tip to back of manđible)</th>
<td id="7692106A450F1A95FCF9FC73FC5AC3B5" box="[852,929,988,1009]" gridcol="1" gridrow="5" pageId="5" pageNumber="32">15.5</td>
<td id="7692106A450F1A95FC64FC73FBFEC3B5" box="[969,1029,988,1009]" gridcol="2" gridrow="5" pageId="5" pageNumber="32">17.2</td>
<td id="7692106A450F1A95FB93FC73FB80C3B5" box="[1086,1147,988,1009]" gridcol="3" gridrow="5" pageId="5" pageNumber="32">15.7</td>
<td id="7692106A450F1A95FB19FC73FB0CC3B5" box="[1204,1271,988,1009]" gridcol="4" gridrow="5" pageId="5" pageNumber="32">12.8</td>
<td id="7692106A450F1A95FA84FC73FA9DC3B5" box="[1321,1382,988,1009]" gridcol="5" gridrow="5" pageId="5" pageNumber="32">14.3</td>
<td id="7692106A450F1A95FA32FC73FA20C3B5" box="[1439,1499,988,1009]" gridcol="6" gridrow="5" pageId="5" pageNumber="32">12.8</td>
<td id="7692106A450F1A95F9B9FC73F9ABC3B5" box="[1556,1616,988,1009]" gridcol="7" gridrow="5" pageId="5" pageNumber="32">10.7</td>
<td id="7692106A450F1A95F927FC73F93DC3B5" box="[1674,1734,988,1009]" gridcol="8" gridrow="5" pageId="5" pageNumber="32">11.2</td>
<td id="7692106A450F1A95F952FC73F8B8C3B5" box="[1791,1859,988,1009]" gridcol="9" gridrow="5" pageId="5" pageNumber="32">(12.5)</td>
</tr>
<tr id="35437916450F1A95FFD0FBAEF8B8C452" box="[125,1859,1025,1046]" gridrow="6" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FBAEFD93C452" box="[125,616,1025,1046]" gridcol="0" gridrow="6" pageId="5" pageNumber="32">Maximum heađ wiđth</th>
<td id="7692106A450F1A95FCF9FBAEFC5AC452" box="[852,929,1025,1046]" gridcol="1" gridrow="6" pageId="5" pageNumber="32">11.0</td>
<td id="7692106A450F1A95FC64FBAEFBFEC452" box="[969,1029,1025,1046]" gridcol="2" gridrow="6" pageId="5" pageNumber="32">10.8</td>
<td id="7692106A450F1A95FB93FBAEFB80C452" box="[1086,1147,1025,1046]" gridcol="3" gridrow="6" pageId="5" pageNumber="32">12.3</td>
<td id="7692106A450F1A95FB19FBAEFB0CC452" box="[1204,1271,1025,1046]" gridcol="4" gridrow="6" pageId="5" pageNumber="32">8.7</td>
<td id="7692106A450F1A95FA84FBAEFA9DC452" box="[1321,1382,1025,1046]" gridcol="5" gridrow="6" pageId="5" pageNumber="32">9.2</td>
<td id="7692106A450F1A95FA32FBAEFA20C452" box="[1439,1499,1025,1046]" gridcol="6" gridrow="6" pageId="5" pageNumber="32">6.9</td>
<td id="7692106A450F1A95F9B9FBAEF9ABC452" box="[1556,1616,1025,1046]" gridcol="7" gridrow="6" pageId="5" pageNumber="32">6.9</td>
<td id="7692106A450F1A95F927FBAEF93DC452" box="[1674,1734,1025,1046]" gridcol="8" gridrow="6" pageId="5" pageNumber="32">6.4</td>
<td id="7692106A450F1A95F952FBAEF8B8C452" box="[1791,1859,1025,1046]" gridcol="9" gridrow="6" pageId="5" pageNumber="32">8.0</td>
</tr>
<tr id="35437916450F1A95FFD0FB89F8B8C47F" box="[125,1859,1062,1083]" gridrow="7" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FB89FD93C47F" box="[125,616,1062,1083]" gridcol="0" gridrow="7" pageId="5" pageNumber="32">Circumference at miđbođy</th>
<td id="7692106A450F1A95FCF9FB89FC5AC47F" box="[852,929,1062,1083]" gridcol="1" gridrow="7" pageId="5" pageNumber="32">55</td>
<td id="7692106A450F1A95FC64FB89FBFEC47F" box="[969,1029,1062,1083]" gridcol="2" gridrow="7" pageId="5" pageNumber="32">58</td>
<td id="7692106A450F1A95FB93FB89FB80C47F" box="[1086,1147,1062,1083]" gridcol="3" gridrow="7" pageId="5" pageNumber="32">50</td>
<td id="7692106A450F1A95FB19FB89FB0CC47F" box="[1204,1271,1062,1083]" gridcol="4" gridrow="7" pageId="5" pageNumber="32">44</td>
<td id="7692106A450F1A95FA84FB89FA9DC47F" box="[1321,1382,1062,1083]" gridcol="5" gridrow="7" pageId="5" pageNumber="32">40</td>
<td id="7692106A450F1A95FA32FB89FA20C47F" box="[1439,1499,1062,1083]" gridcol="6" gridrow="7" pageId="5" pageNumber="32">26</td>
<td id="7692106A450F1A95F9B9FB89F9ABC47F" box="[1556,1616,1062,1083]" gridcol="7" gridrow="7" pageId="5" pageNumber="32">30</td>
<td id="7692106A450F1A95F927FB89F93DC47F" box="[1674,1734,1062,1083]" gridcol="8" gridrow="7" pageId="5" pageNumber="32">24</td>
<td id="7692106A450F1A95F952FB89F8B8C47F" box="[1791,1859,1062,1083]" gridcol="9" gridrow="7" pageId="5" pageNumber="32">35</td>
</tr>
<tr id="35437916450F1A95FFD0FBE4F8B8C424" box="[125,1859,1099,1120]" gridrow="8" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FBE4FD93C424" box="[125,616,1099,1120]" gridcol="0" gridrow="8" pageId="5" pageNumber="32">Nostril-nostril</th>
<td id="7692106A450F1A95FCF9FBE4FC5AC424" box="[852,929,1099,1120]" gridcol="1" gridrow="8" pageId="5" pageNumber="32">5.3</td>
<td id="7692106A450F1A95FC64FBE4FBFEC424" box="[969,1029,1099,1120]" gridcol="2" gridrow="8" pageId="5" pageNumber="32">5.2</td>
<td id="7692106A450F1A95FB93FBE4FB80C424" box="[1086,1147,1099,1120]" gridcol="3" gridrow="8" pageId="5" pageNumber="32">5.6</td>
<td id="7692106A450F1A95FB19FBE4FB0CC424" box="[1204,1271,1099,1120]" gridcol="4" gridrow="8" pageId="5" pageNumber="32">4.5</td>
<td id="7692106A450F1A95FA84FBE4FA9DC424" box="[1321,1382,1099,1120]" gridcol="5" gridrow="8" pageId="5" pageNumber="32">4.6</td>
<td id="7692106A450F1A95FA32FBE4FA20C424" box="[1439,1499,1099,1120]" gridcol="6" gridrow="8" pageId="5" pageNumber="32">3.6</td>
<td id="7692106A450F1A95F9B9FBE4F9ABC424" box="[1556,1616,1099,1120]" gridcol="7" gridrow="8" pageId="5" pageNumber="32">3.7</td>
<td id="7692106A450F1A95F927FBE4F93DC424" box="[1674,1734,1099,1120]" gridcol="8" gridrow="8" pageId="5" pageNumber="32">3.2</td>
<td id="7692106A450F1A95F952FBE4F8B8C424" box="[1791,1859,1099,1120]" gridcol="9" gridrow="8" pageId="5" pageNumber="32">4.7</td>
</tr>
<tr id="35437916450F1A95FFD0FBDFF8B8C4C1" box="[125,1859,1136,1157]" gridrow="9" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FBDFFD93C4C1" box="[125,616,1136,1157]" gridcol="0" gridrow="9" pageId="5" pageNumber="32">Eye-eye</th>
<td id="7692106A450F1A95FCF9FBDFFC5AC4C1" box="[852,929,1136,1157]" gridcol="1" gridrow="9" pageId="5" pageNumber="32">7.2</td>
<td id="7692106A450F1A95FC64FBDFFBFEC4C1" box="[969,1029,1136,1157]" gridcol="2" gridrow="9" pageId="5" pageNumber="32">7.8</td>
<td id="7692106A450F1A95FB93FBDFFB80C4C1" box="[1086,1147,1136,1157]" gridcol="3" gridrow="9" pageId="5" pageNumber="32">7.6</td>
<td id="7692106A450F1A95FB19FBDFFB0CC4C1" box="[1204,1271,1136,1157]" gridcol="4" gridrow="9" pageId="5" pageNumber="32">5.7</td>
<td id="7692106A450F1A95FA84FBDFFA9DC4C1" box="[1321,1382,1136,1157]" gridcol="5" gridrow="9" pageId="5" pageNumber="32">6.6</td>
<td id="7692106A450F1A95FA32FBDFFA20C4C1" box="[1439,1499,1136,1157]" gridcol="6" gridrow="9" pageId="5" pageNumber="32">5.2</td>
<td id="7692106A450F1A95F9B9FBDFF9ABC4C1" box="[1556,1616,1136,1157]" gridcol="7" gridrow="9" pageId="5" pageNumber="32">5.4</td>
<td id="7692106A450F1A95F927FBDFF93DC4C1" box="[1674,1734,1136,1157]" gridcol="8" gridrow="9" pageId="5" pageNumber="32">4.7</td>
<td id="7692106A450F1A95F952FBDFF8B8C4C1" box="[1791,1859,1136,1157]" gridcol="9" gridrow="9" pageId="5" pageNumber="32">6.4</td>
</tr>
<tr id="35437916450F1A95FFD0FB3AF8B8C48A" box="[125,1859,1173,1230]" gridrow="10" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FB3AFD93C48A" box="[125,616,1173,1230]" gridcol="0" gridrow="10" pageId="5" pageNumber="32">Eye-snout tip Diameter of exposeđ part of eye</th>
<td id="7692106A450F1A95FCF9FB3AFC5AC48A" box="[852,929,1173,1230]" gridcol="1" gridrow="10" pageId="5" pageNumber="32">6.5 2.5</td>
<td id="7692106A450F1A95FC64FB3AFBFEC48A" box="[969,1029,1173,1230]" gridcol="2" gridrow="10" pageId="5" pageNumber="32">7.0 2.6</td>
<td id="7692106A450F1A95FB93FB3AFB80C48A" box="[1086,1147,1173,1230]" gridcol="3" gridrow="10" pageId="5" pageNumber="32">6.6 2.7</td>
<td id="7692106A450F1A95FB19FB3AFB0CC48A" box="[1204,1271,1173,1230]" gridcol="4" gridrow="10" pageId="5" pageNumber="32">5.0 2.1</td>
<td id="7692106A450F1A95FA84FB3AFA9DC48A" box="[1321,1382,1173,1230]" gridcol="5" gridrow="10" pageId="5" pageNumber="32">5.7 2.0</td>
<td id="7692106A450F1A95FA32FB3AFA20C48A" box="[1439,1499,1173,1230]" gridcol="6" gridrow="10" pageId="5" pageNumber="32">4.2 1.7</td>
<td id="7692106A450F1A95F9B9FB3AF9ABC48A" box="[1556,1616,1173,1230]" gridcol="7" gridrow="10" pageId="5" pageNumber="32">4.6 2.0</td>
<td id="7692106A450F1A95F927FB3AF93DC48A" box="[1674,1734,1173,1230]" gridcol="8" gridrow="10" pageId="5" pageNumber="32">3.7 1.4</td>
<td id="7692106A450F1A95F952FB3AF8B8C48A" box="[1791,1859,1173,1230]" gridcol="9" gridrow="10" pageId="5" pageNumber="32">(4.1) 1.9</td>
</tr>
<tr id="35437916450F1A95FFD0FB70F8B8C55C" box="[125,1859,1247,1304]" gridrow="11" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FB70FD93C55C" box="[125,616,1247,1304]" gridcol="0" gridrow="11" pageId="5" pageNumber="32">Maximum wiđth of rostral Maximum length of frontal</th>
<td id="7692106A450F1A95FCF9FB70FC5AC55C" box="[852,929,1247,1304]" gridcol="1" gridrow="11" pageId="5" pageNumber="32">5.3 5.8</td>
<td id="7692106A450F1A95FC64FB70FBFEC55C" box="[969,1029,1247,1304]" gridcol="2" gridrow="11" pageId="5" pageNumber="32">5.4 5.2</td>
<td id="7692106A450F1A95FB93FB70FB80C55C" box="[1086,1147,1247,1304]" gridcol="3" gridrow="11" pageId="5" pageNumber="32">5.6 5.6</td>
<td id="7692106A450F1A95FB19FB70FB0CC55C" box="[1204,1271,1247,1304]" gridcol="4" gridrow="11" pageId="5" pageNumber="32">4.3 4.3</td>
<td id="7692106A450F1A95FA84FB70FA9DC55C" box="[1321,1382,1247,1304]" gridcol="5" gridrow="11" pageId="5" pageNumber="32">4.4 4.6</td>
<td id="7692106A450F1A95FA32FB70FA20C55C" box="[1439,1499,1247,1304]" gridcol="6" gridrow="11" pageId="5" pageNumber="32">3.9 3.9</td>
<td id="7692106A450F1A95F9B9FB70F9ABC55C" box="[1556,1616,1247,1304]" gridcol="7" gridrow="11" pageId="5" pageNumber="32">3.7 4.1</td>
<td id="7692106A450F1A95F927FB70F93DC55C" box="[1674,1734,1247,1304]" gridcol="8" gridrow="11" pageId="5" pageNumber="32">3.3 3.4</td>
<td id="7692106A450F1A95F952FB70F8B8C55C" box="[1791,1859,1247,1304]" gridcol="9" gridrow="11" pageId="5" pageNumber="32">(3.6) 5.0</td>
</tr>
<tr id="35437916450F1A95FFD0FA87F8B8C579" box="[125,1859,1320,1341]" gridrow="12" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FA87FD93C579" box="[125,616,1320,1341]" gridcol="0" gridrow="12" pageId="5" pageNumber="32">Maximum wiđth of frontal</th>
<td id="7692106A450F1A95FCF9FA87FC5AC579" box="[852,929,1320,1341]" gridcol="1" gridrow="12" pageId="5" pageNumber="32">4.5</td>
<td id="7692106A450F1A95FC64FA87FBFEC579" box="[969,1029,1320,1341]" gridcol="2" gridrow="12" pageId="5" pageNumber="32">4.3</td>
<td id="7692106A450F1A95FB93FA87FB80C579" box="[1086,1147,1320,1341]" gridcol="3" gridrow="12" pageId="5" pageNumber="32">5.1</td>
<td id="7692106A450F1A95FB19FA87FB0CC579" box="[1204,1271,1320,1341]" gridcol="4" gridrow="12" pageId="5" pageNumber="32">4.0</td>
<td id="7692106A450F1A95FA84FA87FA9DC579" box="[1321,1382,1320,1341]" gridcol="5" gridrow="12" pageId="5" pageNumber="32">4.0</td>
<td id="7692106A450F1A95FA32FA87FA20C579" box="[1439,1499,1320,1341]" gridcol="6" gridrow="12" pageId="5" pageNumber="32">3.4</td>
<td id="7692106A450F1A95F9B9FA87F9ABC579" box="[1556,1616,1320,1341]" gridcol="7" gridrow="12" pageId="5" pageNumber="32">3.4</td>
<td id="7692106A450F1A95F927FA87F93DC579" box="[1674,1734,1320,1341]" gridcol="8" gridrow="12" pageId="5" pageNumber="32">3.1</td>
<td id="7692106A450F1A95F952FA87F8B8C579" box="[1791,1859,1320,1341]" gridcol="9" gridrow="12" pageId="5" pageNumber="32">4.4</td>
</tr>
<tr id="35437916450F1A95FFD0FAE2F8B8C526" box="[125,1859,1357,1378]" gridrow="13" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FAE2FD93C526" box="[125,616,1357,1378]" gridcol="0" gridrow="13" pageId="5" pageNumber="32">Maximum length of parietal</th>
<td id="7692106A450F1A95FCF9FAE2FC5AC526" box="[852,929,1357,1378]" gridcol="1" gridrow="13" pageId="5" pageNumber="32">6.6</td>
<td id="7692106A450F1A95FC64FAE2FBFEC526" box="[969,1029,1357,1378]" gridcol="2" gridrow="13" pageId="5" pageNumber="32">6.4</td>
<td id="7692106A450F1A95FB93FAE2FB80C526" box="[1086,1147,1357,1378]" gridcol="3" gridrow="13" pageId="5" pageNumber="32">7.2</td>
<td id="7692106A450F1A95FB19FAE2FB0CC526" box="[1204,1271,1357,1378]" gridcol="4" gridrow="13" pageId="5" pageNumber="32">6.8</td>
<td id="7692106A450F1A95FA84FAE2FA9DC526" box="[1321,1382,1357,1378]" gridcol="5" gridrow="13" pageId="5" pageNumber="32">6.4</td>
<td id="7692106A450F1A95FA32FAE2FA20C526" box="[1439,1499,1357,1378]" gridcol="6" gridrow="13" pageId="5" pageNumber="32">5.3</td>
<td id="7692106A450F1A95F9B9FAE2F9ABC526" box="[1556,1616,1357,1378]" gridcol="7" gridrow="13" pageId="5" pageNumber="32">4.8</td>
<td id="7692106A450F1A95F927FAE2F93DC526" box="[1674,1734,1357,1378]" gridcol="8" gridrow="13" pageId="5" pageNumber="32">4.8</td>
<td id="7692106A450F1A95F952FAE2F8B8C526" box="[1791,1859,1357,1378]" gridcol="9" gridrow="13" pageId="5" pageNumber="32">6.2</td>
</tr>
<tr id="35437916450F1A95FFD0FADDF8B8C5C3" box="[125,1859,1394,1415]" gridrow="14" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FADDFD93C5C3" box="[125,616,1394,1415]" gridcol="0" gridrow="14" pageId="5" pageNumber="32">No. scales contacting parietal (excluđing heađ shielđs)</th>
<td id="7692106A450F1A95FCF9FADDFC5AC5C3" box="[852,929,1394,1415]" gridcol="1" gridrow="14" pageId="5" pageNumber="32">3,3</td>
<td id="7692106A450F1A95FC64FADDFBFEC5C3" box="[969,1029,1394,1415]" gridcol="2" gridrow="14" pageId="5" pageNumber="32">3,3</td>
<td id="7692106A450F1A95FB93FADDFB80C5C3" box="[1086,1147,1394,1415]" gridcol="3" gridrow="14" pageId="5" pageNumber="32">2,3</td>
<td id="7692106A450F1A95FB19FADDFB0CC5C3" box="[1204,1271,1394,1415]" gridcol="4" gridrow="14" pageId="5" pageNumber="32">3,3</td>
<td id="7692106A450F1A95FA84FADDFA9DC5C3" box="[1321,1382,1394,1415]" gridcol="5" gridrow="14" pageId="5" pageNumber="32">3,4</td>
<td id="7692106A450F1A95FA32FADDFA20C5C3" box="[1439,1499,1394,1415]" gridcol="6" gridrow="14" pageId="5" pageNumber="32">4,4</td>
<td id="7692106A450F1A95F9B9FADDF9ABC5C3" box="[1556,1616,1394,1415]" gridcol="7" gridrow="14" pageId="5" pageNumber="32">3,3</td>
<td id="7692106A450F1A95F927FADDF93DC5C3" box="[1674,1734,1394,1415]" gridcol="8" gridrow="14" pageId="5" pageNumber="32">3,3</td>
<td id="7692106A450F1A95F952FADDF8B8C5C3" box="[1791,1859,1394,1415]" gridcol="9" gridrow="14" pageId="5" pageNumber="32">3,3</td>
</tr>
<tr id="35437916450F1A95FFD0FA38F8B8C5E8" box="[125,1859,1431,1452]" gridrow="15" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FA38FD93C5E8" box="[125,616,1431,1452]" gridcol="0" gridrow="15" pageId="5" pageNumber="32">No. infralabial shielđs</th>
<td id="7692106A450F1A95FCF9FA38FC5AC5E8" box="[852,929,1431,1452]" gridcol="1" gridrow="15" pageId="5" pageNumber="32">8,8</td>
<td id="7692106A450F1A95FC64FA38FBFEC5E8" box="[969,1029,1431,1452]" gridcol="2" gridrow="15" pageId="5" pageNumber="32">7,7</td>
<td id="7692106A450F1A95FB93FA38FB80C5E8" box="[1086,1147,1431,1452]" gridcol="3" gridrow="15" pageId="5" pageNumber="32">7,7</td>
<td id="7692106A450F1A95FB19FA38FB0CC5E8" box="[1204,1271,1431,1452]" gridcol="4" gridrow="15" pageId="5" pageNumber="32">7,7</td>
<td id="7692106A450F1A95FA84FA38FA9DC5E8" box="[1321,1382,1431,1452]" gridcol="5" gridrow="15" pageId="5" pageNumber="32">8,8</td>
<td id="7692106A450F1A95FA32FA38FA20C5E8" box="[1439,1499,1431,1452]" gridcol="6" gridrow="15" pageId="5" pageNumber="32">7,7</td>
<td id="7692106A450F1A95F9B9FA38F9ABC5E8" box="[1556,1616,1431,1452]" gridcol="7" gridrow="15" pageId="5" pageNumber="32">7,7</td>
<td id="7692106A450F1A95F927FA38F93DC5E8" box="[1674,1734,1431,1452]" gridcol="8" gridrow="15" pageId="5" pageNumber="32">7,7</td>
<td id="7692106A450F1A95F952FA38F8B8C5E8" box="[1791,1859,1431,1452]" gridcol="9" gridrow="15" pageId="5" pageNumber="32">7,7</td>
</tr>
<tr id="35437916450F1A95FFD0FA13F8B8C595" box="[125,1859,1468,1489]" gridrow="16" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FA13FD93C595" box="[125,616,1468,1489]" gridcol="0" gridrow="16" pageId="5" pageNumber="32">No. scales between first V anđ posterior genial</th>
<td id="7692106A450F1A95FCF9FA13FC5AC595" box="[852,929,1468,1489]" gridcol="1" gridrow="16" pageId="5" pageNumber="32">5,5</td>
<td id="7692106A450F1A95FC64FA13FBFEC595" box="[969,1029,1468,1489]" gridcol="2" gridrow="16" pageId="5" pageNumber="32">6,6</td>
<td id="7692106A450F1A95FB93FA13FB80C595" box="[1086,1147,1468,1489]" gridcol="3" gridrow="16" pageId="5" pageNumber="32">5,5</td>
<td id="7692106A450F1A95FB19FA13FB0CC595" box="[1204,1271,1468,1489]" gridcol="4" gridrow="16" pageId="5" pageNumber="32">4,4</td>
<td id="7692106A450F1A95FA84FA13FA9DC595" box="[1321,1382,1468,1489]" gridcol="5" gridrow="16" pageId="5" pageNumber="32">6,5</td>
<td id="7692106A450F1A95FA32FA13FA20C595" box="[1439,1499,1468,1489]" gridcol="6" gridrow="16" pageId="5" pageNumber="32">4,4</td>
<td id="7692106A450F1A95F9B9FA13F9ABC595" box="[1556,1616,1468,1489]" gridcol="7" gridrow="16" pageId="5" pageNumber="32">5,5</td>
<td id="7692106A450F1A95F927FA13F93DC595" box="[1674,1734,1468,1489]" gridcol="8" gridrow="16" pageId="5" pageNumber="32">5,5</td>
<td id="7692106A450F1A95F952FA13F8B8C595" box="[1791,1859,1468,1489]" gridcol="9" gridrow="16" pageId="5" pageNumber="32">5,5</td>
</tr>
<tr id="35437916450F1A95FFD0FA4EF8B8C5B2" box="[125,1859,1505,1526]" gridrow="17" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0FA4EFD93C5B2" box="[125,616,1505,1526]" gridcol="0" gridrow="17" pageId="5" pageNumber="32">No. scales between first V anđ last infralabial</th>
<td id="7692106A450F1A95FCF9FA4EFC5AC5B2" box="[852,929,1505,1526]" gridcol="1" gridrow="17" pageId="5" pageNumber="32">6,6</td>
<td id="7692106A450F1A95FC64FA4EFBFEC5B2" box="[969,1029,1505,1526]" gridcol="2" gridrow="17" pageId="5" pageNumber="32">8,8</td>
<td id="7692106A450F1A95FB93FA4EFB80C5B2" box="[1086,1147,1505,1526]" gridcol="3" gridrow="17" pageId="5" pageNumber="32">5,6</td>
<td id="7692106A450F1A95FB19FA4EFB0CC5B2" box="[1204,1271,1505,1526]" gridcol="4" gridrow="17" pageId="5" pageNumber="32">6,6</td>
<td id="7692106A450F1A95FA84FA4EFA9DC5B2" box="[1321,1382,1505,1526]" gridcol="5" gridrow="17" pageId="5" pageNumber="32">7,7</td>
<td id="7692106A450F1A95FA32FA4EFA20C5B2" box="[1439,1499,1505,1526]" gridcol="6" gridrow="17" pageId="5" pageNumber="32">5,6</td>
<td id="7692106A450F1A95F9B9FA4EF9ABC5B2" box="[1556,1616,1505,1526]" gridcol="7" gridrow="17" pageId="5" pageNumber="32">6,7</td>
<td id="7692106A450F1A95F927FA4EF93DC5B2" box="[1674,1734,1505,1526]" gridcol="8" gridrow="17" pageId="5" pageNumber="32">6,7</td>
<td id="7692106A450F1A95F952FA4EF8B8C5B2" box="[1791,1859,1505,1526]" gridcol="9" gridrow="17" pageId="5" pageNumber="32">6,6</td>
</tr>
<tr id="35437916450F1A95FFD0F9A9F8B8C65F" box="[125,1859,1542,1563]" gridrow="18" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0F9A9FD93C65F" box="[125,616,1542,1563]" gridcol="0" gridrow="18" pageId="5" pageNumber="32">No. scales overlappeđ by anal (excluđing subcauđals)</th>
<td id="7692106A450F1A95FCF9F9A9FC5AC65F" box="[852,929,1542,1563]" gridcol="1" gridrow="18" pageId="5" pageNumber="32">4,6</td>
<td id="7692106A450F1A95FC64F9A9FBFEC65F" box="[969,1029,1542,1563]" gridcol="2" gridrow="18" pageId="5" pageNumber="32">6,5</td>
<td id="7692106A450F1A95FB93F9A9FB80C65F" box="[1086,1147,1542,1563]" gridcol="3" gridrow="18" pageId="5" pageNumber="32">5,5</td>
<td id="7692106A450F1A95FB19F9A9FB0CC65F" box="[1204,1271,1542,1563]" gridcol="4" gridrow="18" pageId="5" pageNumber="32">7,5</td>
<td id="7692106A450F1A95FA84F9A9FA9DC65F" box="[1321,1382,1542,1563]" gridcol="5" gridrow="18" pageId="5" pageNumber="32">6,7</td>
<td id="7692106A450F1A95FA32F9A9FA20C65F" box="[1439,1499,1542,1563]" gridcol="6" gridrow="18" pageId="5" pageNumber="32">5,6</td>
<td id="7692106A450F1A95F9B9F9A9F9ABC65F" box="[1556,1616,1542,1563]" gridcol="7" gridrow="18" pageId="5" pageNumber="32">5,5</td>
<td id="7692106A450F1A95F927F9A9F93DC65F" box="[1674,1734,1542,1563]" gridcol="8" gridrow="18" pageId="5" pageNumber="32">5,4</td>
<td id="7692106A450F1A95F952F9A9F8B8C65F" box="[1791,1859,1542,1563]" gridcol="9" gridrow="18" pageId="5" pageNumber="32">6,5</td>
</tr>
<tr id="35437916450F1A95FFD0F984F8B8C604" box="[125,1859,1579,1600]" gridrow="19" pageId="5" pageNumber="32">
<th id="7692106A450F1A95FFD0F984FD93C604" box="[125,616,1579,1600]" gridcol="0" gridrow="19" pageId="5" pageNumber="32">No. unpigmenteđ posterior ventral scales</th>
<td id="7692106A450F1A95FCF9F984FC5AC604" box="[852,929,1579,1600]" gridcol="1" gridrow="19" pageId="5" pageNumber="32">8</td>
<td id="7692106A450F1A95FC64F984FBFEC604" box="[969,1029,1579,1600]" gridcol="2" gridrow="19" pageId="5" pageNumber="32">9</td>
<td id="7692106A450F1A95FB93F984FB80C604" box="[1086,1147,1579,1600]" gridcol="3" gridrow="19" pageId="5" pageNumber="32">13</td>
<td id="7692106A450F1A95FB19F984FB0CC604" box="[1204,1271,1579,1600]" gridcol="4" gridrow="19" pageId="5" pageNumber="32">15</td>
<td id="7692106A450F1A95FA84F984FA9DC604" box="[1321,1382,1579,1600]" gridcol="5" gridrow="19" pageId="5" pageNumber="32">ca.39</td>
<td id="7692106A450F1A95FA32F984FA20C604" box="[1439,1499,1579,1600]" gridcol="6" gridrow="19" pageId="5" pageNumber="32">7</td>
<td id="7692106A450F1A95F9B9F984F9ABC604" box="[1556,1616,1579,1600]" gridcol="7" gridrow="19" pageId="5" pageNumber="32">0</td>
<td id="7692106A450F1A95F927F984F93DC604" box="[1674,1734,1579,1600]" gridcol="8" gridrow="19" pageId="5" pageNumber="32">4</td>
<td id="7692106A450F1A95F952F984F8B8C604" box="[1791,1859,1579,1600]" gridcol="9" gridrow="19" pageId="5" pageNumber="32">0</td>
</tr>
</table>
</paragraph>
<caption id="DF0C2BDC450CE56CFF3AF810FE6BC7B7" httpUri="https://zenodo.org/record/1051971/files/figure.png" pageId="6" pageNumber="33" targetBox="[198,1364,220,1894]" targetPageId="6">
<paragraph id="8BCC7B54450CE56CFF3AF810FE6BC7B7" blockId="6.[151,1435,1983,2035]" pageId="6" pageNumber="33">
<emphasis id="B907A746450CE56CFF3AF810FEE8C790" bold="true" box="[151,275,1983,2005]" pageId="6" pageNumber="33">FIGURE 3.</emphasis>
Holotype of 
<taxonomicName id="4C7300D7450CE56CFE36F810FD91C791" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[411,618,1983,2005]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="33" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A746450CE56CFE36F810FD91C791" box="[411,618,1983,2005]" italics="true" pageId="6" pageNumber="33">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D450CE56CFDDDF810FD3AC790" box="[624,705,1983,2004]" pageId="6" pageNumber="33" rank="species">
<emphasis id="B907A746450CE56CFDDDF810FD3AC790" bold="true" box="[624,705,1983,2004]" pageId="6" pageNumber="33">sp. nov.</emphasis>
</taxonomicNameLabel>
(NCBS-AU163) whole preserved specimen in A) dorsal and B) ventral views. Scale bar 10 mm.
</paragraph>
</caption>
<caption id="DF0C2BDC450DE56DFF3AF81DFD3DC7A3" httpUri="https://zenodo.org/record/1051973/files/figure.png" pageId="7" pageNumber="34" targetBox="[205,1382,224,1914]" targetPageId="7">
<paragraph id="8BCC7B54450DE56DFF3AF81DFD3DC7A3" blockId="7.[151,1435,1970,2023]" pageId="7" pageNumber="34">
<emphasis id="B907A746450DE56DFF3AF81DFEE9C783" bold="true" box="[151,274,1970,1992]" pageId="7" pageNumber="34">FIGURE 4.</emphasis>
Holotype of 
<taxonomicName id="4C7300D7450DE56DFE3AF81DFD9DC78C" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[407,614,1970,1992]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A746450DE56DFE3AF81DFD9DC78C" box="[407,614,1970,1992]" italics="true" pageId="7" pageNumber="34">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D450DE56DFDC6F81CFD40C78C" box="[619,699,1971,1992]" pageId="7" pageNumber="34" rank="species">
<emphasis id="B907A746450DE56DFDC6F81CFD40C78C" bold="true" box="[619,699,1971,1992]" pageId="7" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
(NCBS-AU163) head of preserved specimen in A) dorsal, B) ventral, C) right lateral and D) left lateral views. Scale bar 10 mm.
</paragraph>
</caption>
<caption id="DF0C2BDC4502E562FF3AFBEFFC38C430" httpUri="https://zenodo.org/record/1051975/files/figure.png" pageId="8" pageNumber="35" targetBox="[157,1430,161,1062]" targetPageId="8">
<paragraph id="8BCC7B544502E562FF3AFBEFFC38C430" blockId="8.[151,1436,1088,1141]" pageId="8" pageNumber="35">
<emphasis id="B907A7464502E562FF3AFBEFFEE9C411" bold="true" box="[151,274,1088,1110]" pageId="8" pageNumber="35">FIGURE 5.</emphasis>
Paratypes of 
<taxonomicName id="4C7300D74502E562FE30FBEFFD91C412" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[413,618,1088,1110]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464502E562FE30FBEFFD91C412" box="[413,618,1088,1110]" italics="true" pageId="8" pageNumber="35">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4502E562FDDCFBEFFD39C411" box="[625,706,1088,1109]" pageId="8" pageNumber="35" rank="species">
<emphasis id="B907A7464502E562FDDCFBEFFD39C411" bold="true" box="[625,706,1088,1109]" pageId="8" pageNumber="35">sp. nov.</emphasis>
</taxonomicNameLabel>
(specimen numbers given in first row). Heads of preserved specimen in A) dorsal, B) ventral, C) right lateral and D) left lateral views. Scale bar 10 mm.
</paragraph>
</caption>
<paragraph id="8BCC7B544502E562FF6AFB0EFF07C5D6" blockId="8.[151,1436,1185,2038]" pageId="8" pageNumber="35">External naris seen as C-shaped slit approximately (very slightly dorsal of) central on nasal. Flap behind slit closes to almost level with surface of rest of nasal such that naris not notably countersunk. Naris visible dorsally, anteriorly, mostly laterally. Five supralabials (SLs) on each side, SL5 largest, SL1, SL2 and SL4 subequal, smallest. SL1 contacts rostral, nasal and loreal, SL2 contacts loreal and lower preocular, SL3 contacts lower pre- and postocular as well as eye, SL4 contacts lower postocular and anterior temporal, SL5 contacts anterior and second (and lower third) temporal. Among SLs, eye contacts SL3 only. Eye dorsolateral, pupil elliptical (long axis vertical).</paragraph>
<paragraph id="8BCC7B544502E562FF6AFA32FB2AC602" blockId="8.[151,1436,1185,2038]" pageId="8" pageNumber="35">Temporals 1+1(+2); second largest, longest; posterior temporals less directly situated between parietal and SL5. Parietals larger than other head scales. Midline interparietal suture much longer than parietal projection behind suture, shorter than frontal. Each parietal contacts frontal, supraocular, upper postocular, anterior, second and upper third temporal plus three other scales. Right parietal more rounded posteriorly, left pointed at a right angle. Inside of mouth pale. Teeth partly hidden by gingivae, 10 or 11 positions on each maxilla.</paragraph>
<paragraph id="8BCC7B544502E562FF6AF9FEFF1AC706" blockId="8.[151,1436,1185,2038]" pageId="8" pageNumber="35">Mental small, subtriangular, wider than long. Infralabials (ILs) asymmetric, 8 on right, 7 on left; first ILs in midline contact; fifth largest. On left, IL2 smallest, IL6 longest, only it and IL7 longer than broad; on right, IL6 smallest, seemingly supernumerary, IL6, IL7 and IL8 longer than broad (IL7 longest). Two pairs of genials; first pair largest, longer than broad, in long midline contact; second pair not in contact, separated anteriorly by one small, midline scale. Anterior genials contact IL1–4; posterior genials contact IL5 only. Anteriormost ventral separated from posterior genials by 5 scales; separated from posteriormost infralabials by 6 (left) and 7 (right) scales.</paragraph>
<paragraph id="8BCC7B544502E562FF6AF8E2FF01C7B2" blockId="8.[151,1436,1185,2038]" pageId="8" pageNumber="35">Macroscopically and under low magnification (using a light dissecting microscope) body scales smooth, iridescent. No keels or apical pits. Vertebral scale row not different from other DSRs. Dorsal body scales generally evenly sized on dorsum and along body except for those involved in dorsal scale row reductions; scales in lower rows more broadly rounded distally; scales closer to vent slightly smaller than at midbody, those at anterior of body smallest.</paragraph>
<paragraph id="8BCC7B544503E563FF6AFF38FF34C120" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="36">Posteriormost ventral substantially narrower, mostly restricted to left; penultimate ventral divided (paired). Anals (right overlapping left) slightly larger than posteriormost ventral. Each anal overlaps 4 (left) and 6 (right) small scales along margin of vent, in addition to first pair of subcaudals. Dorsal scale rows 19 at level of first ventral, reducing to 17 via loss of row 3 or 4 by level of third (right) or fourth (left) ventral. Seventeen rows maintained almost to vent; first or second row lost on the right on row level with fifth and sixth ventral anterior to vent.</paragraph>
<paragraph id="8BCC7B544503E563FF6AFEC0FC76C194" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="36">Tail flattened ventrally. Dorsal tail scales more heterogenous in size than on body in association with scale reductions. Subcaudals paired/divided throughout, first pair notably smaller and less regular than second pair; terminal scute conical, longer than wide, pointed, slightly upturned.</paragraph>
<paragraph id="8BCC7B544503E563FF6AFE73FBAEC2EC" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="36">Macroscopically bicoloured, darker above, border between darker dorsum and paler venter sharply demarcated, generally between first and second dorsal scale row along body, lower and more irregular on tail. In preservation grey, brown and green dorsally and creamy white ventrolaterally. Midventral dark charcoal greybrown (on each ventral more grey anteriorly, more brown posteriorly) stripe along most of body, irregular lateral margins. Posteriormost 10 ventrals and anals pale, without darker midventral stripe; Midventral stripe on tail more irregular, especially posteriorly, where absent (occasional dark blotches) on last third.</paragraph>
<paragraph id="8BCC7B544503E563FF6AFD1BFF0CC38C" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="36">Head dark dorsally, generally uniform, rostral slightly paler. Whitish patches on lower edges of head behind rostral on all supralabials, most notably on most of SL2 and SL3 and on two consecutive body scales behind last supralabial (where continuous with pale ventral surface of body). Underside of head mostly pale; mental and anteriormost four infralabials (and smaller parts of other ILs plus anterior margins of anterior genials) with same grey/brown as midventral stripe of body. Midventral darker stripe of body extends anteriorly as far as onto posteriormost preventral midline scale. Pale stripe between darker dorsum and darker midventral stripe interrupted by occasional, small, irregular darker spots on first dorsal scale row or (much more commonly) lateral edge of ventrals.</paragraph>
<paragraph id="8BCC7B544503E563FF6AFC7BFC90C481" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="36">Darker dorsum of body and tail (along whole length) bears row of fairly evenly spaced, darker grey, irregular spots (two or three, occasionally four scales large), mostly confined to second and third dorsal scale rows. Additional row of dark dots also mostly evenly spaced (generally similarly spaced to lower row, though more closely spaced anteriorly, and alternating with upper row) runs along fifth or (less frequently) sixth dorsal scale row. Upper row of dots mostly one, sometimes two scales large. Much more irregular and occasional darker dots occur approximately middorsally on body and (more faintly) tail. Dark scales on dorsum (including head) are coarsely darkly mottled under low power dissecting microscope.</paragraph>
<paragraph id="8BCC7B544503E563FF6AFB60FAC3C574" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="36">
<emphasis id="B907A7464503E563FF6AFB60FDE6C4AC" bold="true" box="[199,541,1231,1256]" pageId="9" pageNumber="36">
Variation among 
<typeStatus id="54C8C5F64503E563FE0DFB60FDE3C4AC" box="[416,536,1231,1256]" pageId="9" pageNumber="36" type="paratype">paratypes</typeStatus>
.
</emphasis>
See 
<tableCitation id="C6F14EEF4503E563FDF0FB60FD4EC4AC" box="[605,693,1231,1256]" captionStart="TABLE 2" captionStartId="5.[125,202,585,606]" captionText="TABLE 2. Meristic anđ morphometric (in millimetres) đata for examineđ specimens of Rhabdops aquaticus sp. nov. All specimens are paratypes except for the holotype (*) anđ a referređ specimen (**). Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because" pageId="9" pageNumber="36">Table 2</tableCitation>
for variation in meristic and morphometric features. 
<typeStatus id="54C8C5F64503E563FA82FB7EFA67C4AD" box="[1327,1436,1233,1257]" pageId="9" pageNumber="36" type="paratype">Paratypes</typeStatus>
generally in moderate to good condition; all with incisions on anterior midventral part of tail, several with incisions into coelom, NCBS- AU162 with eyes removed, several specimens with one or both hemipenes removed.
</paragraph>
<paragraph id="8BCC7B544503E563FF6AFA92FD27C6DC" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="36">Paratypes typically match holotype description except where noted here. One of smallest paratypes (BNHS 3347) with more laterally compressed (rather than subcylindrical, ventrally flattened) body. Head little wider than body and ovate in dorsal view in all paratypes except subequal and straight-sided in dorsal view in NCBS-AU162. Degree of tapering of anterior of head in lateral view slightly variable. Lateral edges of frontal anteriorly diverging in all paratypes except subparallel in BNHS 3255; frontal about as long as (rather than longer than) distance between it and snout tip in NCBS-AU162. Loreal as large as (BNHS 3234) or slightly larger than nasal (BNHS 1724, 3510) in some paratypes. All paratypes differ from holotype in having larger post- rather than preoculars; relative sizes of upper and lower pre- and postoculars typically as in holotype though postoculars suequal in size in BNHS 3234, right postocular larger on right of NCBS-AU162; lower postoculars more ventrally (as well as posteriorly) situated in for example, NCBS-AU162.</paragraph>
<paragraph id="8BCC7B544503E563FF6AF90BFE5FC709" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="36">Naris slightly anterior as well as slightly dorsal of centre of nasal shield in BNHS 3347. Third and fourth supralabial subequal in BNHS 3509, BNHS 3510 and NCBS-AU162. Interparietal suture and frontal subequal in length in BNHS 3255. Posterior margin of parietals rounded or wavy (broadly scalloped) in all paratypes, with each contacting typically three, rarely two (left side of BNHS 3509) or four (BNHS 1724, right side of BNHS 3234) non-head shields.</paragraph>
<paragraph id="8BCC7B544503E563FF6AF8F8FDEAC799" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="36">All paratypes with two pairs of genials, though BNHS 3234 also with symmetrical (bilaterally paired) small scales behind posterior pair. Paratypes typically resemble holotype in having at least the posteriormost ventral scale divided/paired, though entire in BNHS 3234, NCBS-AU162. Terminal scute on tail not (or barely) always slightly upturned, straight in BNHS 3509.</paragraph>
<caption id="DF0C2BDC4500E560FF3AF81DFC11C7A3" httpUri="https://zenodo.org/record/1051977/files/figure.png" pageId="10" pageNumber="37" targetBox="[174,1413,194,1947]" targetPageId="10">
<paragraph id="8BCC7B544500E560FF3AF81DFC11C7A3" blockId="10.[151,1436,1970,2023]" pageId="10" pageNumber="37">
<emphasis id="B907A7464500E560FF3AF81DFEE8C783" bold="true" box="[151,275,1970,1992]" pageId="10" pageNumber="37">FIGURE 6.</emphasis>
<taxonomicName id="4C7300D74500E560FEB6F81DFE11C78C" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[283,490,1970,1992]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="37" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464500E560FEB6F81DFE11C78C" box="[283,490,1970,1992]" italics="true" pageId="10" pageNumber="37">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4500E560FE5DF81CFDB9C78C" box="[496,578,1971,1992]" pageId="10" pageNumber="37" rank="species">
<emphasis id="B907A7464500E560FE5DF81CFDB9C78C" bold="true" box="[496,578,1971,1992]" pageId="10" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
showing colouration in life of: A) larger (presumably adult) specimen from Baraki, and B) smaller (possibly juvenile) specimen from Amboli. Specimens not collected.
</paragraph>
</caption>
<caption id="DF0C2BDC4501E561FF3AF822FBC2C786" httpUri="https://zenodo.org/record/1051979/files/figure.png" pageId="11" pageNumber="38" targetBox="[198,1403,192,1896]" targetPageId="11">
<paragraph id="8BCC7B544501E561FF3AF822FBC2C786" blockId="11.[151,1436,1933,1986]" pageId="11" pageNumber="38">
<emphasis id="B907A7464501E561FF3AF822FEE9C7E7" bold="true" box="[151,274,1933,1955]" pageId="11" pageNumber="38">FIGURE 7.</emphasis>
A) 
<taxonomicName id="4C7300D74501E561FE94F822FDFCC7E0" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[313,519,1933,1956]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="38" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464501E561FE94F822FDFCC7E0" box="[313,519,1933,1956]" italics="true" pageId="11" pageNumber="38">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4501E561FDA1F821FDA7C7E7" box="[524,604,1934,1955]" pageId="11" pageNumber="38" rank="species">
<emphasis id="B907A7464501E561FDA1F821FDA7C7E7" bold="true" box="[524,604,1934,1955]" pageId="11" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
from Amboli, and B) 
<taxonomicName id="4C7300D74501E561FC93F820FC41C7E7" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[830,954,1933,1956]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="38" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464501E561FC93F820FC41C7E7" box="[830,954,1933,1956]" italics="true" pageId="11" pageNumber="38">R. olivaceus</emphasis>
</taxonomicName>
from Pookode. Note differences in body colour pattern in terms of stripes and spots, and differences in eye colour. Specimens not collected.
</paragraph>
</caption>
<caption id="DF0C2BDC4506E566FF3AF98CFE8DC613" httpUri="https://zenodo.org/record/1051981/files/figure.png" pageId="12" pageNumber="39" targetBox="[172,1416,193,1549]" targetPageId="12">
<paragraph id="8BCC7B544506E566FF3AF98CFE8DC613" blockId="12.[151,1435,1571,1623]" pageId="12" pageNumber="39">
<emphasis id="B907A7464506E566FF3AF98CFEEDC67C" bold="true" box="[151,278,1571,1593]" pageId="12" pageNumber="39">FIGURE 8.</emphasis>
Holotype of 
<taxonomicName id="4C7300D74506E566FE04F98CFD83C67D" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[425,632,1571,1593]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="39" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464506E566FE04F98CFD83C67D" box="[425,632,1571,1593]" italics="true" pageId="12" pageNumber="39">Rhabdops olivaceus</emphasis>
</taxonomicName>
(BMNH 1946.1.12.44, previously 74.4.29.64) in largely dorsal (upper) and ventral (lower) views.
</paragraph>
</caption>
<paragraph id="8BCC7B544506E567FF6AF92BFA7BC0BC" blockId="12.[151,1437,1668,2017]" lastBlockId="13.[151,1437,151,2013]" lastPageId="13" lastPageNumber="40" pageId="12" pageNumber="39">
The border between dark dorsum and pale venter varies slightly, partly between ventrals and first dorsal scale row in BNHS 3347 and 3234 (where border partly on first dorsal scale row); on second (occasionally third) dorsal scale row in BNHS 3255; not lower on tail than on body in BNHS 3347. In BNHS 3347 the lowest dark scale row (first dorsal row) is slightly paler than more dorsal scale rows. Dark spots on upper part of pale venter somewhat variable; mostly restricted to posterior of body in NCBS-AU162; mostly on first dorsal scale row (rather than lateral edge of ventrals) in BNHS 3509; absent in BNHS 1724, 3347, 3255 or 3510. Dark spots on dorsal scale rows 5 and 6 often larger (up to four scales) in BNHS 3234 than in holotype. Venter of tail more variable in pattern; almost entirely dark (NCBS-AU162), darkly blotched or mottled rather than striped (BNHS 3234, 3509, 3510), mostly pale (BNHS 3255) or striped only on anterior half (BNHS 3347). Posteriormost ventrals and anals more typically with than without small spots or flecks of dark pigment. Rostral not paler than rest of head in dorsal view in NCBS-AU162, BNHS 1724, 3347, 3255, 3510. Supralabials always with pale patches though mostly pale in BNHS 1724. Underside of head with slightly less dark pigment in NCBS-AU162 (pigment on first two rather than four infralabials); more extensively pigmented in BNHS 
<quantity id="4C8BD6B14507E567FCB8FF70FC91C0BC" box="[789,874,223,248]" metricMagnitude="1" metricUnit="m" metricValue="8.91286" pageId="13" pageNumber="40" unit="in" value="3509.0">3509 in</quantity>
which anterior genials almost completely dark.
</paragraph>
<paragraph id="8BCC7B544507E567FF6AFEABFE59C120" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="40">
<emphasis id="B907A7464507E567FF6AFEABFE97C159" bold="true" box="[199,364,260,285]" pageId="13" pageNumber="40">Colour in life.</emphasis>
Adults olive brown above with blackish spots, off-white ventrolaterally. Eye orange/red-brown. Smaller, younger animals tend to have a more olive green dorsum, yellow ventrolaterally, and a duller green-greybrown eye (
<figureCitation id="134867D14507E567FEB1FEE3FE90C121" box="[284,363,332,357]" captionStart="FIGURE 6" captionStartId="10.[151,250,1970,1992]" captionTargetBox="[174,1413,194,1947]" captionTargetId="figure@10.[173,1414,193,1949]" captionTargetPageId="10" captionText="FIGURE 6. Rhabdops aquaticus sp. nov. showing colouration in life of: A) larger (presumably adult) specimen from Baraki, and B) smaller (possibly juvenile) specimen from Amboli. Specimens not collected." httpUri="https://zenodo.org/record/1051977/files/figure.png" pageId="13" pageNumber="40">Figs. 6</figureCitation>
, 
<figureCitation id="134867D14507E567FEDAFEE3FE6FC120" box="[375,404,332,356]" captionStart="FIGURE 7" captionStartId="11.[151,250,1933,1955]" captionTargetBox="[198,1403,192,1896]" captionTargetId="figure@11.[198,1403,192,1896]" captionTargetPageId="11" captionText="FIGURE 7. A) Rhabdops aquaticus sp. nov. from Amboli, and B) R. olivaceus from Pookode. Note differences in body colour pattern in terms of stripes and spots, and differences in eye colour. Specimens not collected." httpUri="https://zenodo.org/record/1051979/files/figure.png" pageId="13" pageNumber="40">7a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36928DF4507E567FF6AFEC0FBDCC1B1" pageId="13" pageNumber="40" type="etymology">
<paragraph id="8BCC7B544507E567FF6AFEC0FD8BC194" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="40">
<emphasis id="B907A7464507E567FF6AFEC0FEB5C1CC" bold="true" box="[199,334,367,392]" pageId="13" pageNumber="40">Etymology.</emphasis>
The species name is derived from the Latin for relating to water, 
<taxonomicName id="4C7300D74507E567FBDDFEDEFB26C1CC" authorityName="Gower" authorityYear="2017" box="[1136,1245,369,392]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus">
<emphasis id="B907A7464507E567FBDDFEDEFB26C1CC" box="[1136,1245,369,392]" italics="true" pageId="13" pageNumber="40">aquaticus</emphasis>
</taxonomicName>
, in reference to observations of this snake often being observed in freshwater bodies. For nomenclatural purposes, the species epithet is considered a noun in apposition.
</paragraph>
<paragraph id="8BCC7B544507E567FF6AFE73FBDCC1B1" blockId="13.[151,1437,151,2013]" box="[199,1063,476,501]" pageId="13" pageNumber="40">
<emphasis id="B907A7464507E567FF6AFE73FE07C1B1" bold="true" box="[199,508,476,501]" pageId="13" pageNumber="40">Suggested common name.</emphasis>
Water 
<taxonomicName id="4C7300D74507E567FDE1FE73FD47C1B1" authorityName="Boulenger" authorityYear="1893" box="[588,700,476,501]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="genus">Rhabdops</taxonomicName>
or Aquatic 
<taxonomicName id="4C7300D74507E567FCECFE73FBD8C1B1" authority="(English)" authorityName="English" box="[833,1059,476,501]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="genus">Rhabdops (English)</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36928DF4507E567FF6AFE50FCA7C614" pageId="13" pageNumber="40" type="description">
<paragraph id="8BCC7B544507E567FF6AFE50FC80C3C4" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="40">
<emphasis id="B907A7464507E567FF6AFE50FBFDC25C" bold="true" box="[199,1030,511,536]" pageId="13" pageNumber="40">
Differences between 
<taxonomicName id="4C7300D74507E567FE7CFE50FD3EC25C" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[465,709,511,536]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FE7CFE50FD3EC25C" bold="true" box="[465,709,511,536]" italics="true" pageId="13" pageNumber="40">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FD78FE50FCC2C25C" box="[725,825,511,536]" pageId="13" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
and congeners.
</emphasis>
<taxonomicName id="4C7300D74507E567FBBAFE50FAFEC25C" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1047,1285,511,536]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FBBAFE50FAFEC25C" box="[1047,1285,511,536]" italics="true" pageId="13" pageNumber="40">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FAB8FE50FA82C25C" box="[1301,1401,511,536]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FAB8FE50FA82C25C" bold="true" box="[1301,1401,511,536]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished clearly from the northeast Indian 
<taxonomicName id="4C7300D74507E567FD61FD8AFCB9C279" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[716,834,548,573]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B907A7464507E567FD61FD8AFCB9C279" box="[716,834,548,573]" italics="true" pageId="13" pageNumber="40">R. bicolor</emphasis>
</taxonomicName>
in that the latter has a single (rather than paired) internasal and a single prefrontal. The two species also differ in colour pattern, in that 
<taxonomicName id="4C7300D74507E567FB1CFDE6FADCC224" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[1201,1319,583,608]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B907A7464507E567FB1CFDE6FADCC224" box="[1201,1319,583,608]" italics="true" pageId="13" pageNumber="40">R. bicolor</emphasis>
</taxonomicName>
lacks the prominent dark spots of 
<taxonomicName id="4C7300D74507E567FE0BFDC2FDC9C2C0" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[422,562,621,644]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FE0BFDC2FDC9C2C0" box="[422,562,621,644]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FD95FDC3FD6AC2C1" box="[568,657,620,645]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FD95FDC3FD6AC2C1" bold="true" box="[568,657,620,645]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="4C7300D74507E567FD35FDC3FC80C2C0" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[664,891,620,645]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FD35FDC3FC80C2C0" box="[664,891,620,645]" italics="true" pageId="13" pageNumber="40">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FC2FFDC3FC20C2C1" box="[898,987,620,645]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FC2FFDC3FC20C2C1" bold="true" box="[898,987,620,645]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
is very similar to 
<taxonomicName id="4C7300D74507E567FB0EFDC2FAD0C2C1" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[1187,1323,620,645]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FB0EFDC2FAD0C2C1" box="[1187,1323,620,645]" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
in overall phenotype, including details of scalation (
<tableCitation id="C6F14EEF4507E567FD2CFD20FD1CC2EC" box="[641,743,655,680]" captionStart="TABLE 2" captionStartId="5.[125,202,585,606]" captionText="TABLE 2. Meristic anđ morphometric (in millimetres) đata for examineđ specimens of Rhabdops aquaticus sp. nov. All specimens are paratypes except for the holotype (*) anđ a referređ specimen (**). Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because" pageId="13" pageNumber="40">Tables 2</tableCitation>
, 
<tableCitation id="C6F14EEF4507E567FD55FD3FFCF3C2EC" box="[760,776,656,680]" captionStart="TABLE 3" captionStartId="17.[125,201,497,516]" captionText="TABLE 3. Meristic anđ morphometric (in millimetres) đata for examineđ specimens of Rhabdops olivaceus. Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because of slight đamage to specimen. Invariant characters not recorđeđ here incluđe: number of postoculars (2,2), prefrontals (2), internasals (2); anal shielđs (paiređ, right overlapping left); number of unpigmenteđ ventrals (none). ZSI in this table is an abbreviation of &quot; ZSI / WGRS / IR / v. ¨. Holotype (*) was previously numbeređ BMNH 74.4. 29.64. Blank cell inđicates đata not recorđeđ. Tail incomplete in BNHS 3411. BMNH 1922.5. 25.24 is interpreteđ as lacking preoculars but having loreals that contact the eye ̅ this is baseđ on the ̒ loreals ̓ having the approximately same shape anđ size anđ relationship with the supralabials as the unambiguous loreals of the other specimens (an alternative view might be that this specimen lacks loreals anđ has a single, elongate preocular on each siđe). See Appenđix 2 for localities." pageId="13" pageNumber="40">3</tableCitation>
; Figs. 7,8). Our recognition of 
<taxonomicName id="4C7300D74507E567FBD2FD3EFAEBC2EC" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1151,1296,657,680]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FBD2FD3EFAEBC2EC" box="[1151,1296,657,680]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FAB6FD20FA82C2EC" box="[1307,1401,655,680]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FAB6FD20FA82C2EC" bold="true" box="[1307,1401,655,680]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
as distinct from 
<taxonomicName id="4C7300D74507E567FE99FD1AFE3BC289" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[308,448,692,717]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FE99FD1AFE3BC289" box="[308,448,692,717]" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
is based on substantial and concordant discontinuity in variation in geography, DNA sequences, number of ventrals, pattern and colour. Among the vouchers that we have examined, 
<taxonomicName id="4C7300D74507E567FB49FD76FA89C2B4" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1252,1394,729,752]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FB49FD76FA89C2B4" box="[1252,1394,729,752]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FAD7FD78FF33C351" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FAD7FD78FF33C351" bold="true" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
has more ventral scales than 
<taxonomicName id="4C7300D74507E567FDBAFD52FD5AC351" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[535,673,764,789]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FDBAFD52FD5AC351" box="[535,673,764,789]" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
(224–230, mean 227, SD = 3.94, n = 9 versus 205–217, mean 211, SD = 5.92, n = 8). There are also differences in subcaudal counts (
<tableCitation id="C6F14EEF4507E567FC3BFCB0FC01C37C" box="[918,1018,799,824]" captionStart="TABLE 2" captionStartId="5.[125,202,585,606]" captionText="TABLE 2. Meristic anđ morphometric (in millimetres) đata for examineđ specimens of Rhabdops aquaticus sp. nov. All specimens are paratypes except for the holotype (*) anđ a referređ specimen (**). Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because" pageId="13" pageNumber="40">Tables 2</tableCitation>
, 
<tableCitation id="C6F14EEF4507E567FBA5FC8FFBECC37C" box="[1032,1047,800,824]" captionStart="TABLE 3" captionStartId="17.[125,201,497,516]" captionText="TABLE 3. Meristic anđ morphometric (in millimetres) đata for examineđ specimens of Rhabdops olivaceus. Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because of slight đamage to specimen. Invariant characters not recorđeđ here incluđe: number of postoculars (2,2), prefrontals (2), internasals (2); anal shielđs (paiređ, right overlapping left); number of unpigmenteđ ventrals (none). ZSI in this table is an abbreviation of &quot; ZSI / WGRS / IR / v. ¨. Holotype (*) was previously numbeređ BMNH 74.4. 29.64. Blank cell inđicates đata not recorđeđ. Tail incomplete in BNHS 3411. BMNH 1922.5. 25.24 is interpreteđ as lacking preoculars but having loreals that contact the eye ̅ this is baseđ on the ̒ loreals ̓ having the approximately same shape anđ size anđ relationship with the supralabials as the unambiguous loreals of the other specimens (an alternative view might be that this specimen lacks loreals anđ has a single, elongate preocular on each siđe). See Appenđix 2 for localities." pageId="13" pageNumber="40">3</tableCitation>
), with male 
<taxonomicName id="4C7300D74507E567FB04FC8EFAC3C37C" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1193,1336,801,824]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FB04FC8EFAC3C37C" box="[1193,1336,801,824]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FAEDFCB0FA67C37C" box="[1344,1436,799,824]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FAEDFCB0FA67C37C" bold="true" box="[1344,1436,799,824]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
having 75–81 and male 
<taxonomicName id="4C7300D74507E567FE1FFCEAFDC4C319" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[434,575,836,861]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FE1FFCEAFDC4C319" box="[434,575,836,861]" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
65–74. Uncorrected mt p-distances between 
<taxonomicName id="4C7300D74507E567FBE2FCEAFB24C318" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1103,1247,837,860]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FBE2FCEAFB24C318" box="[1103,1247,837,860]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FB44FCEBFABDC319" box="[1257,1350,836,861]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FB44FCEBFABDC319" bold="true" box="[1257,1350,836,861]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C7300D74507E567FA29FCEAFEFAC3C4" baseAuthorityName="Beddome" baseAuthorityYear="1863" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FA29FCEAFEFAC3C4" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
are as follows: 
<emphasis id="B907A7464507E567FE1FFCC8FE23C3C4" box="[434,472,871,896]" italics="true" pageId="13" pageNumber="40">
16 
<geoCoordinate id="EE471D934507E567FE63FCC8FE23C3C4" box="[462,472,871,896]" direction="south" orientation="latitude" pageId="13" pageNumber="40" precision="5555" value="-1.5">s</geoCoordinate>
</emphasis>
1.5%, 
<emphasis id="B907A7464507E567FD85FCC8FDAAC3C4" box="[552,593,871,896]" italics="true" pageId="13" pageNumber="40">nd4</emphasis>
4.2–4.9%, 
<emphasis id="B907A7464507E567FD7FFCC8FCFAC3C4" box="[722,769,871,896]" italics="true" pageId="13" pageNumber="40">cytb</emphasis>
7.6–7.7%.
</paragraph>
<paragraph id="8BCC7B544507E567FF6AFC23FBF5C549" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="40">
The most immediately obvious external phenotypic difference between 
<taxonomicName id="4C7300D74507E567FC50FC23FB18C3E0" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1021,1251,908,933]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FC50FC23FB18C3E0" box="[1021,1251,908,933]" italics="true" pageId="13" pageNumber="40">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FB40FC23FAB2C3E1" box="[1261,1353,908,933]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FB40FC23FAB2C3E1" bold="true" box="[1261,1353,908,933]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C7300D74507E567FA29FC22FEFAC38C" baseAuthorityName="Beddome" baseAuthorityYear="1863" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FA29FC22FEFAC38C" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
lies in the pattern and colour (
<figureCitation id="134867D14507E567FDF0FC00FD5BC38C" box="[605,672,943,968]" captionStart="FIGURE 6" captionStartId="10.[151,250,1970,1992]" captionTargetBox="[174,1413,194,1947]" captionTargetId="figure@10.[173,1414,193,1949]" captionTargetPageId="10" captionText="FIGURE 6. Rhabdops aquaticus sp. nov. showing colouration in life of: A) larger (presumably adult) specimen from Baraki, and B) smaller (possibly juvenile) specimen from Amboli. Specimens not collected." httpUri="https://zenodo.org/record/1051977/files/figure.png" pageId="13" pageNumber="40">Fig. 6</figureCitation>
&amp; 
<figureCitation id="134867D14507E567FD6BFC1FFD2DC38C" box="[710,726,944,968]" captionStart="FIGURE 7" captionStartId="11.[151,250,1933,1955]" captionTargetBox="[198,1403,192,1896]" captionTargetId="figure@11.[198,1403,192,1896]" captionTargetPageId="11" captionText="FIGURE 7. A) Rhabdops aquaticus sp. nov. from Amboli, and B) R. olivaceus from Pookode. Note differences in body colour pattern in terms of stripes and spots, and differences in eye colour. Specimens not collected." httpUri="https://zenodo.org/record/1051979/files/figure.png" pageId="13" pageNumber="40">7</figureCitation>
). In large (presumably adult) 
<taxonomicName id="4C7300D74507E567FB85FC1EFB4EC38C" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1064,1205,945,968]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FB85FC1EFB4EC38C" box="[1064,1205,945,968]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FB10FC00FAE3C38C" box="[1213,1304,943,968]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FB10FC00FAE3C38C" bold="true" box="[1213,1304,943,968]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
, there is an abrupt demarcation between the darker dorsum and pale venter, with substantial ventrolateral, largely unblemished pale bands bordering a typically narrower and irregular dark midventral stripe, and parts of the supralabials and most of the underside of the head are pale. In 
<taxonomicName id="4C7300D74507E567FD33FBB2FCD3C471" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[670,808,1052,1077]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FD33FBB2FCD3C471" box="[670,808,1052,1077]" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
, in contrast, the supralabials, underside of the head and body are primarily darkly pigmented (
<figureCitation id="134867D14507E567FDE4FB90FD75C41C" box="[585,654,1087,1112]" captionStart="FIGURE 7" captionStartId="11.[151,250,1933,1955]" captionTargetBox="[198,1403,192,1896]" captionTargetId="figure@11.[198,1403,192,1896]" captionTargetPageId="11" captionText="FIGURE 7. A) Rhabdops aquaticus sp. nov. from Amboli, and B) R. olivaceus from Pookode. Note differences in body colour pattern in terms of stripes and spots, and differences in eye colour. Specimens not collected." httpUri="https://zenodo.org/record/1051979/files/figure.png" pageId="13" pageNumber="40">Fig. 7</figureCitation>
b), there are no substantial pale patches, and along the body the dark dorsum ends ventrolaterally at least two narrowly separated, subparallel, narrow dark lines (shared by the lateral edges of the ventrals and lower edges of first dorsal scale row, and the upper edges of the first and bottom edges of the second dorsal scale row: 
<figureCitation id="134867D14507E567FE45FB03FDD4C480" box="[488,559,1196,1221]" captionStart="FIGURE 7" captionStartId="11.[151,250,1933,1955]" captionTargetBox="[198,1403,192,1896]" captionTargetId="figure@11.[198,1403,192,1896]" captionTargetPageId="11" captionText="FIGURE 7. A) Rhabdops aquaticus sp. nov. from Amboli, and B) R. olivaceus from Pookode. Note differences in body colour pattern in terms of stripes and spots, and differences in eye colour. Specimens not collected." httpUri="https://zenodo.org/record/1051979/files/figure.png" pageId="13" pageNumber="40">Fig. 7</figureCitation>
). Dark spots on the dorsal scale rows in 
<taxonomicName id="4C7300D74507E567FBA3FB02FB65C480" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1038,1182,1197,1220]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FBA3FB02FB65C480" box="[1038,1182,1197,1220]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FB0AFB03FAFFC481" box="[1191,1284,1196,1221]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FB0AFB03FAFFC481" bold="true" box="[1191,1284,1196,1221]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
are typically larger. Larger (presumably adult) 
<taxonomicName id="4C7300D74507E567FDA3FB7EFD62C4AC" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[526,665,1233,1256]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FDA3FB7EFD62C4AC" box="[526,665,1233,1256]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FD0DFB60FD02C4AC" box="[672,761,1231,1256]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FD0DFB60FD02C4AC" bold="true" box="[672,761,1231,1256]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
have conspicuously red-orange eyes (
<figureCitation id="134867D14507E567FB0FFB60FB0BC4AC" box="[1186,1264,1231,1256]" captionStart="FIGURE 6" captionStartId="10.[151,250,1970,1992]" captionTargetBox="[174,1413,194,1947]" captionTargetId="figure@10.[173,1414,193,1949]" captionTargetPageId="10" captionText="FIGURE 6. Rhabdops aquaticus sp. nov. showing colouration in life of: A) larger (presumably adult) specimen from Baraki, and B) smaller (possibly juvenile) specimen from Amboli. Specimens not collected." httpUri="https://zenodo.org/record/1051977/files/figure.png" pageId="13" pageNumber="40">Figs. 6</figureCitation>
, 
<figureCitation id="134867D14507E567FB51FB7FFAE2C4AC" box="[1276,1305,1232,1256]" captionStart="FIGURE 7" captionStartId="11.[151,250,1933,1955]" captionTargetBox="[198,1403,192,1896]" captionTargetId="figure@11.[198,1403,192,1896]" captionTargetPageId="11" captionText="FIGURE 7. A) Rhabdops aquaticus sp. nov. from Amboli, and B) R. olivaceus from Pookode. Note differences in body colour pattern in terms of stripes and spots, and differences in eye colour. Specimens not collected." httpUri="https://zenodo.org/record/1051979/files/figure.png" pageId="13" pageNumber="40">7a</figureCitation>
), unlike the only live specimen of 
<taxonomicName id="4C7300D74507E567FE3DFB5AFDE2C549" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[400,537,1268,1293]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FE3DFB5AFDE2C549" box="[400,537,1268,1293]" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
that we have seen a photograph of (
<figureCitation id="134867D14507E567FC03FB5BFC0AC549" box="[942,1009,1268,1293]" captionStart="FIGURE 7" captionStartId="11.[151,250,1933,1955]" captionTargetBox="[198,1403,192,1896]" captionTargetId="figure@11.[198,1403,192,1896]" captionTargetPageId="11" captionText="FIGURE 7. A) Rhabdops aquaticus sp. nov. from Amboli, and B) R. olivaceus from Pookode. Note differences in body colour pattern in terms of stripes and spots, and differences in eye colour. Specimens not collected." httpUri="https://zenodo.org/record/1051979/files/figure.png" pageId="13" pageNumber="40">Fig. 7</figureCitation>
b).
</paragraph>
<paragraph id="8BCC7B544507E567FF6AFAB8FCA7C614" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="40">
Considering all specimens that we have examined, 
<taxonomicName id="4C7300D74507E567FCBAFAB6FC5DC574" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[791,934,1305,1328]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FCBAFAB6FC5DC574" box="[791,934,1305,1328]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FC02FAB8FBF0C574" box="[943,1035,1303,1328]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FC02FAB8FBF0C574" bold="true" box="[943,1035,1303,1328]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
reaches a substantially larger total length than 
<taxonomicName id="4C7300D74507E567FEB6FA92FE5EC511" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[283,421,1340,1365]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FEB6FA92FE5EC511" box="[283,421,1340,1365]" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
(ca. 
<quantity id="4C8BD6B14507E567FE71FA93FDC2C510" box="[476,569,1340,1364]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="40" unit="mm" value="900.0">900 mm</quantity>
versus ca. 
<quantity id="4C8BD6B14507E567FD1BFA93FCECC510" box="[694,791,1340,1365]" metricMagnitude="-1" metricUnit="m" metricValue="5.7" pageId="13" pageNumber="40" unit="mm" value="570.0">570 mm</quantity>
: 
<tableCitation id="C6F14EEF4507E567FC8CFA93FC7AC510" box="[801,897,1340,1365]" captionStart="TABLE 2" captionStartId="5.[125,202,585,606]" captionText="TABLE 2. Meristic anđ morphometric (in millimetres) đata for examineđ specimens of Rhabdops aquaticus sp. nov. All specimens are paratypes except for the holotype (*) anđ a referređ specimen (**). Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because" pageId="13" pageNumber="40">Tables 2</tableCitation>
, 
<tableCitation id="C6F14EEF4507E567FC20FA93FC66C510" box="[909,925,1340,1364]" captionStart="TABLE 3" captionStartId="17.[125,201,497,516]" captionText="TABLE 3. Meristic anđ morphometric (in millimetres) đata for examineđ specimens of Rhabdops olivaceus. Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because of slight đamage to specimen. Invariant characters not recorđeđ here incluđe: number of postoculars (2,2), prefrontals (2), internasals (2); anal shielđs (paiređ, right overlapping left); number of unpigmenteđ ventrals (none). ZSI in this table is an abbreviation of &quot; ZSI / WGRS / IR / v. ¨. Holotype (*) was previously numbeređ BMNH 74.4. 29.64. Blank cell inđicates đata not recorđeđ. Tail incomplete in BNHS 3411. BMNH 1922.5. 25.24 is interpreteđ as lacking preoculars but having loreals that contact the eye ̅ this is baseđ on the ̒ loreals ̓ having the approximately same shape anđ size anđ relationship with the supralabials as the unambiguous loreals of the other specimens (an alternative view might be that this specimen lacks loreals anđ has a single, elongate preocular on each siđe). See Appenđix 2 for localities." pageId="13" pageNumber="40">3</tableCitation>
). 
<bibRefCitation id="EFE206A54507E567FC1EFA93FB6CC510" box="[947,1175,1340,1365]" pageId="13" pageNumber="40" refString="Ganesh, S. R., Chandramouli, S. R. &amp; Gowrishankar, P. (2012) Record lengths of two endemic caenophidian snakes from the Western Ghats Mountains, India. Hamadryad, 36, 44 - 46." type="journal article">Ganesh et al. (2012)</bibRefCitation>
reported a 
<quantity id="4C8BD6B14507E567FABBFA93FA88C510" box="[1302,1395,1340,1365]" metricMagnitude="-1" metricUnit="m" metricValue="9.85" pageId="13" pageNumber="40" unit="mm" value="985.0">985 mm</quantity>
TL 
<taxonomicName id="4C7300D74507E567FF3AFACEFEDBC53C" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[151,288,1375,1400]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FF3AFACEFEDBC53C" box="[151,288,1375,1400]" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
, however, no specimen was collected, and it was from a locality in 
<collectingRegion id="49B7B5B64507E567FBBBFAF0FB72C53C" box="[1046,1161,1375,1400]" country="India" name="Karnataka" pageId="13" pageNumber="40">Karnataka</collectingRegion>
state at the far northerly limit for vouchered records of the species (
<figureCitation id="134867D14507E567FD3EFA2BFD22C5D8" box="[659,729,1412,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1468,1490]" captionTargetBox="[182,1403,275,1428]" captionTargetId="figure@2.[151,1436,253,1446]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of the genus Rhabdops in the Western Ghats region of peninsular India. Circles denote records of Rhabdops olivaceus and squares denote records of Rhabdops aquaticus sp. nov. Green symbols indicates vouchered records and white symbols indicate non-vouchered reports. Arrow indicates the type locality of the respective species. See Appendix 3 for locality details." httpUri="https://zenodo.org/record/1051967/files/figure.png" pageId="13" pageNumber="40">Fig. 1</figureCitation>
). In addition, this large specimen and one other uncollected animal from a nearby locality are reported as having more ventral scales (223 and 230) than known for 
<taxonomicName id="4C7300D74507E567FAB9FA06FA67C584" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[1300,1436,1447,1472]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FAB9FA06FA67C584" box="[1300,1436,1447,1472]" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
vouchers (205–217, see 
<tableCitation id="C6F14EEF4507E567FE0AFA63FE01C5A0" box="[423,506,1484,1509]" captionStart="TABLE 3" captionStartId="17.[125,201,497,516]" captionText="TABLE 3. Meristic anđ morphometric (in millimetres) đata for examineđ specimens of Rhabdops olivaceus. Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because of slight đamage to specimen. Invariant characters not recorđeđ here incluđe: number of postoculars (2,2), prefrontals (2), internasals (2); anal shielđs (paiređ, right overlapping left); number of unpigmenteđ ventrals (none). ZSI in this table is an abbreviation of &quot; ZSI / WGRS / IR / v. ¨. Holotype (*) was previously numbeređ BMNH 74.4. 29.64. Blank cell inđicates đata not recorđeđ. Tail incomplete in BNHS 3411. BMNH 1922.5. 25.24 is interpreteđ as lacking preoculars but having loreals that contact the eye ̅ this is baseđ on the ̒ loreals ̓ having the approximately same shape anđ size anđ relationship with the supralabials as the unambiguous loreals of the other specimens (an alternative view might be that this specimen lacks loreals anđ has a single, elongate preocular on each siđe). See Appenđix 2 for localities." pageId="13" pageNumber="40">Table 3</tableCitation>
) and the larger specimen differs (S.R. Ganesh, pers. comm., 2017) from the colour and patterns documented here for 
<taxonomicName id="4C7300D74507E567FD81FA5EFD41C64C" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[556,698,1519,1544]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464507E567FD81FA5EFD41C64C" box="[556,698,1519,1544]" italics="true" pageId="13" pageNumber="40">R. olivaceus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7300D74507E567FD54FA5EFC71C64C" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[761,906,1521,1544]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FD54FA5EFC71C64C" box="[761,906,1521,1544]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FC3BFA40FC0FC64C" box="[918,1012,1519,1544]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FC3BFA40FC0FC64C" bold="true" box="[918,1012,1519,1544]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
We suggest that the identity of the population(s) from 
<collectingRegion id="49B7B5B64507E567FED4F9BBFE17C669" box="[377,492,1556,1581]" country="India" name="Karnataka" pageId="13" pageNumber="40">Karnataka</collectingRegion>
reported by 
<bibRefCitation id="EFE206A54507E567FD2AF9BBFC83C668" box="[647,888,1556,1581]" pageId="13" pageNumber="40" refString="Ganesh, S. R., Chandramouli, S. R. &amp; Gowrishankar, P. (2012) Record lengths of two endemic caenophidian snakes from the Western Ghats Mountains, India. Hamadryad, 36, 44 - 46." type="journal article">Ganesh et al. (2012)</bibRefCitation>
requires closer investigation (which is already underway: S.R. Ganesh &amp; Gowri Shankar, pers. comm., 2017).
</paragraph>
</subSubSection>
<subSubSection id="C36928DF4507E578FF6AF9F3FBD1C2C1" lastPageId="18" lastPageNumber="45" pageId="13" pageNumber="40" type="distribution">
<paragraph id="8BCC7B544507E567FF6AF9F3FAE6C740" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="40">
<emphasis id="B907A7464507E567FF6AF9F3FCFAC631" bold="true" box="[199,769,1628,1653]" pageId="13" pageNumber="40">Distribution, natural history and conservation.</emphasis>
<taxonomicName id="4C7300D74507E567FCA6F9F3FC0AC630" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[779,1009,1628,1653]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FCA6F9F3FC0AC630" box="[779,1009,1628,1653]" italics="true" pageId="13" pageNumber="40">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FC56F9F3FBACC631" box="[1019,1111,1628,1653]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FC56F9F3FBACC631" bold="true" box="[1019,1111,1628,1653]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
is known from the 
<collectingRegion id="49B7B5B64507E567FAECF9F2FA67C630" box="[1345,1436,1629,1652]" country="Sri Lanka" name="Western" pageId="13" pageNumber="40">Western</collectingRegion>
Ghats region of 
<collectingRegion id="49B7B5B64507E567FEE1F92FFE86C6DC" box="[332,381,1664,1688]" country="India" name="Goa" pageId="13" pageNumber="40">Goa</collectingRegion>
, northernmost 
<collectingRegion id="49B7B5B64507E567FD8BF9D0FD62C6DC" box="[550,665,1663,1688]" country="India" name="Karnataka" pageId="13" pageNumber="40">Karnataka</collectingRegion>
and southern 
<collectingRegion id="49B7B5B64507E567FC9AF9D0FC3DC6DC" box="[823,966,1663,1688]" country="India" name="Maharashtra" pageId="13" pageNumber="40">Maharashtra</collectingRegion>
, from elevations of 
<quantity id="4C8BD6B14507E567FB0BF9D0FACDC6D3" box="[1190,1334,1663,1688]" metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="10.0" metricValueMin="7.5" pageId="13" pageNumber="40" unit="m" value="875.0" valueMax="1000.0" valueMin="750.0">750–1000 m</quantity>
. (
<figureCitation id="134867D14507E567FAE4F9D0FA70C6DC" box="[1353,1419,1663,1688]" captionStart="FIGURE 1" captionStartId="2.[151,250,1468,1490]" captionTargetBox="[182,1403,275,1428]" captionTargetId="figure@2.[151,1436,253,1446]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of the genus Rhabdops in the Western Ghats region of peninsular India. Circles denote records of Rhabdops olivaceus and squares denote records of Rhabdops aquaticus sp. nov. Green symbols indicates vouchered records and white symbols indicate non-vouchered reports. Arrow indicates the type locality of the respective species. See Appendix 3 for locality details." httpUri="https://zenodo.org/record/1051967/files/figure.png" pageId="13" pageNumber="40">Fig. 1</figureCitation>
). The new species has been encountered mostly during the monsoon season, when it is predominantly aquatic or semi-aquatic, seen in or along streams in semi-evergreen forest and waterlogged habitats on lateritic plateaus (
<bibRefCitation id="EFE206A54507E567FF33F943FDE0C741" author="Bhosale" box="[158,539,1772,1797]" pageId="13" pageNumber="40" refString="Bhosale, H. &amp; Joshi, D. (2014) Notes on distribution, natural history and habitat use of a Colubridae snake, Rhabdops olivaceus (Beddome, 1863). Russian Journal of Herpetology, 21, 166 - 168." type="journal article" year="2014">Bhosale &amp; Joshi, 2014; pers. obs.</bibRefCitation>
). We have not encountered the species in forests away from streams.
</paragraph>
<paragraph id="8BCC7B544507E564FF6AF8BFFE7FC1CC" blockId="13.[151,1437,151,2013]" lastBlockId="14.[151,1437,151,2013]" lastPageId="14" lastPageNumber="41" pageId="13" pageNumber="40">
We have examined and identified preserved voucher specimens of 
<taxonomicName id="4C7300D74507E567FC03F8BEFBC2C76C" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[942,1081,1809,1832]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FC03F8BEFBC2C76C" box="[942,1081,1809,1832]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FBEDF8A0FB62C76C" box="[1088,1177,1807,1832]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FBEDF8A0FB62C76C" bold="true" box="[1088,1177,1807,1832]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
from three localities in 
<collectingRegion id="49B7B5B64507E567FF3AF89BFED8C709" box="[151,291,1844,1869]" country="India" name="Maharashtra" pageId="13" pageNumber="40">Maharashtra</collectingRegion>
(Appendix 3). At the 
<typeStatus id="54C8C5F64507E567FDBAF89AFDBCC709" box="[535,583,1845,1869]" pageId="13" pageNumber="40">type</typeStatus>
locality, Amboli, adults have been seen mostly in streams in semi-evergreen forest, and juveniles seen mostly on lateritic plateaus during the monsoon. A few individuals were also encountered in a well close to the village. At Baraki, both adults and juveniles of 
<taxonomicName id="4C7300D74507E567FC03F8D2FBC7C7D0" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[942,1084,1917,1940]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464507E567FC03F8D2FBC7C7D0" box="[942,1084,1917,1940]" italics="true" pageId="13" pageNumber="40">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4507E567FBE9F8D3FB5BC7D1" box="[1092,1184,1916,1941]" pageId="13" pageNumber="40" rank="species">
<emphasis id="B907A7464507E567FBE9F8D3FB5BC7D1" bold="true" box="[1092,1184,1916,1941]" pageId="13" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
were observed on the summit plateau (Fig. 10B), where smaller, greener individuals were seen mostly under rocks, always in moist or waterlogged areas, and larger, browner individuals were encountered mostly in large, seasonal pools, though also during the day in wet soil under rocks. At Koyna the new species has been observed both in streams in the semievergreen forest (Fig. 10A) and on plateaus. Multiple uncollected individuals of 
<taxonomicName id="4C7300D74504E564FBF6FF12FB16C090" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1115,1261,189,212]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FBF6FF12FB16C090" box="[1115,1261,189,212]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FB57FF13FAA2C091" box="[1274,1369,188,213]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FB57FF13FAA2C091" bold="true" box="[1274,1369,188,213]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
were observed also at multiple additional localities in 
<collectingRegion id="49B7B5B64504E564FD41FF70FC83C0BC" box="[748,888,223,248]" country="India" name="Maharashtra" pageId="14" pageNumber="41">Maharashtra</collectingRegion>
state: Idarganj plateau (Radhanagari Wildlife Sanctuary, Kolhapur district), Manoli plateau (near Amba, Kolhapur district), Zolabmi plateau (Chandoli National Park, Sangli district), Koyna (Satara district) and Vankusavade plateau (Satara district) (
<figureCitation id="134867D14504E564FB23FE88FB2FC104" box="[1166,1236,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1468,1490]" captionTargetBox="[182,1403,275,1428]" captionTargetId="figure@2.[151,1436,253,1446]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of the genus Rhabdops in the Western Ghats region of peninsular India. Circles denote records of Rhabdops olivaceus and squares denote records of Rhabdops aquaticus sp. nov. Green symbols indicates vouchered records and white symbols indicate non-vouchered reports. Arrow indicates the type locality of the respective species. See Appendix 3 for locality details." httpUri="https://zenodo.org/record/1051967/files/figure.png" pageId="14" pageNumber="41">Fig. 1</figureCitation>
; Appendix 3). In addition, there are other published records of this species occurring at other localities in the northern 
<collectingRegion id="49B7B5B64504E564FB54FEE2FAAFC120" box="[1273,1364,333,356]" country="Sri Lanka" name="Western" pageId="14" pageNumber="41">Western</collectingRegion>
Ghats (
<figureCitation id="134867D14504E564FF32FEC0FF18C1CC" box="[159,227,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1468,1490]" captionTargetBox="[182,1403,275,1428]" captionTargetId="figure@2.[151,1436,253,1446]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of the genus Rhabdops in the Western Ghats region of peninsular India. Circles denote records of Rhabdops olivaceus and squares denote records of Rhabdops aquaticus sp. nov. Green symbols indicates vouchered records and white symbols indicate non-vouchered reports. Arrow indicates the type locality of the respective species. See Appendix 3 for locality details." httpUri="https://zenodo.org/record/1051967/files/figure.png" pageId="14" pageNumber="41">Fig. 1</figureCitation>
; Appendix 3).
</paragraph>
<paragraph id="8BCC7B544504E564FF6AFE3BFC8AC373" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="41">
<taxonomicName id="4C7300D74504E564FF6AFE3BFE56C1E8" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[199,429,404,429]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FF6AFE3BFE56C1E8" box="[199,429,404,429]" italics="true" pageId="14" pageNumber="41">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FE1BFE3BFDE9C1E9" box="[438,530,404,429]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FE1BFE3BFDE9C1E9" bold="true" box="[438,530,404,429]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
is chiefly nocturnal, when we have observed specimens seemingly foraging in lentic or very slowly flowing water on plateaus and in forest streams. During these observations, 
<taxonomicName id="4C7300D74504E564FB48FE16FA89C194" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1253,1394,441,464]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FB48FE16FA89C194" box="[1253,1394,441,464]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FAD7FE18FF33C1B1" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FAD7FE18FF33C1B1" bold="true" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
flicked their tongues under water and moved among inundated leaf litter or under rocks, seemingly searching for prey. Based on casual, opportunistic observation of five actively foraging individuals in a stream in Koyna, V.B.G. observed that they stayed submerged for 4–12 minutes. We have never observed this assumed foraging behaviour during the day. At Vanakusavade, V.B.G. observed approximately 20 individuals at night, close to a drying steam on the border of a plateau. Most of these individuals were actively foraging in the stream, and a few were under rocks close to the stream. At least as many 
<taxonomicName id="4C7300D74504E564FD57FD20FBE0C2EC" baseAuthorityName="Schneider" baseAuthorityYear="1799" box="[762,1051,655,680]" class="Reptilia" family="Colubridae" genus="Xenochrophis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="piscator">
<emphasis id="B907A7464504E564FD57FD20FC6EC2EC" box="[762,917,655,680]" italics="true" pageId="14" pageNumber="41">Xenochrophis</emphasis>
cf. 
<emphasis id="B907A7464504E564FC12FD3EFBE0C2EC" box="[959,1051,657,680]" italics="true" pageId="14" pageNumber="41">piscator</emphasis>
</taxonomicName>
were also seen in the same habitat on this occasion. During the day we have seen 
<taxonomicName id="4C7300D74504E564FD12FD1AFCB5C288" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[703,846,693,716]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FD12FD1AFCB5C288" box="[703,846,693,716]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FCF5FD1AFC57C288" box="[856,940,693,716]" pageId="14" pageNumber="41" rank="species">sp. nov</taxonomicNameLabel>
mostly under rocks close to streams or on plateaus. In Koyna on 
<date id="FFCD5D944504E564FE39FD77FD8AC2B4" box="[404,625,727,752]" pageId="14" pageNumber="41" value="2010-09-28">28 September 2010</date>
, V.B.G. observed 35 individuals along a 
<quantity id="4C8BD6B14504E564FB94FD78FB7BC2AB" box="[1081,1152,727,752]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="41" unit="m" value="500.0">500 m</quantity>
stretch of a forest stream in four hours of intensive searching at night. During the day, individuals were seen basking on exposed rocks in the same stream, and on approach they moved rapidly into the water.
</paragraph>
<paragraph id="8BCC7B544504E564FF6AFCEBFB40C439" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="41">
The diet of 
<taxonomicName id="4C7300D74504E564FEF8FCEAFE18C319" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[341,483,836,861]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464504E564FEF8FCEAFE18C319" box="[341,483,836,861]" italics="true" pageId="14" pageNumber="41">R. olivaceus</emphasis>
</taxonomicName>
is reported as fish, slugs, earthworms and soft-bodied invertebrates (
<bibRefCitation id="EFE206A54504E564FABCFCEBFECBC3C4" author="Whitaker" pageId="14" pageNumber="41" refString="Whitaker, R. &amp; Captain, A. (2004) Snakes of India: The Field Guide. Draco Books, Chennai, 479 pp." type="book" year="2004">Whitaker &amp; Captain 2004</bibRefCitation>
; 
<bibRefCitation id="EFE206A54504E564FE91FCC8FDDDC3C4" author="Radhakrishnan" box="[316,550,871,896]" pageId="14" pageNumber="41" refString="Radhakrishnan, C. (1997) Notes on a new distributional record and the ecology of Rhabdops olivaceus (Beddome) (Reptilia- Serpentes- Colubridae). Journal of Bombay Natural History Society, 94, 167 - 168." type="journal article" year="1997">Radhakrishnan 1997</bibRefCitation>
; 
<bibRefCitation id="EFE206A54504E564FD9FFCC8FCCEC3C4" author="Srinivasulu" box="[562,821,871,896]" pageId="14" pageNumber="41" refString="Srinivasulu, C., Srinivasulu, B. &amp; Molur, S. (2014) The status and distribution of reptiles in the Western Ghats, India. Conservation Assessment and Management Plan (CAMP). Wildlife Information Liaison Development Society, Coimbatore, Tamil Nadu, 148 pp." type="book" year="2014">
Srinivasulu 
<emphasis id="B907A7464504E564FD14FCC6FD12C3C4" box="[697,745,871,896]" italics="true" pageId="14" pageNumber="41">et al</emphasis>
. 2014
</bibRefCitation>
). However, we can find no primary published report to confirm this, and it is possible that these summaries of prey of 
<taxonomicName id="4C7300D74504E564FC28FC22FBE9C3E1" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[901,1042,908,933]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464504E564FC28FC22FBE9C3E1" box="[901,1042,908,933]" italics="true" pageId="14" pageNumber="41">R. olivaceus</emphasis>
</taxonomicName>
were extended, in part, from, for example, Wall’s (1908, 1912) first-hand report of earthworms in the diet of 
<taxonomicName id="4C7300D74504E564FB99FC1EFB52C38C" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[1076,1193,943,968]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B907A7464504E564FB99FC1EFB52C38C" box="[1076,1193,943,968]" italics="true" pageId="14" pageNumber="41">R. bicolor</emphasis>
</taxonomicName>
. We have no direct observations on the diet of 
<taxonomicName id="4C7300D74504E564FE63FC7AFDA0C3A8" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[462,603,981,1004]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FE63FC7AFDA0C3A8" box="[462,603,981,1004]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FDC9FC7BFD44C3A9" box="[612,703,980,1005]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FDC9FC7BFD44C3A9" bold="true" box="[612,703,980,1005]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
, but remnants of arthropod (seemingly crustacean) exoskeletons were observed in the faeces of one specimen kept captive temporarily for observation and photography. On several occasions, especially on plateaus, the new species was seen in ephemeral pools under rocks directly alongside remnants of freshwater crabs. Specimens of 
<taxonomicName id="4C7300D74504E564FD07FBEEFCC7C41C" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[682,828,1089,1112]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FD07FBEEFCC7C41C" box="[682,828,1089,1112]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FCE5FB90FC5CC41C" box="[840,935,1087,1112]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FCE5FB90FC5CC41C" bold="true" box="[840,935,1087,1112]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
of all sizes are docile when handled, the specimens we observed never made any attempt to bite or displayed any aggressive behaviour.
</paragraph>
<paragraph id="8BCC7B544504E564FF6AFB28FC3DC741" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="41">
Ontogenetic colour change (OCC) occurs in 
<taxonomicName id="4C7300D74504E564FD63FB28FC4DC4E4" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[718,950,1159,1184]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FD63FB28FC4DC4E4" box="[718,950,1159,1184]" italics="true" pageId="14" pageNumber="41">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FC6DFB28FBE7C4E4" box="[960,1052,1159,1184]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FC6DFB28FBE7C4E4" bold="true" box="[960,1052,1159,1184]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
Small (presumably mostly or all juvenile) animals are olive green dorsally and yellow ventrally and ventrolaterally with dull eyes, while larger animals (presumably mostly or all adults) are olive brown dorsally and off-white ventrally and ventrolaterally with red-orange eyes (
<figureCitation id="134867D14504E564FECCFB5BFE52C549" box="[353,425,1268,1293]" captionStart="FIGURE 6" captionStartId="10.[151,250,1970,1992]" captionTargetBox="[174,1413,194,1947]" captionTargetId="figure@10.[173,1414,193,1949]" captionTargetPageId="10" captionText="FIGURE 6. Rhabdops aquaticus sp. nov. showing colouration in life of: A) larger (presumably adult) specimen from Baraki, and B) smaller (possibly juvenile) specimen from Amboli. Specimens not collected." httpUri="https://zenodo.org/record/1051977/files/figure.png" pageId="14" pageNumber="41">Fig. 6</figureCitation>
) and, as far as we know, without sexual dichromatism. Correlations have been found between the dorsal colour pattern of snakes and their antipredatory behaviour (e.g. 
<bibRefCitation id="EFE206A54504E564FBCCFAB8FABEC574" author="Jackson" box="[1121,1349,1303,1328]" pageId="14" pageNumber="41" refString="Jackson, J. F., Ingram, W. &amp; Campbell, H. W. (1976) The dorsal pigmentation pattern of snakes as an antipredator strategy: a multivariate approach. American Naturalist, 110, 1029 - 1053." type="journal article" year="1976">
Jackson 
<emphasis id="B907A7464504E564FB69FAB6FB02C574" box="[1220,1273,1303,1328]" italics="true" pageId="14" pageNumber="41">et al</emphasis>
. 1976
</bibRefCitation>
; 
<bibRefCitation id="EFE206A54504E564FAF9FAB8FF28C511" author="Pough" pageId="14" pageNumber="41" refString="Pough, F. H. (1976) Multiple cryptic effects of crossbanded and ringed patterns of snakes. Copeia, 1976, 834 - 836. https: // doi. org / 10.2307 / 1443481" type="journal article" year="1976">Pough 1976</bibRefCitation>
). Snakes with disruptive patterns tend to rely on concealment and aggression for defence (
<bibRefCitation id="EFE206A54504E564FB6AFA93FAB1C511" author="Creer" box="[1223,1354,1340,1365]" pageId="14" pageNumber="41" refString="Creer, D. (2005) Correlations between ontogenetic change in color pattern and antipredator behavior in the racer, Coluber constrictor. Ethology, 111, 287 - 300. https: // doi. org / 10.1111 / j. 1439 - 0310.2004.01062. x" type="journal article" year="2005">Creer 2005</bibRefCitation>
), while those with stripes, speckles and/or uniform colour are more likely to rely on flight (
<bibRefCitation id="EFE206A54504E564FBE1FAF0FAE9C53C" author="Allen" box="[1100,1298,1375,1400]" pageId="14" pageNumber="41" refString="Allen W. L., Baddeley, R., Scott-Samuel, N. E. &amp; Cuthill, I. C. (2013) The evolution and function of pattern diversity in snakes. Behavioural Ecol ogy, 24, 1237 - 1250. https: // doi. org / 10.1093 / beheco / art 058" type="journal article" year="2013">
Allen 
<emphasis id="B907A7464504E564FB39FACEFB36C53C" box="[1172,1229,1375,1400]" italics="true" pageId="14" pageNumber="41">et al.</emphasis>
2013
</bibRefCitation>
). Given that OCC in 
<taxonomicName id="4C7300D74504E564FF5EFA2AFE87C5D8" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[243,380,1413,1436]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FF5EFA2AFE87C5D8" box="[243,380,1413,1436]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FE29FA2BFE20C5D9" box="[388,475,1412,1437]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FE29FA2BFE20C5D9" bold="true" box="[388,475,1412,1437]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
does not involve a change in basic pattern (darker with dark spots above, paler below), it can be argued that it is not related causally to ontogenetic changes in antipredatory behaviour. Perhaps more likely 
<taxonomicName id="4C7300D74504E564FF3AFA62FEDFC5A0" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[151,292,1485,1508]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FF3AFA62FEDFC5A0" box="[151,292,1485,1508]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FE82FA63FE70C5A1" box="[303,395,1484,1509]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FE82FA63FE70C5A1" bold="true" box="[303,395,1484,1509]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
undergoes ontogenetic changes in habits and/or microhabitats, with the OCC occurring to maintain crypsis (
<bibRefCitation id="EFE206A54504E564FECCFA40FE12C64C" author="Booth" box="[353,489,1519,1544]" pageId="14" pageNumber="41" refString="Booth, C. L. (1990) Evolutionary significance of ontogenetic colour change in animals. Biological Journal of the Linnean Society, 40, 125 - 163. https: // doi. org / 10.1111 / j. 1095 - 8312.1990. tb 01973. x" type="journal article" year="1990">Booth 1990</bibRefCitation>
) and avoid predation (Wilson 
<emphasis id="B907A7464504E564FCE6FA5EFC85C64C" box="[843,894,1519,1544]" italics="true" pageId="14" pageNumber="41">et al</emphasis>
. 2007). Our observations are opportunistic and preliminary, but we believe that smaller individuals of 
<taxonomicName id="4C7300D74504E564FD41F9BAFC8FC668" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[748,884,1557,1580]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FD41F9BAFC8FC668" box="[748,884,1557,1580]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FCD6F9BBFC28C669" box="[891,979,1556,1581]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FCD6F9BBFC28C669" bold="true" box="[891,979,1556,1581]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
are more likely to be seen in waterlogged places or in shallow water with ephemeral vegetation on plateaus during the monsoon, while larger individuals are more often encountered in streams or deep pools with less or no aquatic vegetation. We have seen only a single smaller, greener individual in the forest, that being in a stream close to a plateau. Thus, we speculate that maintaining crypsis during ontogenetic changes in microhabitat use (at least partly) explains OCC in 
<taxonomicName id="4C7300D74504E564FBC4F90AFB09C6F8" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1129,1266,1701,1724]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FBC4F90AFB09C6F8" box="[1129,1266,1701,1724]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FB54F90BFAA9C6F9" box="[1273,1362,1700,1725]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FB54F90BFAA9C6F9" bold="true" box="[1273,1362,1700,1725]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
Given the local abundance of 
<taxonomicName id="4C7300D74504E564FE39F966FDE6C6A4" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[404,541,1737,1760]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FE39F966FDE6C6A4" box="[404,541,1737,1760]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FD89F968FD80C6A4" box="[548,635,1735,1760]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FD89F968FD80C6A4" bold="true" box="[548,635,1735,1760]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
, this species should be amenable to quantitative analyses of microhabitat use, as well as the discovery of, for e.g., the main predators of this species.
</paragraph>
<paragraph id="8BCC7B544504E578FF6AF8A0FCB1C159" blockId="14.[151,1437,151,2013]" lastBlockId="18.[151,1437,151,645]" lastPageId="18" lastPageNumber="45" pageId="14" pageNumber="41">
The IUCN red list (IUCN 2017) contains 
<taxonomicName id="4C7300D74504E564FD07F8A0FC76C76C" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[682,909,1807,1832]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464504E564FD07F8A0FC76C76C" box="[682,909,1807,1832]" italics="true" pageId="14" pageNumber="41">Rhabdops olivaceus</emphasis>
</taxonomicName>
as “Least Concern” by virtue of its extensive distribution. Our recognition and description of 
<taxonomicName id="4C7300D74504E564FD77F89AFC96C708" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[730,877,1845,1868]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FD77F89AFC96C708" box="[730,877,1845,1868]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FCD6F89BFC27C709" box="[891,988,1844,1869]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FCD6F89BFC27C709" bold="true" box="[891,988,1844,1869]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
has substantially reduced the known distribution of 
<taxonomicName id="4C7300D74504E564FEEDF8F6FE30C734" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[320,459,1879,1904]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464504E564FEEDF8F6FE30C734" box="[320,459,1879,1904]" italics="true" pageId="14" pageNumber="41">R. olivaceus</emphasis>
</taxonomicName>
, such that a reassessment is required. Srinavasulu 
<emphasis id="B907A7464504E564FBA0F8F6FBBCC734" box="[1037,1095,1879,1904]" italics="true" pageId="14" pageNumber="41">et al.</emphasis>
(2013) state that 
<taxonomicName id="4C7300D74504E564FABCF8F6FA67C734" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[1297,1436,1879,1904]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464504E564FABCF8F6FA67C734" box="[1297,1436,1879,1904]" italics="true" pageId="14" pageNumber="41">R. olivaceus</emphasis>
</taxonomicName>
“population declines have been reported” and that in 
<collectingRegion id="49B7B5B64504E564FD43F8D3FCE7C7D0" box="[750,796,1916,1940]" country="India" name="Goa" pageId="14" pageNumber="41">Goa</collectingRegion>
and 
<collectingRegion id="49B7B5B64504E564FCF9F8D3FC1BC7D1" box="[852,992,1916,1941]" country="India" name="Maharashtra" pageId="14" pageNumber="41">Maharashtra</collectingRegion>
(thus, 
<taxonomicName id="4C7300D74504E564FB83F8D2FB41C7D0" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1070,1210,1917,1940]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464504E564FB83F8D2FB41C7D0" box="[1070,1210,1917,1940]" italics="true" pageId="14" pageNumber="41">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4504E564FB6FF8D3FAE7C7D1" box="[1218,1308,1916,1941]" pageId="14" pageNumber="41" rank="species">
<emphasis id="B907A7464504E564FB6FF8D3FAE7C7D1" bold="true" box="[1218,1308,1916,1941]" pageId="14" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
rather than 
<taxonomicName id="4C7300D74504E564FF3AF80EFED8C7FD" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[151,291,1952,1977]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="41" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464504E564FF3AF80EFED8C7FD" box="[151,291,1952,1977]" italics="true" pageId="14" pageNumber="41">R. olivaceus</emphasis>
</taxonomicName>
) studies “have reported short-term declines linked to changes in habitat quality. Recent records are lacking from historical localities, and this may be associated with longer-term response to habitat degradation”, but no reports are cited or details provided, and we are unaware of any population estimates or monitoring within the range of either 
<taxonomicName id="4C7300D74518E578FEE3FF12FE20C091" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[334,475,188,213]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="45" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464518E578FEE3FF12FE20C091" box="[334,475,188,213]" italics="true" pageId="18" pageNumber="45">R. olivaceus</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C7300D74518E578FDA4FF12FD62C090" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[521,665,189,212]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="45" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464518E578FDA4FF12FD62C090" box="[521,665,189,212]" italics="true" pageId="18" pageNumber="45">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4518E578FD09FF13FCF9C091" box="[676,770,188,213]" pageId="18" pageNumber="45" rank="species">
<emphasis id="B907A7464518E578FD09FF13FCF9C091" bold="true" box="[676,770,188,213]" pageId="18" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
<bibRefCitation id="EFE206A54518E578FCA0FF13FBD1C090" box="[781,1066,188,213]" pageId="18" pageNumber="45" refString="Srinivasulu, B., Srinivasulu, C., Das, A., Achyuthan, N. S. &amp; Kulkarni, N. U. (2013) Rhabdops olivaceus. The IUCN Red List of Threatened Species 2013: e. T 172649 A 1358911." type="journal article">
Srinivasulu 
<emphasis id="B907A7464518E578FC35FF12FC2FC091" box="[920,980,188,213]" italics="true" pageId="18" pageNumber="45">et al.</emphasis>
(2013)
</bibRefCitation>
report also that 
<taxonomicName id="4C7300D74518E578FB5CFF12FA85C091" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[1265,1406,188,213]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="45" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464518E578FB5CFF12FA85C091" box="[1265,1406,188,213]" italics="true" pageId="18" pageNumber="45">R. olivaceus</emphasis>
</taxonomicName>
is threatened by water pollution and siltation from human activities, but do not specify whether that is in parts of the range that pertain to 
<taxonomicName id="4C7300D74518E578FED3FEAAFDFCC159" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[382,519,260,285]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="45" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464518E578FED3FEAAFDFCC159" box="[382,519,260,285]" italics="true" pageId="18" pageNumber="45">R. olivaceus</emphasis>
</taxonomicName>
and/or 
<taxonomicName id="4C7300D74518E578FDF0FEAAFD12C158" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[605,745,261,284]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="45" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464518E578FDF0FEAAFD12C158" box="[605,745,261,284]" italics="true" pageId="18" pageNumber="45">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4518E578FD5DFEABFCB1C159" box="[752,842,260,285]" pageId="18" pageNumber="45" rank="species">
<emphasis id="B907A7464518E578FD5DFEABFCB1C159" bold="true" box="[752,842,260,285]" pageId="18" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<caption id="DF0C2BDC4505E565FF3AF81DFB56C7A3" httpUri="https://zenodo.org/record/1051983/files/figure.png" pageId="15" pageNumber="42" targetBox="[174,1413,194,1948]" targetPageId="15">
<paragraph id="8BCC7B544505E565FF3AF81DFB56C7A3" blockId="15.[151,1436,1970,2023]" pageId="15" pageNumber="42">
<emphasis id="B907A7464505E565FF3AF81DFEE8C783" bold="true" box="[151,275,1970,1992]" pageId="15" pageNumber="42">FIGURE 9.</emphasis>
Habitat of 
<taxonomicName id="4C7300D74505E565FE2AF81DFDACC78C" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[391,599,1970,1992]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="42" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464505E565FE2AF81DFDACC78C" box="[391,599,1970,1992]" italics="true" pageId="15" pageNumber="42">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4505E565FDF0F81CFD55C78C" box="[605,686,1971,1992]" pageId="15" pageNumber="42" rank="species">
<emphasis id="B907A7464505E565FDF0F81CFD55C78C" bold="true" box="[605,686,1971,1992]" pageId="15" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
during the monsoon. A) Stream in Koyna showing (right of image) more lentic water where snakes were seen. B) Summit plateau at Baraki showing rocks and ephemeral pools.
</paragraph>
</caption>
<paragraph id="8BCC7B54451AE57AFF3AF833FE6FC7B4" pageId="16" pageNumber="43">
<table id="F97389F4451A1A95FF3AF833FA67C7B4" box="[151,1436,1948,2032]" gridcols="5" gridrows="3" pageId="16" pageNumber="43">
<tr id="35437916451A1A95FF3AF833FA67C7F7" box="[151,1436,1948,1971]" gridrow="0" pageId="16" pageNumber="43">
<th id="7692106A451A1A95FF3AF833FD8DC7F7" box="[151,630,1948,1971]" gridcol="0" gridrow="0" pageId="16" pageNumber="43">
<emphasis id="B907A746451AE57AFF3AF833FED9C7F5" bold="true" box="[151,290,1948,1970]" pageId="16" pageNumber="43">FIGURE 10.</emphasis>
<taxonomicName id="4C7300D7451AE57AFE86F833FE00C7F6" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[299,507,1948,1970]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="43" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A746451AE57AFE86F833FE00C7F6" box="[299,507,1948,1970]" italics="true" pageId="16" pageNumber="43">Rhabdops aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D451AE57AFDA9F832FDACC7F6" box="[516,599,1949,1970]" pageId="16" pageNumber="43" rank="species">
<emphasis id="B907A746451AE57AFDA9F832FDACC7F6" bold="true" box="[516,599,1949,1970]" pageId="16" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
in
</th>
<th id="7692106A451A1A95FDD0F833FC79C7F7" box="[637,898,1948,1971]" gridcol="1" gridrow="0" pageId="16" pageNumber="43">life in natural habitat. A)</th>
<th id="7692106A451A1A95FC26F833FB61C7F7" box="[907,1178,1948,1971]" gridcol="2" gridrow="0" pageId="16" pageNumber="43">Larger, browner specimen</th>
<th id="7692106A451A1A95FB0FF833FAB2C7F7" box="[1186,1353,1948,1971]" gridcol="3" gridrow="0" pageId="16" pageNumber="43">in an ephemeral</th>
<th id="7692106A451A1A95FAFFF833FA67C7F7" box="[1362,1436,1948,1971]" gridcol="4" gridrow="0" pageId="16" pageNumber="43">pool in</th>
</tr>
<tr id="35437916451A1A95FF3AF814FA67C795" box="[151,1436,1979,2001]" gridrow="1" pageId="16" pageNumber="43">
<th id="7692106A451A1A95FF3AF814FD8DC795" box="[151,630,1979,2001]" gridcol="0" gridrow="1" pageId="16" pageNumber="43">Baraki, and B) smaller, greener (though soon</th>
<td id="7692106A451A1A95FDD0F814FC79C795" box="[637,898,1979,2001]" gridcol="1" gridrow="1" pageId="16" pageNumber="43">to shed) individual from</td>
<td id="7692106A451A1A95FC26F814FB61C795" box="[907,1178,1979,2001]" gridcol="2" gridrow="1" pageId="16" pageNumber="43">Idarganj in a waterlogged</td>
<td id="7692106A451A1A95FB0FF814FAB2C795" box="[1186,1353,1979,2001]" gridcol="3" gridrow="1" pageId="16" pageNumber="43">area underneath</td>
<td id="7692106A451A1A95FAFFF814FA67C795" box="[1362,1436,1979,2001]" gridcol="4" gridrow="1" pageId="16" pageNumber="43">a rock.</td>
</tr>
<tr id="35437916451A1A95FF3AF875FA67C7B4" box="[151,1436,2010,2032]" gridrow="2" pageId="16" pageNumber="43" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="7692106A451A1A95FF3AF875FD8DC7B4" box="[151,630,2010,2032]" gridcol="0" gridrow="2" pageId="16" pageNumber="43">Specimens not collected.</th>
</tr>
</table>
</paragraph>
<caption id="DF0C2BDC451BE57BFFD0FE5EFA1EC2C2" pageId="17" pageNumber="44">
<paragraph id="8BCC7B54451BE57BFFD0FE5EFA1EC2C2" blockId="17.[125,1883,497,647]" pageId="17" pageNumber="44">
<emphasis id="B907A746451BE57BFFD0FE5EFF1AC240" bold="true" box="[125,225,497,516]" pageId="17" pageNumber="44">TABLE 3.</emphasis>
Meristic anđ morphometric (in millimetres) đata for examineđ specimens of 
<taxonomicName id="4C7300D7451BE57BFC08FE5EFBA7C240" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[933,1116,497,516]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="44" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A746451BE57BFC08FE5EFBA7C240" box="[933,1116,497,516]" italics="true" pageId="17" pageNumber="44">Rhabdops olivaceus</emphasis>
</taxonomicName>
. Numbers separateđ by comma are for left, right siđe. Hyphen (-) inđicates shortest đistance between two points. Values in parentheses likely associateđ with greater error because of slight đamage to specimen. Invariant characters not recorđeđ here incluđe: number of postoculars (2,2), prefrontals (2), internasals (2); anal shielđs (paiređ, right overlapping left); number of unpigmenteđ ventrals (none). ZSI in this table is an abbreviation of &quot;ZSI/WGRS/IR/v.¨. Holotype (*) was previously numbeređ BMNH 74.4.29.64. Blank cell inđicates đata not recorđeđ. Tail incomplete in BNHS 3411. BMNH 1922.5.25.24 is interpreteđ as lacking preoculars but having loreals that contact the eye ̅ this is baseđ on the ̒loreals̓ having the approximately same shape anđ size anđ relationship with the supralabials as the unambiguous loreals of the other specimens (an alternative view might be that this specimen lacks loreals anđ has a single, elongate preocular on each siđe). See Appenđix 2 for localities.
</paragraph>
</caption>
<paragraph id="8BCC7B54451BE57BFFD0FD72F8EFC6CA" pageId="17" pageNumber="44">
<table id="F97389F4451B1A95FFD0FD07F8BFC6CA" box="[125,1860,680,1678]" gridcols="9" gridrows="26" pageId="17" pageNumber="44">
<tr id="35437916451B1A95FFD0FD07F8BFC2B5" box="[125,1860,680,753]" gridrow="0" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FD07FD93C2B5" box="[125,616,680,753]" gridcol="0" gridrow="0" pageId="17" pageNumber="44">Sex</th>
<th id="7692106A451B1A95FC9CFD07FC53C2B5" box="[817,936,680,753]" gridcol="1" gridrow="0" pageId="17" pageNumber="44">*BMNH 1946.1.12.44 M</th>
<th id="7692106A451B1A95FC4BFD07FBD2C2B5" box="[998,1065,680,753]" gridcol="2" gridrow="0" pageId="17" pageNumber="44">NCBS- AU164 M</th>
<th id="7692106A451B1A95FBFAFD07FB61C2B5" box="[1111,1178,680,753]" gridcol="3" gridrow="0" pageId="17" pageNumber="44">NCBS- AU165 M</th>
<th id="7692106A451B1A95FB65FD07FAFEC2B5" box="[1224,1285,680,753]" gridcol="4" gridrow="0" pageId="17" pageNumber="44">BNHS 1723 M</th>
<th id="7692106A451B1A95FA97FD07FA4AC2B5" box="[1338,1457,680,753]" gridcol="5" gridrow="0" pageId="17" pageNumber="44">BMNH 1922.5.25.24 F</th>
<th id="7692106A451B1A95FA42FD07F9ABC2B5" box="[1519,1616,680,753]" gridcol="6" gridrow="0" pageId="17" pageNumber="44">BMNH 82.8.26.31 F</th>
<th id="7692106A451B1A95F92BFD07F941C2B5" box="[1670,1722,680,753]" gridcol="7" gridrow="0" pageId="17" pageNumber="44">ZSI 518 F</th>
<th id="7692106A451B1A95F955FD07F8BFC2B5" box="[1784,1860,680,753]" gridcol="8" gridrow="0" pageId="17" pageNumber="44">BNHS 3411 F</th>
</tr>
<tr id="35437916451B1A95FFD0FCAEF8BFC350" box="[125,1860,769,788]" gridrow="1" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FCAEFD93C350" box="[125,616,769,788]" gridcol="0" gridrow="1" pageId="17" pageNumber="44">Snout tip-vent length length</th>
<td id="7692106A451B1A95FC9CFCAEFC53C350" box="[817,936,769,788]" gridcol="1" gridrow="1" pageId="17" pageNumber="44">422</td>
<td id="7692106A451B1A95FC4BFCAEFBD2C350" box="[998,1065,769,788]" gridcol="2" gridrow="1" pageId="17" pageNumber="44">480</td>
<td id="7692106A451B1A95FBFAFCAEFB61C350" box="[1111,1178,769,788]" gridcol="3" gridrow="1" pageId="17" pageNumber="44">420</td>
<td id="7692106A451B1A95FB65FCAEFAFEC350" box="[1224,1285,769,788]" gridcol="4" gridrow="1" pageId="17" pageNumber="44">325</td>
<td id="7692106A451B1A95FA97FCAEFA4AC350" box="[1338,1457,769,788]" gridcol="5" gridrow="1" pageId="17" pageNumber="44">668</td>
<td id="7692106A451B1A95FA42FCAEF9ABC350" box="[1519,1616,769,788]" gridcol="6" gridrow="1" pageId="17" pageNumber="44">447</td>
<td id="7692106A451B1A95F92BFCAEF941C350" box="[1670,1722,769,788]" gridcol="7" gridrow="1" pageId="17" pageNumber="44">468</td>
<td id="7692106A451B1A95F955FCAEF8BFC350" box="[1784,1860,769,788]" gridcol="8" gridrow="1" pageId="17" pageNumber="44">330</td>
</tr>
<tr id="35437916451B1A95FFD0FC8BF8BFC37C" box="[125,1860,804,824]" gridrow="2" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FC8BFD93C37C" box="[125,616,804,824]" gridcol="0" gridrow="2" pageId="17" pageNumber="44">Tail length</th>
<td id="7692106A451B1A95FC9CFC8BFC53C37C" box="[817,936,804,824]" gridcol="1" gridrow="2" pageId="17" pageNumber="44">98</td>
<td id="7692106A451B1A95FC4BFC8BFBD2C37C" box="[998,1065,804,824]" gridcol="2" gridrow="2" pageId="17" pageNumber="44">90</td>
<td id="7692106A451B1A95FBFAFC8BFB61C37C" box="[1111,1178,804,824]" gridcol="3" gridrow="2" pageId="17" pageNumber="44">94</td>
<td id="7692106A451B1A95FB65FC8BFAFEC37C" box="[1224,1285,804,824]" gridcol="4" gridrow="2" pageId="17" pageNumber="44">65</td>
<td id="7692106A451B1A95FA97FC8BFA4AC37C" box="[1338,1457,804,824]" gridcol="5" gridrow="2" pageId="17" pageNumber="44">117</td>
<td id="7692106A451B1A95FA42FC8BF9ABC37C" box="[1519,1616,804,824]" gridcol="6" gridrow="2" pageId="17" pageNumber="44">95</td>
<td id="7692106A451B1A95F92BFC8BF941C37C" box="[1670,1722,804,824]" gridcol="7" gridrow="2" pageId="17" pageNumber="44">89</td>
<td id="7692106A451B1A95F955FC8BF8BFC37C" box="[1784,1860,804,824]" gridcol="8" gridrow="2" pageId="17" pageNumber="44">65+</td>
</tr>
<tr id="35437916451B1A95FFD0FCE7F8BFC31F" box="[125,1860,840,859]" gridrow="3" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FCE7FD93C31F" box="[125,616,840,859]" gridcol="0" gridrow="3" pageId="17" pageNumber="44">Total ventral scales (Dowling methođ)</th>
<td id="7692106A451B1A95FC9CFCE7FC53C31F" box="[817,936,840,859]" gridcol="1" gridrow="3" pageId="17" pageNumber="44">214</td>
<td id="7692106A451B1A95FC4BFCE7FBD2C31F" box="[998,1065,840,859]" gridcol="2" gridrow="3" pageId="17" pageNumber="44">219</td>
<td id="7692106A451B1A95FBFAFCE7FB61C31F" box="[1111,1178,840,859]" gridcol="3" gridrow="3" pageId="17" pageNumber="44">217</td>
<td id="7692106A451B1A95FB65FCE7FAFEC31F" box="[1224,1285,840,859]" gridcol="4" gridrow="3" pageId="17" pageNumber="44">212</td>
<td id="7692106A451B1A95FA97FCE7FA4AC31F" box="[1338,1457,840,859]" gridcol="5" gridrow="3" pageId="17" pageNumber="44">205</td>
<td id="7692106A451B1A95FA42FCE7F9ABC31F" box="[1519,1616,840,859]" gridcol="6" gridrow="3" pageId="17" pageNumber="44">205</td>
<td id="7692106A451B1A95F92BFCE7F941C31F" box="[1670,1722,840,859]" gridcol="7" gridrow="3" pageId="17" pageNumber="44">217</td>
<td id="7692106A451B1A95F955FCE7F8BFC31F" box="[1784,1860,840,859]" gridcol="8" gridrow="3" pageId="17" pageNumber="44">210</td>
</tr>
<tr id="35437916451B1A95FFD0FCC4F8BFC33B" box="[125,1860,875,895]" gridrow="4" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FCC4FD93C33B" box="[125,616,875,895]" gridcol="0" gridrow="4" pageId="17" pageNumber="44">Total subcauđal scales</th>
<td id="7692106A451B1A95FC9CFCC4FC53C33B" box="[817,936,875,895]" gridcol="1" gridrow="4" pageId="17" pageNumber="44">72,72</td>
<td id="7692106A451B1A95FC4BFCC4FBD2C33B" box="[998,1065,875,895]" gridcol="2" gridrow="4" pageId="17" pageNumber="44">72,72</td>
<td id="7692106A451B1A95FBFAFCC4FB61C33B" box="[1111,1178,875,895]" gridcol="3" gridrow="4" pageId="17" pageNumber="44">74,74</td>
<td id="7692106A451B1A95FB65FCC4FAFEC33B" box="[1224,1285,875,895]" gridcol="4" gridrow="4" pageId="17" pageNumber="44">65,65</td>
<td id="7692106A451B1A95FA97FCC4FA4AC33B" box="[1338,1457,875,895]" gridcol="5" gridrow="4" pageId="17" pageNumber="44">63,63</td>
<td id="7692106A451B1A95FA42FCC4F9ABC33B" box="[1519,1616,875,895]" gridcol="6" gridrow="4" pageId="17" pageNumber="44">67,68</td>
<td id="7692106A451B1A95F92BFCC4F941C33B" box="[1670,1722,875,895]" gridcol="7" gridrow="4" pageId="17" pageNumber="44">65,65</td>
<td id="7692106A451B1A95F955FCC4F8BFC33B" box="[1784,1860,875,895]" gridcol="8" gridrow="4" pageId="17" pageNumber="44">34+,35+</td>
</tr>
<tr id="35437916451B1A95FFD0FC20F8BFC3E7" box="[125,1860,911,931]" gridrow="5" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FC20FD93C3E7" box="[125,616,911,931]" gridcol="0" gridrow="5" pageId="17" pageNumber="44">Heađ length (snout tip to back of manđible)</th>
<td id="7692106A451B1A95FC9CFC20FC53C3E7" box="[817,936,911,931]" gridcol="1" gridrow="5" pageId="17" pageNumber="44">11.1</td>
<td id="7692106A451B1A95FC4BFC20FBD2C3E7" box="[998,1065,911,931]" gridcol="2" gridrow="5" pageId="17" pageNumber="44">12.5</td>
<td id="7692106A451B1A95FBFAFC20FB61C3E7" box="[1111,1178,911,931]" gridcol="3" gridrow="5" pageId="17" pageNumber="44">10.2</td>
<td id="7692106A451B1A95FB65FC20FAFEC3E7" box="[1224,1285,911,931]" gridcol="4" gridrow="5" pageId="17" pageNumber="44">9.4</td>
<td id="7692106A451B1A95FA97FC20FA4AC3E7" box="[1338,1457,911,931]" gridcol="5" gridrow="5" pageId="17" pageNumber="44">15.8</td>
<td id="7692106A451B1A95FA42FC20F9ABC3E7" box="[1519,1616,911,931]" gridcol="6" gridrow="5" pageId="17" pageNumber="44">13.3</td>
<td id="7692106A451B1A95F92BFC20F941C3E7" box="[1670,1722,911,931]" gridcol="7" gridrow="5" pageId="17" pageNumber="44">11.2</td>
<td id="7692106A451B1A95F955FC20F8BFC3E7" box="[1784,1860,911,931]" gridcol="8" gridrow="5" pageId="17" pageNumber="44">10.6</td>
</tr>
<tr id="35437916451B1A95FFD0FC1CF8BFC382" box="[125,1860,947,966]" gridrow="6" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FC1CFD93C382" box="[125,616,947,966]" gridcol="0" gridrow="6" pageId="17" pageNumber="44">Maximum heađ wiđth</th>
<td id="7692106A451B1A95FC9CFC1CFC53C382" box="[817,936,947,966]" gridcol="1" gridrow="6" pageId="17" pageNumber="44">7.5</td>
<td id="7692106A451B1A95FC4BFC1CFBD2C382" box="[998,1065,947,966]" gridcol="2" gridrow="6" pageId="17" pageNumber="44">8.4</td>
<td id="7692106A451B1A95FBFAFC1CFB61C382" box="[1111,1178,947,966]" gridcol="3" gridrow="6" pageId="17" pageNumber="44">7.9</td>
<td id="7692106A451B1A95FB65FC1CFAFEC382" box="[1224,1285,947,966]" gridcol="4" gridrow="6" pageId="17" pageNumber="44">6.9</td>
<td id="7692106A451B1A95FA97FC1CFA4AC382" box="[1338,1457,947,966]" gridcol="5" gridrow="6" pageId="17" pageNumber="44">9.6</td>
<td id="7692106A451B1A95FA42FC1CF9ABC382" box="[1519,1616,947,966]" gridcol="6" gridrow="6" pageId="17" pageNumber="44">8.5</td>
<td id="7692106A451B1A95F92BFC1CF941C382" box="[1670,1722,947,966]" gridcol="7" gridrow="6" pageId="17" pageNumber="44">8.4</td>
<td id="7692106A451B1A95F955FC1CF8BFC382" box="[1784,1860,947,966]" gridcol="8" gridrow="6" pageId="17" pageNumber="44">8.4</td>
</tr>
<tr id="35437916451B1A95FFD0FC79F8BFC3AE" box="[125,1860,982,1002]" gridrow="7" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FC79FD93C3AE" box="[125,616,982,1002]" gridcol="0" gridrow="7" pageId="17" pageNumber="44">Circumference at miđbođy</th>
<td id="7692106A451B1A95FC9CFC79FC53C3AE" box="[817,936,982,1002]" gridcol="1" gridrow="7" pageId="17" pageNumber="44">29</td>
<td id="7692106A451B1A95FC4BFC79FBD2C3AE" box="[998,1065,982,1002]" gridcol="2" gridrow="7" pageId="17" pageNumber="44">28</td>
<td id="7692106A451B1A95FBFAFC79FB61C3AE" box="[1111,1178,982,1002]" gridcol="3" gridrow="7" pageId="17" pageNumber="44">(31)</td>
<td id="7692106A451B1A95FB65FC79FAFEC3AE" box="[1224,1285,982,1002]" gridcol="4" gridrow="7" pageId="17" pageNumber="44">25</td>
<td id="7692106A451B1A95FA97FC79FA4AC3AE" box="[1338,1457,982,1002]" gridcol="5" gridrow="7" pageId="17" pageNumber="44">53</td>
<td id="7692106A451B1A95FA42FC79F9ABC3AE" box="[1519,1616,982,1002]" gridcol="6" gridrow="7" pageId="17" pageNumber="44">38</td>
<td id="7692106A451B1A95F92BFC79F941C3AE" box="[1670,1722,982,1002]" gridcol="7" gridrow="7" pageId="17" pageNumber="44">38</td>
<td id="7692106A451B1A95F955FC79F8BFC3AE" box="[1784,1860,982,1002]" gridcol="8" gridrow="7" pageId="17" pageNumber="44">35</td>
</tr>
<tr id="35437916451B1A95FFD0FC55F8BFC449" box="[125,1860,1018,1037]" gridrow="8" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FC55FD93C449" box="[125,616,1018,1037]" gridcol="0" gridrow="8" pageId="17" pageNumber="44">Nostril-nostril</th>
<td id="7692106A451B1A95FC9CFC55FC53C449" box="[817,936,1018,1037]" gridcol="1" gridrow="8" pageId="17" pageNumber="44">3.1</td>
<td id="7692106A451B1A95FC4BFC55FBD2C449" box="[998,1065,1018,1037]" gridcol="2" gridrow="8" pageId="17" pageNumber="44">3.8</td>
<td id="7692106A451B1A95FBFAFC55FB61C449" box="[1111,1178,1018,1037]" gridcol="3" gridrow="8" pageId="17" pageNumber="44">3.3</td>
<td id="7692106A451B1A95FB65FC55FAFEC449" box="[1224,1285,1018,1037]" gridcol="4" gridrow="8" pageId="17" pageNumber="44">3.2</td>
<td id="7692106A451B1A95FA97FC55FA4AC449" box="[1338,1457,1018,1037]" gridcol="5" gridrow="8" pageId="17" pageNumber="44">4.4</td>
<td id="7692106A451B1A95FA42FC55F9ABC449" box="[1519,1616,1018,1037]" gridcol="6" gridrow="8" pageId="17" pageNumber="44">3.7</td>
<td id="7692106A451B1A95F92BFC55F941C449" box="[1670,1722,1018,1037]" gridcol="7" gridrow="8" pageId="17" pageNumber="44">3.4</td>
<td id="7692106A451B1A95F955FC55F8BFC449" box="[1784,1860,1018,1037]" gridcol="8" gridrow="8" pageId="17" pageNumber="44">3.6</td>
</tr>
<tr id="35437916451B1A95FFD0FBB1F8BFC475" box="[125,1860,1054,1073]" gridrow="9" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FBB1FD93C475" box="[125,616,1054,1073]" gridcol="0" gridrow="9" pageId="17" pageNumber="44">Eye-eye</th>
<td id="7692106A451B1A95FC9CFBB1FC53C475" box="[817,936,1054,1073]" gridcol="1" gridrow="9" pageId="17" pageNumber="44">4.9</td>
<td id="7692106A451B1A95FC4BFBB1FBD2C475" box="[998,1065,1054,1073]" gridcol="2" gridrow="9" pageId="17" pageNumber="44">5.7</td>
<td id="7692106A451B1A95FBFAFBB1FB61C475" box="[1111,1178,1054,1073]" gridcol="3" gridrow="9" pageId="17" pageNumber="44">5.2</td>
<td id="7692106A451B1A95FB65FBB1FAFEC475" box="[1224,1285,1054,1073]" gridcol="4" gridrow="9" pageId="17" pageNumber="44">4.8</td>
<td id="7692106A451B1A95FA97FBB1FA4AC475" box="[1338,1457,1054,1073]" gridcol="5" gridrow="9" pageId="17" pageNumber="44">6.7</td>
<td id="7692106A451B1A95FA42FBB1F9ABC475" box="[1519,1616,1054,1073]" gridcol="6" gridrow="9" pageId="17" pageNumber="44">5.5</td>
<td id="7692106A451B1A95F92BFBB1F941C475" box="[1670,1722,1054,1073]" gridcol="7" gridrow="9" pageId="17" pageNumber="44">4.8</td>
<td id="7692106A451B1A95F955FBB1F8BFC475" box="[1784,1860,1054,1073]" gridcol="8" gridrow="9" pageId="17" pageNumber="44">5.5</td>
</tr>
<tr id="35437916451B1A95FFD0FBEEF8BFC411" box="[125,1860,1089,1109]" gridrow="10" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FBEEFD93C411" box="[125,616,1089,1109]" gridcol="0" gridrow="10" pageId="17" pageNumber="44">Eye-snout tip</th>
<td id="7692106A451B1A95FC9CFBEEFC53C411" box="[817,936,1089,1109]" gridcol="1" gridrow="10" pageId="17" pageNumber="44">4.0</td>
<td id="7692106A451B1A95FC4BFBEEFBD2C411" box="[998,1065,1089,1109]" gridcol="2" gridrow="10" pageId="17" pageNumber="44">5.1</td>
<td id="7692106A451B1A95FBFAFBEEFB61C411" box="[1111,1178,1089,1109]" gridcol="3" gridrow="10" pageId="17" pageNumber="44">4.1</td>
<td id="7692106A451B1A95FB65FBEEFAFEC411" box="[1224,1285,1089,1109]" gridcol="4" gridrow="10" pageId="17" pageNumber="44">4.2</td>
<td id="7692106A451B1A95FA97FBEEFA4AC411" box="[1338,1457,1089,1109]" gridcol="5" gridrow="10" pageId="17" pageNumber="44">6.0</td>
<td id="7692106A451B1A95FA42FBEEF9ABC411" box="[1519,1616,1089,1109]" gridcol="6" gridrow="10" pageId="17" pageNumber="44">4.7</td>
<td id="7692106A451B1A95F92BFBEEF941C411" box="[1670,1722,1089,1109]" gridcol="7" gridrow="10" pageId="17" pageNumber="44">4.6</td>
<td id="7692106A451B1A95F955FBEEF8BFC411" box="[1784,1860,1089,1109]" gridcol="8" gridrow="10" pageId="17" pageNumber="44">5.0</td>
</tr>
<tr id="35437916451B1A95FFD0FBCAF8BFC43C" box="[125,1860,1125,1144]" gridrow="11" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FBCAFD93C43C" box="[125,616,1125,1144]" gridcol="0" gridrow="11" pageId="17" pageNumber="44">Diameter of exposeđ part of eye</th>
<td id="7692106A451B1A95FC9CFBCAFC53C43C" box="[817,936,1125,1144]" gridcol="1" gridrow="11" pageId="17" pageNumber="44">1.5</td>
<td id="7692106A451B1A95FC4BFBCAFBD2C43C" box="[998,1065,1125,1144]" gridcol="2" gridrow="11" pageId="17" pageNumber="44">1.8</td>
<td id="7692106A451B1A95FBFAFBCAFB61C43C" box="[1111,1178,1125,1144]" gridcol="3" gridrow="11" pageId="17" pageNumber="44">1.6</td>
<td id="7692106A451B1A95FB65FBCAFAFEC43C" box="[1224,1285,1125,1144]" gridcol="4" gridrow="11" pageId="17" pageNumber="44">1.7</td>
<td id="7692106A451B1A95FA97FBCAFA4AC43C" box="[1338,1457,1125,1144]" gridcol="5" gridrow="11" pageId="17" pageNumber="44">2.0</td>
<td id="7692106A451B1A95FA42FBCAF9ABC43C" box="[1519,1616,1125,1144]" gridcol="6" gridrow="11" pageId="17" pageNumber="44">1.6</td>
<td id="7692106A451B1A95F92BFBCAF941C43C" box="[1670,1722,1125,1144]" gridcol="7" gridrow="11" pageId="17" pageNumber="44">1.4</td>
<td id="7692106A451B1A95F955FBCAF8BFC43C" box="[1784,1860,1125,1144]" gridcol="8" gridrow="11" pageId="17" pageNumber="44">2.0</td>
</tr>
<tr id="35437916451B1A95FFD0FB27F8BFC4D8" box="[125,1860,1160,1180]" gridrow="12" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FB27FD93C4D8" box="[125,616,1160,1180]" gridcol="0" gridrow="12" pageId="17" pageNumber="44">Maximum wiđth of rostral</th>
<td id="7692106A451B1A95FC9CFB27FC53C4D8" box="[817,936,1160,1180]" gridcol="1" gridrow="12" pageId="17" pageNumber="44">3.5</td>
<td id="7692106A451B1A95FC4BFB27FBD2C4D8" box="[998,1065,1160,1180]" gridcol="2" gridrow="12" pageId="17" pageNumber="44">4.3</td>
<td id="7692106A451B1A95FBFAFB27FB61C4D8" box="[1111,1178,1160,1180]" gridcol="3" gridrow="12" pageId="17" pageNumber="44">3.8</td>
<td id="7692106A451B1A95FB65FB27FAFEC4D8" box="[1224,1285,1160,1180]" gridcol="4" gridrow="12" pageId="17" pageNumber="44">3.4</td>
<td id="7692106A451B1A95FA97FB27FA4AC4D8" box="[1338,1457,1160,1180]" gridcol="5" gridrow="12" pageId="17" pageNumber="44">4.4</td>
<td id="7692106A451B1A95FA42FB27F9ABC4D8" box="[1519,1616,1160,1180]" gridcol="6" gridrow="12" pageId="17" pageNumber="44">3.9</td>
<td id="7692106A451B1A95F92BFB27F941C4D8" box="[1670,1722,1160,1180]" gridcol="7" gridrow="12" pageId="17" pageNumber="44">3.6</td>
<td id="7692106A451B1A95F955FB27F8BFC4D8" box="[1784,1860,1160,1180]" gridcol="8" gridrow="12" pageId="17" pageNumber="44">3.5</td>
</tr>
<tr id="35437916451B1A95FFD0FB03F8BFC4FB" box="[125,1860,1196,1215]" gridrow="13" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FB03FD93C4FB" box="[125,616,1196,1215]" gridcol="0" gridrow="13" pageId="17" pageNumber="44">Maximum length of frontal</th>
<td id="7692106A451B1A95FC9CFB03FC53C4FB" box="[817,936,1196,1215]" gridcol="1" gridrow="13" pageId="17" pageNumber="44">4.0</td>
<td id="7692106A451B1A95FC4BFB03FBD2C4FB" box="[998,1065,1196,1215]" gridcol="2" gridrow="13" pageId="17" pageNumber="44">4.6</td>
<td id="7692106A451B1A95FBFAFB03FB61C4FB" box="[1111,1178,1196,1215]" gridcol="3" gridrow="13" pageId="17" pageNumber="44">3.9</td>
<td id="7692106A451B1A95FB65FB03FAFEC4FB" box="[1224,1285,1196,1215]" gridcol="4" gridrow="13" pageId="17" pageNumber="44">3.6</td>
<td id="7692106A451B1A95FA97FB03FA4AC4FB" box="[1338,1457,1196,1215]" gridcol="5" gridrow="13" pageId="17" pageNumber="44">5.2</td>
<td id="7692106A451B1A95FA42FB03F9ABC4FB" box="[1519,1616,1196,1215]" gridcol="6" gridrow="13" pageId="17" pageNumber="44">4.0</td>
<td id="7692106A451B1A95F92BFB03F941C4FB" box="[1670,1722,1196,1215]" gridcol="7" gridrow="13" pageId="17" pageNumber="44">3.5</td>
<td id="7692106A451B1A95F955FB03F8BFC4FB" box="[1784,1860,1196,1215]" gridcol="8" gridrow="13" pageId="17" pageNumber="44">4.3</td>
</tr>
<tr id="35437916451B1A95FFD0FB7FF8BFC4A7" box="[125,1860,1232,1251]" gridrow="14" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FB7FFD93C4A7" box="[125,616,1232,1251]" gridcol="0" gridrow="14" pageId="17" pageNumber="44">Maximum wiđth of frontal</th>
<td id="7692106A451B1A95FC9CFB7FFC53C4A7" box="[817,936,1232,1251]" gridcol="1" gridrow="14" pageId="17" pageNumber="44">3.3</td>
<td id="7692106A451B1A95FC4BFB7FFBD2C4A7" box="[998,1065,1232,1251]" gridcol="2" gridrow="14" pageId="17" pageNumber="44">3.3</td>
<td id="7692106A451B1A95FBFAFB7FFB61C4A7" box="[1111,1178,1232,1251]" gridcol="3" gridrow="14" pageId="17" pageNumber="44">3.6</td>
<td id="7692106A451B1A95FB65FB7FFAFEC4A7" box="[1224,1285,1232,1251]" gridcol="4" gridrow="14" pageId="17" pageNumber="44">3.1</td>
<td id="7692106A451B1A95FA97FB7FFA4AC4A7" box="[1338,1457,1232,1251]" gridcol="5" gridrow="14" pageId="17" pageNumber="44">4.6</td>
<td id="7692106A451B1A95FA42FB7FF9ABC4A7" box="[1519,1616,1232,1251]" gridcol="6" gridrow="14" pageId="17" pageNumber="44">3.5</td>
<td id="7692106A451B1A95F92BFB7FF941C4A7" box="[1670,1722,1232,1251]" gridcol="7" gridrow="14" pageId="17" pageNumber="44">3.3</td>
<td id="7692106A451B1A95F955FB7FF8BFC4A7" box="[1784,1860,1232,1251]" gridcol="8" gridrow="14" pageId="17" pageNumber="44">3.7</td>
</tr>
<tr id="35437916451B1A95FFD0FB5CF8BFC542" box="[125,1860,1267,1286]" gridrow="15" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FB5CFD93C542" box="[125,616,1267,1286]" gridcol="0" gridrow="15" pageId="17" pageNumber="44">Maximum length of parietal</th>
<td id="7692106A451B1A95FC9CFB5CFC53C542" box="[817,936,1267,1286]" gridcol="1" gridrow="15" pageId="17" pageNumber="44">5.4</td>
<td id="7692106A451B1A95FC4BFB5CFBD2C542" box="[998,1065,1267,1286]" gridcol="2" gridrow="15" pageId="17" pageNumber="44">6.3</td>
<td id="7692106A451B1A95FBFAFB5CFB61C542" box="[1111,1178,1267,1286]" gridcol="3" gridrow="15" pageId="17" pageNumber="44">5.5</td>
<td id="7692106A451B1A95FB65FB5CFAFEC542" box="[1224,1285,1267,1286]" gridcol="4" gridrow="15" pageId="17" pageNumber="44">4.9</td>
<td id="7692106A451B1A95FA97FB5CFA4AC542" box="[1338,1457,1267,1286]" gridcol="5" gridrow="15" pageId="17" pageNumber="44">7.6</td>
<td id="7692106A451B1A95FA42FB5CF9ABC542" box="[1519,1616,1267,1286]" gridcol="6" gridrow="15" pageId="17" pageNumber="44">6.0</td>
<td id="7692106A451B1A95F92BFB5CF941C542" box="[1670,1722,1267,1286]" gridcol="7" gridrow="15" pageId="17" pageNumber="44">5.7</td>
<td id="7692106A451B1A95F955FB5CF8BFC542" box="[1784,1860,1267,1286]" gridcol="8" gridrow="15" pageId="17" pageNumber="44">5.7</td>
</tr>
<tr id="35437916451B1A95FFD0FAB8F8BFC56E" box="[125,1860,1303,1322]" gridrow="16" pageId="17" pageNumber="44" rowspan-7="1">
<th id="7692106A451B1A95FFD0FAB8FD93C56E" box="[125,616,1303,1322]" gridcol="0" gridrow="16" pageId="17" pageNumber="44">No. scales contacting parietal (excluđing heađ shielđs)</th>
<td id="7692106A451B1A95FC9CFAB8FC53C56E" box="[817,936,1303,1322]" gridcol="1" gridrow="16" pageId="17" pageNumber="44">2,2</td>
<td id="7692106A451B1A95FC4BFAB8FBD2C56E" box="[998,1065,1303,1322]" gridcol="2" gridrow="16" pageId="17" pageNumber="44">4,3</td>
<td id="7692106A451B1A95FBFAFAB8FB61C56E" box="[1111,1178,1303,1322]" gridcol="3" gridrow="16" pageId="17" pageNumber="44">2,4</td>
<td id="7692106A451B1A95FB65FAB8FAFEC56E" box="[1224,1285,1303,1322]" gridcol="4" gridrow="16" pageId="17" pageNumber="44">3,3</td>
<td id="7692106A451B1A95FA97FAB8FA4AC56E" box="[1338,1457,1303,1322]" gridcol="5" gridrow="16" pageId="17" pageNumber="44">4,5</td>
<td id="7692106A451B1A95FA42FAB8F9ABC56E" box="[1519,1616,1303,1322]" gridcol="6" gridrow="16" pageId="17" pageNumber="44">3,4</td>
<td id="7692106A451B1A95F955FAB8F8BFC56E" box="[1784,1860,1303,1322]" gridcol="8" gridrow="16" pageId="17" pageNumber="44">4,3</td>
</tr>
<tr id="35437916451B1A95FFD0FA95F8BFC50A" box="[125,1860,1338,1358]" gridrow="17" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FA95FD93C50A" box="[125,616,1338,1358]" gridcol="0" gridrow="17" pageId="17" pageNumber="44">No. preocular shielđs</th>
<td id="7692106A451B1A95FC9CFA95FC53C50A" box="[817,936,1338,1358]" gridcol="1" gridrow="17" pageId="17" pageNumber="44">2,2</td>
<td id="7692106A451B1A95FC4BFA95FBD2C50A" box="[998,1065,1338,1358]" gridcol="2" gridrow="17" pageId="17" pageNumber="44">2,2</td>
<td id="7692106A451B1A95FBFAFA95FB61C50A" box="[1111,1178,1338,1358]" gridcol="3" gridrow="17" pageId="17" pageNumber="44">1,2</td>
<td id="7692106A451B1A95FB65FA95FAFEC50A" box="[1224,1285,1338,1358]" gridcol="4" gridrow="17" pageId="17" pageNumber="44">2,2</td>
<td id="7692106A451B1A95FA97FA95FA4AC50A" box="[1338,1457,1338,1358]" gridcol="5" gridrow="17" pageId="17" pageNumber="44">0,0</td>
<td id="7692106A451B1A95FA42FA95F9ABC50A" box="[1519,1616,1338,1358]" gridcol="6" gridrow="17" pageId="17" pageNumber="44">1,2</td>
<td id="7692106A451B1A95F92BFA95F941C50A" box="[1670,1722,1338,1358]" gridcol="7" gridrow="17" pageId="17" pageNumber="44">2,2</td>
<td id="7692106A451B1A95F955FA95F8BFC50A" box="[1784,1860,1338,1358]" gridcol="8" gridrow="17" pageId="17" pageNumber="44">2,2</td>
</tr>
<tr id="35437916451B1A95FFD0FAF1F8BFC535" box="[125,1860,1374,1393]" gridrow="18" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FAF1FD93C535" box="[125,616,1374,1393]" gridcol="0" gridrow="18" pageId="17" pageNumber="44">Scales surrounđing eye</th>
<td id="7692106A451B1A95FC9CFAF1FC53C535" box="[817,936,1374,1393]" gridcol="1" gridrow="18" pageId="17" pageNumber="44">6,6</td>
<td id="7692106A451B1A95FC4BFAF1FBD2C535" box="[998,1065,1374,1393]" gridcol="2" gridrow="18" pageId="17" pageNumber="44">6,6</td>
<td id="7692106A451B1A95FBFAFAF1FB61C535" box="[1111,1178,1374,1393]" gridcol="3" gridrow="18" pageId="17" pageNumber="44">5,6</td>
<td id="7692106A451B1A95FB65FAF1FAFEC535" box="[1224,1285,1374,1393]" gridcol="4" gridrow="18" pageId="17" pageNumber="44">6,6</td>
<td id="7692106A451B1A95FA97FAF1FA4AC535" box="[1338,1457,1374,1393]" gridcol="5" gridrow="18" pageId="17" pageNumber="44">6,6</td>
<td id="7692106A451B1A95FA42FAF1F9ABC535" box="[1519,1616,1374,1393]" gridcol="6" gridrow="18" pageId="17" pageNumber="44">6,6</td>
<td id="7692106A451B1A95F92BFAF1F941C535" box="[1670,1722,1374,1393]" gridcol="7" gridrow="18" pageId="17" pageNumber="44">6,6</td>
<td id="7692106A451B1A95F955FAF1F8BFC535" box="[1784,1860,1374,1393]" gridcol="8" gridrow="18" pageId="17" pageNumber="44">6,6</td>
</tr>
<tr id="35437916451B1A95FFD0FA2EF8BFC5D1" box="[125,1860,1409,1429]" gridrow="19" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FA2EFD93C5D1" box="[125,616,1409,1429]" gridcol="0" gridrow="19" pageId="17" pageNumber="44">Supralabial in contact with eye</th>
<td id="7692106A451B1A95FC9CFA2EFC53C5D1" box="[817,936,1409,1429]" gridcol="1" gridrow="19" pageId="17" pageNumber="44">3,3</td>
<td id="7692106A451B1A95FC4BFA2EFBD2C5D1" box="[998,1065,1409,1429]" gridcol="2" gridrow="19" pageId="17" pageNumber="44">3,3</td>
<td id="7692106A451B1A95FBFAFA2EFB61C5D1" box="[1111,1178,1409,1429]" gridcol="3" gridrow="19" pageId="17" pageNumber="44">3,3</td>
<td id="7692106A451B1A95FB65FA2EFAFEC5D1" box="[1224,1285,1409,1429]" gridcol="4" gridrow="19" pageId="17" pageNumber="44">3,3</td>
<td id="7692106A451B1A95FA97FA2EFA4AC5D1" box="[1338,1457,1409,1429]" gridcol="5" gridrow="19" pageId="17" pageNumber="44">3,3</td>
<td id="7692106A451B1A95FA42FA2EF9ABC5D1" box="[1519,1616,1409,1429]" gridcol="6" gridrow="19" pageId="17" pageNumber="44">2,3</td>
<td id="7692106A451B1A95F92BFA2EF941C5D1" box="[1670,1722,1409,1429]" gridcol="7" gridrow="19" pageId="17" pageNumber="44">3,3</td>
<td id="7692106A451B1A95F955FA2EF8BFC5D1" box="[1784,1860,1409,1429]" gridcol="8" gridrow="19" pageId="17" pageNumber="44">3,3</td>
</tr>
<tr id="35437916451B1A95FFD0FA0AF8BFC5FD" box="[125,1860,1445,1465]" gridrow="20" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FA0AFD93C5FD" box="[125,616,1445,1465]" gridcol="0" gridrow="20" pageId="17" pageNumber="44">Supralabials in contact with each loreal</th>
<td id="7692106A451B1A95FC9CFA0AFC53C5FD" box="[817,936,1445,1465]" gridcol="1" gridrow="20" pageId="17" pageNumber="44">1&amp;2</td>
<td id="7692106A451B1A95FC4BFA0AFBD2C5FD" box="[998,1065,1445,1465]" gridcol="2" gridrow="20" pageId="17" pageNumber="44">1&amp;2</td>
<td id="7692106A451B1A95FBFAFA0AFB61C5FD" box="[1111,1178,1445,1465]" gridcol="3" gridrow="20" pageId="17" pageNumber="44">1&amp;2</td>
<td id="7692106A451B1A95FB65FA0AFAFEC5FD" box="[1224,1285,1445,1465]" gridcol="4" gridrow="20" pageId="17" pageNumber="44">1&amp;2</td>
<td id="7692106A451B1A95FA97FA0AFA4AC5FD" box="[1338,1457,1445,1465]" gridcol="5" gridrow="20" pageId="17" pageNumber="44">1,2&amp;3</td>
<td id="7692106A451B1A95FA42FA0AF9ABC5FD" box="[1519,1616,1445,1465]" gridcol="6" gridrow="20" pageId="17" pageNumber="44">1&amp;2</td>
<td id="7692106A451B1A95F92BFA0AF941C5FD" box="[1670,1722,1445,1465]" gridcol="7" gridrow="20" pageId="17" pageNumber="44">1&amp;2</td>
<td id="7692106A451B1A95F955FA0AF8BFC5FD" box="[1784,1860,1445,1465]" gridcol="8" gridrow="20" pageId="17" pageNumber="44">1&amp;2</td>
</tr>
<tr id="35437916451B1A95FFD0FA66F8BFC598" box="[125,1860,1481,1500]" gridrow="21" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FA66FD93C598" box="[125,616,1481,1500]" gridcol="0" gridrow="21" pageId="17" pageNumber="44">No. supralabial shielđs</th>
<td id="7692106A451B1A95FC9CFA66FC53C598" box="[817,936,1481,1500]" gridcol="1" gridrow="21" pageId="17" pageNumber="44">5,5</td>
<td id="7692106A451B1A95FC4BFA66FBD2C598" box="[998,1065,1481,1500]" gridcol="2" gridrow="21" pageId="17" pageNumber="44">5,5</td>
<td id="7692106A451B1A95FBFAFA66FB61C598" box="[1111,1178,1481,1500]" gridcol="3" gridrow="21" pageId="17" pageNumber="44">5,5</td>
<td id="7692106A451B1A95FB65FA66FAFEC598" box="[1224,1285,1481,1500]" gridcol="4" gridrow="21" pageId="17" pageNumber="44">5,5</td>
<td id="7692106A451B1A95FA97FA66FA4AC598" box="[1338,1457,1481,1500]" gridcol="5" gridrow="21" pageId="17" pageNumber="44">5,5</td>
<td id="7692106A451B1A95FA42FA66F9ABC598" box="[1519,1616,1481,1500]" gridcol="6" gridrow="21" pageId="17" pageNumber="44">4,5</td>
<td id="7692106A451B1A95F92BFA66F941C598" box="[1670,1722,1481,1500]" gridcol="7" gridrow="21" pageId="17" pageNumber="44">5,5</td>
<td id="7692106A451B1A95F955FA66F8BFC598" box="[1784,1860,1481,1500]" gridcol="8" gridrow="21" pageId="17" pageNumber="44">5,5</td>
</tr>
<tr id="35437916451B1A95FFD0FA43F8BFC644" box="[125,1860,1516,1536]" gridrow="22" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0FA43FD93C644" box="[125,616,1516,1536]" gridcol="0" gridrow="22" pageId="17" pageNumber="44">No. infralabial shielđs</th>
<td id="7692106A451B1A95FC9CFA43FC53C644" box="[817,936,1516,1536]" gridcol="1" gridrow="22" pageId="17" pageNumber="44">7,7</td>
<td id="7692106A451B1A95FC4BFA43FBD2C644" box="[998,1065,1516,1536]" gridcol="2" gridrow="22" pageId="17" pageNumber="44">7,7</td>
<td id="7692106A451B1A95FBFAFA43FB61C644" box="[1111,1178,1516,1536]" gridcol="3" gridrow="22" pageId="17" pageNumber="44">7,7</td>
<td id="7692106A451B1A95FB65FA43FAFEC644" box="[1224,1285,1516,1536]" gridcol="4" gridrow="22" pageId="17" pageNumber="44">7,7</td>
<td id="7692106A451B1A95FA97FA43FA4AC644" box="[1338,1457,1516,1536]" gridcol="5" gridrow="22" pageId="17" pageNumber="44">7,7</td>
<td id="7692106A451B1A95FA42FA43F9ABC644" box="[1519,1616,1516,1536]" gridcol="6" gridrow="22" pageId="17" pageNumber="44">7,6</td>
<td id="7692106A451B1A95F92BFA43F941C644" box="[1670,1722,1516,1536]" gridcol="7" gridrow="22" pageId="17" pageNumber="44">7,7</td>
<td id="7692106A451B1A95F955FA43F8BFC644" box="[1784,1860,1516,1536]" gridcol="8" gridrow="22" pageId="17" pageNumber="44">7,7</td>
</tr>
<tr id="35437916451B1A95FFD0F9BFF8BFC603" box="[125,1860,1552,1607]" gridrow="23" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0F9BFFD93C603" box="[125,616,1552,1607]" gridcol="0" gridrow="23" pageId="17" pageNumber="44">Largest infralabial shielđ No. scales between first V anđ posterior genial</th>
<td id="7692106A451B1A95FC9CF9BFFC53C603" box="[817,936,1552,1607]" gridcol="1" gridrow="23" pageId="17" pageNumber="44">5,5 4,4</td>
<td id="7692106A451B1A95FC4BF9BFFBD2C603" box="[998,1065,1552,1607]" gridcol="2" gridrow="23" pageId="17" pageNumber="44">5,5 4,4</td>
<td id="7692106A451B1A95FBFAF9BFFB61C603" box="[1111,1178,1552,1607]" gridcol="3" gridrow="23" pageId="17" pageNumber="44">5,5 4,4</td>
<td id="7692106A451B1A95FB65F9BFFAFEC603" box="[1224,1285,1552,1607]" gridcol="4" gridrow="23" pageId="17" pageNumber="44">5,5 4,4</td>
<td id="7692106A451B1A95FA97F9BFFA4AC603" box="[1338,1457,1552,1607]" gridcol="5" gridrow="23" pageId="17" pageNumber="44">5,5 4,5</td>
<td id="7692106A451B1A95FA42F9BFF9ABC603" box="[1519,1616,1552,1607]" gridcol="6" gridrow="23" pageId="17" pageNumber="44">5,4 5,5</td>
<td id="7692106A451B1A95F92BF9BFF941C603" box="[1670,1722,1552,1607]" gridcol="7" gridrow="23" pageId="17" pageNumber="44">5,5 4,4</td>
<td id="7692106A451B1A95F955F9BFF8BFC603" box="[1784,1860,1552,1607]" gridcol="8" gridrow="23" pageId="17" pageNumber="44">5,5 4,4</td>
</tr>
<tr id="35437916451B1A95FFD0F9F8F8BFC62F" box="[125,1860,1623,1643]" gridrow="24" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0F9F8FD93C62F" box="[125,616,1623,1643]" gridcol="0" gridrow="24" pageId="17" pageNumber="44">No. scales between first V anđ last infralabial</th>
<td id="7692106A451B1A95FC9CF9F8FC53C62F" box="[817,936,1623,1643]" gridcol="1" gridrow="24" pageId="17" pageNumber="44">6,5</td>
<td id="7692106A451B1A95FC4BF9F8FBD2C62F" box="[998,1065,1623,1643]" gridcol="2" gridrow="24" pageId="17" pageNumber="44">6,6</td>
<td id="7692106A451B1A95FBFAF9F8FB61C62F" box="[1111,1178,1623,1643]" gridcol="3" gridrow="24" pageId="17" pageNumber="44">5,6</td>
<td id="7692106A451B1A95FB65F9F8FAFEC62F" box="[1224,1285,1623,1643]" gridcol="4" gridrow="24" pageId="17" pageNumber="44">6,6</td>
<td id="7692106A451B1A95FA97F9F8FA4AC62F" box="[1338,1457,1623,1643]" gridcol="5" gridrow="24" pageId="17" pageNumber="44">5,6</td>
<td id="7692106A451B1A95FA42F9F8F9ABC62F" box="[1519,1616,1623,1643]" gridcol="6" gridrow="24" pageId="17" pageNumber="44">6,6</td>
<td id="7692106A451B1A95F92BF9F8F941C62F" box="[1670,1722,1623,1643]" gridcol="7" gridrow="24" pageId="17" pageNumber="44">6,6</td>
<td id="7692106A451B1A95F955F9F8F8BFC62F" box="[1784,1860,1623,1643]" gridcol="8" gridrow="24" pageId="17" pageNumber="44">5,5</td>
</tr>
<tr id="35437916451B1A95FFD0F9D4F8BFC6CA" box="[125,1860,1659,1678]" gridrow="25" pageId="17" pageNumber="44">
<th id="7692106A451B1A95FFD0F9D4FD93C6CA" box="[125,616,1659,1678]" gridcol="0" gridrow="25" pageId="17" pageNumber="44">No. scales overlappeđ by anal (excluđing subcauđals)</th>
<td id="7692106A451B1A95FC9CF9D4FC53C6CA" box="[817,936,1659,1678]" gridcol="1" gridrow="25" pageId="17" pageNumber="44">4,5</td>
<td id="7692106A451B1A95FC4BF9D4FBD2C6CA" box="[998,1065,1659,1678]" gridcol="2" gridrow="25" pageId="17" pageNumber="44">6,5</td>
<td id="7692106A451B1A95FBFAF9D4FB61C6CA" box="[1111,1178,1659,1678]" gridcol="3" gridrow="25" pageId="17" pageNumber="44">7,6</td>
<td id="7692106A451B1A95FB65F9D4FAFEC6CA" box="[1224,1285,1659,1678]" gridcol="4" gridrow="25" pageId="17" pageNumber="44">5,5</td>
<td id="7692106A451B1A95FA97F9D4FA4AC6CA" box="[1338,1457,1659,1678]" gridcol="5" gridrow="25" pageId="17" pageNumber="44">5,5</td>
<td id="7692106A451B1A95FA42F9D4F9ABC6CA" box="[1519,1616,1659,1678]" gridcol="6" gridrow="25" pageId="17" pageNumber="44">4,4</td>
<td id="7692106A451B1A95F92BF9D4F941C6CA" box="[1670,1722,1659,1678]" gridcol="7" gridrow="25" pageId="17" pageNumber="44">4,4</td>
<td id="7692106A451B1A95F955F9D4F8BFC6CA" box="[1784,1860,1659,1678]" gridcol="8" gridrow="25" pageId="17" pageNumber="44">4,4</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCC7B544518E578FF6AFE88FBD1C2C1" blockId="18.[151,1437,151,645]" pageId="18" pageNumber="45">
Although the species that 
<taxonomicName id="4C7300D74518E578FE55FE86FD7CC104" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[504,647,297,320]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="45" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464518E578FE55FE86FD7CC104" box="[504,647,297,320]" italics="true" pageId="18" pageNumber="45">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4518E578FD3FFE88FD14C104" box="[658,751,295,320]" pageId="18" pageNumber="45" rank="species">
<emphasis id="B907A7464518E578FD3FFE88FD14C104" bold="true" box="[658,751,295,320]" pageId="18" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
was previously mistakenly identified as (
<taxonomicName id="4C7300D74518E578FB7BFE86FA98C104" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[1238,1379,295,320]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="45" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B907A7464518E578FB7BFE86FA98C104" box="[1238,1379,295,320]" italics="true" pageId="18" pageNumber="45">R. olivaceus</emphasis>
</taxonomicName>
) has rarely been collected or reported, the new species is not uncommon at many of its known localities. Localities with vouchers of 
<taxonomicName id="4C7300D74518E578FE89FEDEFE4AC1CC" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[292,433,369,392]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="45" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464518E578FE89FEDEFE4AC1CC" box="[292,433,369,392]" italics="true" pageId="18" pageNumber="45">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4518E578FE17FEC0FDEEC1CC" box="[442,533,367,392]" pageId="18" pageNumber="45" rank="species">
<emphasis id="B907A7464518E578FE17FEC0FDEEC1CC" bold="true" box="[442,533,367,392]" pageId="18" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
occur along a ca. 
<quantity id="4C8BD6B14518E578FD4AFEDFFCC6C1CC" box="[743,829,367,392]" metricMagnitude="5" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="45" unit="km" value="170.0">170 km</quantity>
stretch of the 
<collectingRegion id="49B7B5B64518E578FC4EFEDFFBC5C1C3" box="[995,1086,368,391]" country="Sri Lanka" name="Western" pageId="18" pageNumber="45">Western</collectingRegion>
Ghats, with additional records (Srinavasulu 
<emphasis id="B907A7464518E578FE86FE3AFEA0C1E9" box="[299,347,404,429]" italics="true" pageId="18" pageNumber="45">et al</emphasis>
. 2014: see Appendix 3 and 
<figureCitation id="134867D14518E578FD3FFE3BFD2AC1E8" box="[658,721,404,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1468,1490]" captionTargetBox="[182,1403,275,1428]" captionTargetId="figure@2.[151,1436,253,1446]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of the genus Rhabdops in the Western Ghats region of peninsular India. Circles denote records of Rhabdops olivaceus and squares denote records of Rhabdops aquaticus sp. nov. Green symbols indicates vouchered records and white symbols indicate non-vouchered reports. Arrow indicates the type locality of the respective species. See Appendix 3 for locality details." httpUri="https://zenodo.org/record/1051967/files/figure.png" pageId="18" pageNumber="45">Fig. 1</figureCitation>
) extending this to ca. 
<quantity id="4C8BD6B14518E578FC67FE3BFBDAC1E9" box="[970,1057,404,429]" metricMagnitude="5" metricUnit="m" metricValue="2.9" pageId="18" pageNumber="45" unit="km" value="290.0">290 km</quantity>
. However, the new species is thus far confirmed from relatively few localities (this paper, Srinavasulu 
<emphasis id="B907A7464518E578FC37FE16FC37C194" box="[922,972,439,464]" italics="true" pageId="18" pageNumber="45">et al</emphasis>
. 2014), and all of these are habitats with upland laterite plateaus associated with semi-evergreen forest (and it may occur in similar habitats in the region that we have not sampled). Although rich in geographically restricted biodiversity, many of these plateau and forest habitats, except those in protected areas, are impacted negatively by habitat degradation pressure and the lack of legal protection (
<bibRefCitation id="EFE206A54518E578FEF8FDE7FE1AC224" author="Watve" box="[341,481,584,608]" pageId="18" pageNumber="45" refString="Watve, A. (2013) Status review of Rocky plateaus in the northern Western Ghats and Konkan region of Maharashtra, India with recommendations for conservation and management. Journal of Threatened Taxa, 5, 3935 - 3962. http: // doi. org / 10.11609 / JoTT. o 3372.3935 - 62" type="journal article" year="2013">Watve 2013</bibRefCitation>
, 
<bibRefCitation id="EFE206A54518E578FE41FDE8FD16C224" author="Thorpe" box="[492,749,583,608]" pageId="18" pageNumber="45" refString="Thorpe, C. J. &amp; Watve, A. (2015) Lateritic plateaus in the northern Western Ghats, India: a review of bauxite mining restoration practices. Journal of Ecological Society, 28, 25 - 44." type="journal article" year="2015">Thorpe &amp; Watve 2015</bibRefCitation>
). It is possible that effective conservation of 
<taxonomicName id="4C7300D74518E578FB4BFDE6FA88C224" authority="Gower, 2017" authorityName="Gower" authorityYear="2017" box="[1254,1395,585,608]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="45" phylum="Chordata" rank="species" species="aquaticus" status="sp. nov.">
<emphasis id="B907A7464518E578FB4BFDE6FA88C224" box="[1254,1395,585,608]" italics="true" pageId="18" pageNumber="45">R. aquaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2341A3D4518E578FAD7FDE8FF33C2C1" pageId="18" pageNumber="45" rank="species">
<emphasis id="B907A7464518E578FAD7FDE8FF33C2C1" bold="true" pageId="18" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
will require protection of additional areas of suitable habitat across its range.
</paragraph>
</subSubSection>
</treatment>
</document>