195 lines
20 KiB
XML
195 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/CompCytogen.v10i5.10944" ID-GBIF-Dataset="ba2552e1-e11b-4cbf-b242-689cbe66e3e3" ID-PMC="PMC5220643" ID-Pensoft-Pub="1993-078X-5-1" ID-PubMed="28105291" ID-ZBK="F9DCCAEDFA5548F2BDA851054261F1C5" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1993-078X-5-1" ModsDocOrigin="Comparative Cytogenetics 10 (5)" ModsDocTitle="Karyosystematics and molecular taxonomy of the anomalous blue butterflies (Lepidoptera, Lycaenidae) from the Balkan Peninsula" checkinTime="1482265596800" checkinUser="pensoft" docAuthor="Vishnevskaya, Maria S., Saifitdinova, Alsu F. & Lukhtanov, Vladimir A." docDate="2016" docId="0BAF352EA850378FDD78CE777765285B" docLanguage="en" docName="CompCytogen 10 (5): 1-85" docOrigin="CompCytogen 5" docSource="http://dx.doi.org/10.3897/CompCytogen.v10i5.10944" docTitle="Polyommatus (Agrodiaetus) timfristos Lukhtanov, Vishnevskaya & Shapoval, sp. n." docType="treatment" docUuid="58B77480-1FD0-423B-8FF9-BF39E79F177C" docUuidSource="ZooBank" docVersion="7" lastPageNumber="27" masterDocId="5576FFF5FF8450305E339C0D9F67FFEA" masterDocTitle="Karyosystematics and molecular taxonomy of the anomalous blue butterflies (Lepidoptera, Lycaenidae) from the Balkan Peninsula" masterLastPageNumber="85" masterPageNumber="1" pageNumber="25" updateTime="1668127016051" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Karyosystematics and molecular taxonomy of the anomalous blue butterflies (Lepidoptera, Lycaenidae) from the Balkan Peninsula</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Vishnevskaya, Maria S.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Saifitdinova, Alsu F.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Lukhtanov, Vladimir A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>CompCytogen</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2016</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>5</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>85</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/CompCytogen.v10i5.10944</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/CompCytogen.v10i5.10944</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1993-078X-5-1</mods:identifier>
|
||
<mods:identifier type="ZBK">F9DCCAEDFA5548F2BDA851054261F1C5</mods:identifier>
|
||
<mods:identifier type="ZooBank">F9DCCAEDFA5548F2BDA851054261F1C5</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="127878561" LSID="urn:lsid:zoobank.org:act:58B77480-1FD0-423B-8FF9-BF39E79F177C" httpUri="http://treatment.plazi.org/id/0BAF352EA850378FDD78CE777765285B" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
|
||
<subSubSection pageId="24" pageNumber="25" type="nomenclature">
|
||
<paragraph pageId="24" pageNumber="25">
|
||
<taxonomicName LSID="http://zoobank.org/58B77480-1FD0-423B-8FF9-BF39E79F177C" authority="Lukhtanov, Vishnevskaya & Shapoval" class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus (Agrodiaetus) timfristos" order="Lepidoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="timfristos" subGenus="Agrodiaetus">
|
||
<pageBreakToken pageId="24" pageNumber="25" start="start">Polyommatus</pageBreakToken>
|
||
(Agrodiaetus) timfristos Lukhtanov, Vishnevskaya & Shapoval
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="24" pageNumber="25">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph pageId="24" pageNumber="25">Holotype</paragraph>
|
||
<paragraph pageId="24" pageNumber="25">
|
||
(Fig. 16h). male, field code LR-08-247, GenBank accession number KY066725 for COI and KY081279 for ITS2; Greece, Timfristos Mt, Karpenisi,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="38.92433">38°55.460'N</geoCoordinate>
|
||
; 21°
|
||
<geoCoordinate direction="east" orientation="longitude" precision="55" value="47.605">47.605 E</geoCoordinate>
|
||
, 1270 m, 20 July 2008, V.A. Lukhtanov and N.A. Shapoval leg., deposited in Zoological Institute of the Russian Academy of Science (St. Petersburg).
|
||
</paragraph>
|
||
<paragraph pageId="24" pageNumber="25">COI barcode sequence of the holotype, 657 base pairs.</paragraph>
|
||
<paragraph pageId="24" pageNumber="25">ACATTATATTTTATTTTTGGAATTTGAGCAGGAATAGTAGGAACATCTCTAAGAATTTTAATTCGTATGGAATTAAGAACTCCTGGATCCTTAATTGGAAATGATCAAATTTATAATACTATTGTTACAGCCCATGCATTTATTATAATTTTTTTTATGGTTATACCTATTATAATTGGAGGATTTGGTAACTGATTAGTTCCCTTAATATTAGGAGCACCTGATATAGCTTTTCCACGATTAAATAATATGAGATTTTGATTATTACCGCCATCATTAATACTACTAATTTCTAGAAGAATTGTAGAAAATGGAGCAGGAACAGGATGAACAGTTTACCCCCCACTTTCATCAAATATTGCACATGGAGGATCATCTGTAGATTTAGCAATTTTCTCTCTTCATTTAGCGGGAATTTCTTCAATTTTAGGAGCAATTAATTTTATTACAACTATCATTAATATACGAGTAAATAATTTATCTTTTGATCAAATATCATTATTTATTTGAGCAGTGGGAATTACAGCATTATTATTACTTTTATCATTGCCTGTATTAGCTGGGGCAATTACCATATTATTAACAGATCGAAATCTTAATACCTCATTCTTTGACCCAGCTGGTGGAGGAGATCCAATTTTATATCAACATTTATTT</paragraph>
|
||
<paragraph pageId="24" pageNumber="25">Haploid chromosome number of the holotype n=38 (Fig. 4c).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph pageId="24" pageNumber="25">Paratypes.</paragraph>
|
||
<paragraph pageId="24" pageNumber="25">
|
||
Four males, field codes LR-08-255, LR-08-258, LR-08-273, LR-08-274, forewing length 17-18 mm, the same data as holotype. Male: field code LR-08-205, Greece, Parnassos,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="38.555183">38°33.311'N</geoCoordinate>
|
||
; 22°
|
||
<geoCoordinate direction="east" orientation="longitude" precision="55" value="34.3">34.300 E</geoCoordinate>
|
||
, 1750 m, 19 July 2008, V.A. Lukhtanov and N.A.Shapoval leg. Five females: forewing length 15-16 mm; Greece, Timfristos Mt, Karpenisi,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="38.9259">38°55.554'N</geoCoordinate>
|
||
; 21°
|
||
<geoCoordinate direction="east" orientation="longitude" precision="55" value="48.46">48.460 E</geoCoordinate>
|
||
, 1490 m, 21 July 2008, V.A. Lukhtanov and N.A. Shapoval leg. Two females: forewing length 14.5-15.5 mm; Greece, Parnassos,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="38.555183">38°33.311'N</geoCoordinate>
|
||
; 22°
|
||
<geoCoordinate direction="east" orientation="longitude" precision="55" value="34.3">34.300 E</geoCoordinate>
|
||
, 1750 m, 19 July 2008, V.A. Lukhtanov and N.A. Shapoval leg.. All paratypes are deposited in Zoological Institute of the Russian Academy of Science (St. Petersburg).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
|
||
<paragraph pageId="24" pageNumber="25">Males</paragraph>
|
||
<paragraph pageId="24" pageNumber="25">
|
||
(Fig. 16
|
||
<normalizedToken originalValue="d–i">d-i</normalizedToken>
|
||
). Forewing length 16.2-18.2 mm. Upperside: ground color completely brown. Discoidal, submarginal and antemarginal marking absent on both fore- and hindwings. Forewings with a developed sex brand and scaletuft. Fringe brown as ground color.
|
||
</paragraph>
|
||
<paragraph lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
Underside: ground color light brown with yellowish coffee-milk tint. Greenish blue basal suffusion very slight, nearly lacking. One basal black spot is present only on hindwings. Discoidal black spot is present on the forewings, but can be slightly seen on the hindwings (absent or vestigial). Postdiscal black ocelli are encircled by a whitish border. They are prominent on the forewings, forming a strongly curved row. Postdiscal black ocelli on the hindwing small. Submarginal and antemarginal mark
|
||
<pageBreakToken pageId="25" pageNumber="26" start="start">ing</pageBreakToken>
|
||
is absent on the forewings, and absent or vestigial on the hindwings. White streak on hindwings clearly visible. In one specimen the white streak is vestigial, in one the white streak is almost absent (can be slightly distinguished), and in one specimen there is an additional short streak between postdiscal and submarginal areas of the wing, straight under the main white streak. Fringe brown, slightly darker than the underside ground color.
|
||
</paragraph>
|
||
<paragraph pageId="25" pageNumber="26">
|
||
Genitalia: the male genitalia have a structure typical for other species of the subgenus
|
||
<taxonomicName lsidName="(Agrodiaetus)" pageId="25" pageNumber="26" rank="subGenus" subGenus="Agrodiaetus">Agrodiaetus</taxonomicName>
|
||
(Coutsis, 1986).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="25" pageNumber="26" type="description">
|
||
<paragraph pageId="25" pageNumber="26">Females</paragraph>
|
||
<paragraph pageId="25" pageNumber="26">
|
||
(Fig. 17
|
||
<normalizedToken originalValue="a–g">a-g</normalizedToken>
|
||
). Forewing length 15.8-17.5 mm.Upperside: ground color as in males, but lighter dark brown and without sex brand and scaletuft. Fringe greyish brown. Underside: ground color and general design as in males but fringes lighter-colored. Greenish blue basal suffusion almost invisible. White streak on hindwing underside is present in all paratypes and demonstrates a variable level of reduction.
|
||
</paragraph>
|
||
<caption pageId="25" pageNumber="26">
|
||
<paragraph pageId="25" pageNumber="26">
|
||
Figure 17. Paratypes of
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus timfristos" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="timfristos">Polyommatus timfristos</taxonomicName>
|
||
sp. n. (females). a, b samples from Parnassos Mt
|
||
<normalizedToken originalValue="c–g">c-g</normalizedToken>
|
||
samples from Timfristos Mt.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="diagnosis">
|
||
<paragraph pageId="25" pageNumber="26">Diagnosis.</paragraph>
|
||
<paragraph pageId="25" pageNumber="26">
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus timfristos" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="timfristos">Polyommatus timfristos</taxonomicName>
|
||
(n=38) differs by at least three fixed chromosome fusions/fissions from the most closely related and allopatric
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus orphicus" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subspecies" species="orphicus" subspecies="orphicus">Polyommatus orphicus orphicus</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus orphicus" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subspecies" species="orphicus" subspecies="eleniae">Polyommatus orphicus eleniae</taxonomicName>
|
||
(n=41-42).
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus timfristos" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="timfristos">Polyommatus timfristos</taxonomicName>
|
||
(n=38) differs by at least nine fixed chromosome fusions/fissions from allopatric
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus aroaniensis" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="aroaniensis">Polyommatus aroaniensis</taxonomicName>
|
||
(n=47). From the closely related
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus orphicus" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="orphicus">Polyommatus orphicus</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus aroaniensis" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="aroaniensis">Polyommatus aroaniensis</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus timfristos" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="timfristos">Polyommatus timfristos</taxonomicName>
|
||
differs also by a number of nucleotide substitutions within the studied 657-bp fragment of the mitochondrial COI gene.
|
||
</paragraph>
|
||
<paragraph pageId="26" pageNumber="27">
|
||
<pageBreakToken pageId="26" pageNumber="27" start="start">The</pageBreakToken>
|
||
chromosome number in
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus timfristos" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="timfristos">Polyommatus timfristos</taxonomicName>
|
||
(n=38) is similar (but not identical) to that found in
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus humedasae" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="humedasae">Polyommatus humedasae</taxonomicName>
|
||
(n=39,
|
||
<bibRefCitation author="Vila, R" journalOrPublisher="Biological Journal of the Linnean Society" pageId="37" pageNumber="38" pagination="130 - 154" title="How common are dot-like distribution ranges? Taxonomical oversplitting in Western European Agrodiaetus (Lepidoptera, Lycaenidae) revealed by chromosomal and molecular markers." url="https://doi.org/10.1111/j.1095-8312.2010.01481.x" volume="101" year="2010">Vila et al. 2010</bibRefCitation>
|
||
). However, we are not sure that these karyotypes are related in their origin because they are not found in proximity and separated by an area where
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus orphicus" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="orphicus">Polyommatus orphicus</taxonomicName>
|
||
with n=41-42 is distributed. With respect to COI barcodes, the pair
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus timfristos" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="timfristos">Polyommatus timfristos</taxonomicName>
|
||
/
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus humedasae" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="humedasae">Polyommatus humedasae</taxonomicName>
|
||
is more differentiated than pairs
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus timfristos" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="timfristos">Polyommatus timfristos</taxonomicName>
|
||
/
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus aroaniensis" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="aroaniensis">Polyommatus aroaniensis</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus timfristos" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="timfristos">Polyommatus timfristos</taxonomicName>
|
||
/
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus orphicus" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="orphicus">Polyommatus orphicus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="26" pageNumber="27">
|
||
From sympatric and syntopic
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus ripartii" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subspecies" species="ripartii" subspecies="pelopi">Polyommatus ripartii pelopi</taxonomicName>
|
||
the new species can usually be distinguished by the absence of submarginal marking and strong reduction of greenish blue basal suffusion. These characteristics are usually (but not always) better expressed in
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus ripartii" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subspecies" species="ripartii" subspecies="pelopi">Polyommatus ripartii pelopi</taxonomicName>
|
||
specimens. In doubtful cases, the separation is only possible on the base of chromosomal and molecular markers since these species are different: the chromosome number of
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus ripartii" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subspecies" species="ripartii" subspecies="pelopi">Polyommatus ripartii pelopi</taxonomicName>
|
||
is n=90; they also have fixed differences in 33 positions within the studied 657-bp fragment of COI gene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="26" pageNumber="27" type="ecology">
|
||
<paragraph pageId="26" pageNumber="27">Ecology.</paragraph>
|
||
<paragraph pageId="26" pageNumber="27">
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus timfristos" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="timfristos">Polyommatus timfristos</taxonomicName>
|
||
sp. n. inhabits xerothermic and xeromontane localities and dry meadows from 1200 to 1800 m altitude (Figs 32-35). It was found in complete syntopy with
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus ripartii" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subspecies" species="ripartii" subspecies="pelopi">Polyommatus ripartii pelopi</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Polyommatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyommatus admetus" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="admetus">Polyommatus admetus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="26" pageNumber="27" type="etymology">
|
||
<paragraph pageId="26" pageNumber="27">Etymology.</paragraph>
|
||
<paragraph pageId="26" pageNumber="27">Timfristos is a mountain in the eastern part of Evrytania and the western part of Phthiotis in Central Greece. The name is a noun.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |