220 lines
16 KiB
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220 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1145.96724" ID-Pensoft-Pub="1313-2970-1145-1" ID-Pensoft-UUID="ACBBD138B7375B0D9F63CE792A82F653" ID-ZooBank="830628C276CD4641BFC6144CD775ED6B" ModsDocID="1313-2970-1145-1" checkinTime="1675506069289" checkinUser="pensoft" docAuthor="Hedin, Marshal & Milne, Marc A." docDate="2023" docId="A9344A70202B59188ADE99CC5FFAAA85" docLanguage="en" docName="ZooKeys 1145: 1-130" docOrigin="ZooKeys 1145" docPubDate="2023-02-03" docSource="http://dx.doi.org/10.3897/zookeys.1145.96724" docTitle="Nesticus georgia Gertsch 1984" docType="treatment" docVersion="1" id="ACBBD138B7375B0D9F63CE792A82F653" lastPageNumber="1" masterDocId="ACBBD138B7375B0D9F63CE792A82F653" masterDocTitle="New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains" masterLastPageNumber="130" masterPageNumber="1" pageNumber="1" updateTime="1675506069289" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hedin, Marshal</mods:namePart>
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<mods:affiliation>Department of Biology, San Diego State University, San Diego, California 92182 - 4614, USA</mods:affiliation>
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<mods:nameIdentifier type="email">mhedin@sdsu.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Milne, Marc A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1943-0161</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, University of Indianapolis, Indianapolis, Indiana 46227, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-03</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1145</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>130</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1145.96724</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1145.96724</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1145-1</mods:identifier>
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<mods:identifier type="ZooBank">830628C276CD4641BFC6144CD775ED6B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">ACBBD138B7375B0D9F63CE792A82F653</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:A9344A70202B59188ADE99CC5FFAAA85" httpUri="http://treatment.plazi.org/id/A9344A70202B59188ADE99CC5FFAAA85" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="478XL" ID-ENA="60730" LSID="A9344A70-202B-5918-8ADE-99CC5FFAAA85" authority="Gertsch, 1984" authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus georgia" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="georgia">Nesticus georgia Gertsch, 1984</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Nesticus georgia genitalia A Georgia, Dade Co., Sitton's Cave, MCH specimen # 1015, ♂ palp, ventral. Georgia, Dade Co., Sitton's Cave, MCH specimen # 1014, epigynum, ventral (B), dorsal (C). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure51" httpUri="https://binary.pensoft.net/fig/804502" pageId="0" pageNumber="1">Fig. 51A-C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus georgia" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="georgia">Nesticus georgia</taxonomicName>
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Gertsch, 1984: 39, figs 156-158, 164-166.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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Type material:
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<materialsCitation collectingDate="1952-11-28" collectingDateMax="1992-09-23" collectingDateMin="1952-11-28" collectorName="E. J. Kuenzler, New, M. Hedin, K. Crandall, S. O'Kane" country="Georgia" county="Dade Co." location="Sitton's Cave" municipality="Sitton's Cave" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="Holotype">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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: USA -
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingCountry name="Georgia">Georgia</collectingCountry>
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,
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<collectingCounty>Dade Co.</collectingCounty>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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<typeStatus>holotype</typeStatus>
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;
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<location LSID="urn:lsid:plazi:treatment:A9344A70202B59188ADE99CC5FFAAA85:CF9C74B805B207867A1D8FBC57F016F1" country="Georgia" county="Dade Co." municipality="Sitton's Cave" name="Sitton's Cave">
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<normalizedToken originalValue="Sitton’s">Sitton's</normalizedToken>
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Cave
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</location>
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, near
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<location LSID="urn:lsid:plazi:treatment:A9344A70202B59188ADE99CC5FFAAA85:D3187BF3BF837E9EB8DE9086BAAE7B22" country="Georgia" county="Dade Co." municipality="Sitton's Cave" name="Trenton">Trenton</location>
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;
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<collectingDate value="1952-11-28">28 Nov. 1952</collectingDate>
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,
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<collectorName>E.J. Kuenzler</collectorName>
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leg.; AMNH.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectorName>New</collectorName>
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collections from type locality
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</emphasis>
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: -
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<emphasis bold="true" pageId="0" pageNumber="1">
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<location LSID="urn:lsid:plazi:treatment:A9344A70202B59188ADE99CC5FFAAA85:0FAD298115A87282C4D5CE96DCEC5F0D" country="Georgia" county="Dade Co." municipality="Sitton's Cave" name="Dade Co.">Dade Co.</location>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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,
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<specimenCount>♀</specimenCount>
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;
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<location LSID="urn:lsid:plazi:treatment:A9344A70202B59188ADE99CC5FFAAA85:B9EDE725D26B4C8347338F24FC2DE95F" country="Georgia" county="Dade Co." municipality="Sitton's Cave" name="Sitton's Cave">
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<normalizedToken originalValue="Sitton’s">Sitton's</normalizedToken>
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Cave
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</location>
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,
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<locationDeviation location="Trenton">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.609344" unit="mi" value="1.0">1 mi</quantity>
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E of Trenton
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</locationDeviation>
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;
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<collectingDate value="1991-09-30">30 Sep. 1991</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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,
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<collectorName>K. Crandall</collectorName>
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leg.; •
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<specimenCount type="male">2♂</specimenCount>
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,
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<specimenCount type="female">3♀</specimenCount>
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;
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<collectingMunicipality>
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<normalizedToken originalValue="Sitton’s">Sitton's</normalizedToken>
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Cave
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</collectingMunicipality>
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;
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<collectingDate value="1992-09-23">23 Sep. 1992</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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,
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<collectorName>
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S.
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<normalizedToken originalValue="O’Kane">O'Kane</normalizedToken>
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</collectorName>
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leg.
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</materialsCitation>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Nearly eyeless, long-legged taxon. Male palp most similar to that of
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<taxonomicName authorityName="Zigler & Milne" authorityYear="2022" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus lula" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lula">
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<emphasis italics="true" pageId="0" pageNumber="1">Nesticus lula</emphasis>
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</taxonomicName>
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, but with a spatulate tegular apophysis and details of the distal edge of the paracymbial ventral process with a sclerotized process projecting dorsally (Fig.
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<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Nesticus georgia genitalia A Georgia, Dade Co., Sitton's Cave, MCH specimen # 1015, ♂ palp, ventral. Georgia, Dade Co., Sitton's Cave, MCH specimen # 1014, epigynum, ventral (B), dorsal (C). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure51" httpUri="https://binary.pensoft.net/fig/804502" pageId="0" pageNumber="1">51A</figureCitation>
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). Ventral epigynum very close to similarly troglomorphic
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<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. lula" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lula">
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<emphasis italics="true" pageId="0" pageNumber="1">N. lula</emphasis>
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</taxonomicName>
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, difficult to separate based on epigynal morphology alone (compare Fig.
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<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Nesticus georgia genitalia A Georgia, Dade Co., Sitton's Cave, MCH specimen # 1015, ♂ palp, ventral. Georgia, Dade Co., Sitton's Cave, MCH specimen # 1014, epigynum, ventral (B), dorsal (C). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure51" httpUri="https://binary.pensoft.net/fig/804502" pageId="0" pageNumber="1">51B, C</figureCitation>
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to
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<bibRefCitation DOI="https://doi.org/10.1636/JoA-S-21-035" author="Zigler, KS" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="1" pagination="292 - 300" refId="B64" refString="Zigler, KS, Milne, MA, 2022. Two new Nesticus Thorell, 1869 (Araneae: Nesticidae) species from caves in northwest Georgia, USA. The Journal of Arachnology 50 (3): 292 - 300, DOI: https://doi.org/10.1636/JoA-S-21-035" title="Two new Nesticus Thorell, 1869 (Araneae: Nesticidae) species from caves in northwest Georgia, USA." url="https://doi.org/10.1636/JoA-S-21-035" volume="50" year="2022">Zigler and Milne 2022</bibRefCitation>
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: figs 2, 3). Both
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<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. georgia" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="georgia">
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<emphasis italics="true" pageId="0" pageNumber="1">N. georgia</emphasis>
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</taxonomicName>
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and
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<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. lula" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lula">
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<emphasis italics="true" pageId="0" pageNumber="1">N. lula</emphasis>
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</taxonomicName>
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are also similar in overall epigynal morphology to the more distantly related (but geographically proximate)
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<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. barri" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barri">
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<emphasis italics="true" pageId="0" pageNumber="1">N. barri</emphasis>
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</taxonomicName>
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(Fig.
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<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Nesticus barri and N. furtivus genitalia. N. barri - Tennessee, Marion Co., Tate Spring Cave, MCH 04 _ 050, ♂ palp ventral (A), epigynum ventral (C). N. furtivus - Tennessee, Hamilton Co., Raccoon Mountain Caverns, SE Chattanooga, MCH 00 _ 137, ♂ palp ventral (B), epigynum ventral (# 1660) (D)." figureDoi="10.3897/zookeys.1145.96724.figure48" httpUri="https://binary.pensoft.net/fig/804499" pageId="0" pageNumber="1">48C</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1145.96724.figure51" httpUri="https://binary.pensoft.net/fig/804502" pageId="0" pageNumber="1" start="Figure 51" startId="F51">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 51.</emphasis>
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<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus georgia" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="georgia">
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<emphasis italics="true" pageId="0" pageNumber="1">Nesticus georgia</emphasis>
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</taxonomicName>
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genitalia
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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Georgia, Dade Co.,
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<normalizedToken originalValue="Sitton’s">Sitton's</normalizedToken>
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Cave, MCH specimen #1015, ♂ palp, ventral. Georgia, Dade Co.,
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<normalizedToken originalValue="Sitton’s">Sitton's</normalizedToken>
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Cave, MCH specimen #1014, epigynum, ventral (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), dorsal (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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). Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution and natural history.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Known only from a handful of limestone caves from three adjacent counties in northwest Georgia (Fig.
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<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Distribution of Nesticus barri and Nesticus carteri groups. Type localities designated with yellow circles. State boundaries and major cities shown for geographic context. Dashed lines circumscribe known species distributions." figureDoi="10.3897/zookeys.1145.96724.figure49" httpUri="https://binary.pensoft.net/fig/804500" pageId="0" pageNumber="1">49</figureCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.904.1.1" author="Hedin, M" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 19" refId="B17" refString="Hedin, M, Dellinger, B, 2005. Descriptions of a new species and previously unknown males of Nesticus (Araneae: Nesticidae) from caves in eastern North America, with comments on species rarity. Zootaxa 904: 1 - 19, DOI: https://doi.org/10.11646/zootaxa.904.1.1" title="Descriptions of a new species and previously unknown males of Nesticus (Araneae: Nesticidae) from caves in eastern North America, with comments on species rarity." url="https://doi.org/10.11646/zootaxa.904.1.1" volume="904" year="2005">Hedin and Dellinger 2005</bibRefCitation>
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: fig. 1;
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<bibRefCitation DOI="https://doi.org/10.1371/journal.pone.0156751" author="Carver, LM" journalOrPublisher="Molecular Ecology" pageId="0" pageNumber="1" refId="B5" refString="Carver, LM, Perlaky, P, Cressler, A, Zigler, KS, 2016. Reproductive seasonality in Nesticus (Araneae: Nesticidae) cave spiders. PLoS ONE 11(6): e0156751. https://doi.org/10.1371/journal.pone.0156751" title="Reproductive seasonality in Nesticus (Araneae: Nesticidae) cave spiders. PLoS ONE 11 (6): e 0156751." url="https://doi.org/10.1371/journal.pone.0156751" year="2016">Carver et al. 2016</bibRefCitation>
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: fig. 2).
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<bibRefCitation author="Reeves, W" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" pagination="93 - 94" refId="B49" refString="Reeves, W, 1999. Cave-dwelling Nesticidae (Araneae) in the southeastern United States: New distribution records and notes on their bionomics. Insecta Mundi 13: 93 - 94" title="Cave-dwelling Nesticidae (Araneae) in the southeastern United States: New distribution records and notes on their bionomics." volume="13" year="1999">Reeves (1999)</bibRefCitation>
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summarized natural history information (microhabitat preference, fecundity, prey items, etc.) for topotypic
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<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus georgia" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="georgia">
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<emphasis italics="true" pageId="0" pageNumber="1">Nesticus georgia</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |