235 lines
41 KiB
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235 lines
41 KiB
XML
<document ID-CLB-Dataset="6376" ID-DOI="http://dx.doi.org/10.3897/asp.79.e62735" ID-GBIF-Dataset="2fae9b0a-ea4d-4715-9ee6-93a215740e7c" ID-Pensoft-Pub="1864-8312-79-151" ID-Pensoft-UUID="8842F15397AF5A1BBD35D4ECB5A874CE" ID-ZooBank="414052252BB34048A62903AF1569C616" ModsDocID="1864-8312-79-151" checkinTime="1622492420826" checkinUser="pensoft" docAuthor="Molineri, Carlos, Dias, Lucimar G. & Zuniga, Maria del Carmen" docDate="2021" docId="F1E5B4CD8E9955169A748D682313B230" docLanguage="en" docName="ArthropodSystPhyl 79: 151-170" docOrigin="Arthropod Systematics & amp; Phylogeny 79" docPubDate="2021-05-31" docSource="http://dx.doi.org/10.3897/asp.79.e62735" docTitle="Tortopsis toro Molineri & Dias & Zúñiga 2021, sp. nov." docType="treatment" docUuid="834A4E7E-D5A7-42F5-B786-930927D0CC43" docUuidSource="ZooBank" docVersion="5" id="8842F15397AF5A1BBD35D4ECB5A874CE" lastPageNumber="151" masterDocId="8842F15397AF5A1BBD35D4ECB5A874CE" masterDocTitle="New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species" masterLastPageNumber="170" masterPageNumber="151" pageNumber="151" updateTime="1732802382458" updateUser="ExternalLinkService">
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<mods:title id="B8B43071D8D993504595C43F11928A9D">New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species</mods:title>
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<mods:namePart id="C55C68612AF47CE36D57D55089EDFB55">Molineri, Carlos</mods:namePart>
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<mods:affiliation id="421DE8FDBF17AFB381AA3CD6FAE76539">Instituto de Biodiversidad Neotropical, CONICET (National Council of Scientific Research), Universidad Nacional de Tucuman, Ciudad Universitaria Horco Molle, CP 4000, Argentina; Carlos Molineri [carlosmolineri @ gmail. com]</mods:affiliation>
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<mods:nameIdentifier id="5008B77FE9E47707A51553678DB70342" type="email">carlosmolineri@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C6BDE2252FFFB0E2D6E0B31DC0A1971B">Dias, Lucimar G.</mods:namePart>
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<mods:affiliation id="850A1057C11F49FB17A818F2592EDF22">Facultad de Ciencias Exactas y Naturales, Grupo de Investigacion BIONAT, Universidad de Caldas, Caldas, Colombia; Lucimar G. Dias [lucimar. dias @ ucaldas. edu. co]</mods:affiliation>
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<mods:namePart id="85768D74F7196CA8229AE6D316C70775">Zuniga, Maria del Carmen</mods:namePart>
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<mods:affiliation id="45E80CA75D54F7F01D7980247D1CC09F">Grupo de Investigaciones Entomologicas, Departamento de Biologia, Universidad del Valle, Cali, Colombia; Maria del Carmen Zuniga [maczuniga @ gmail. com]</mods:affiliation>
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<mods:title id="CE125CD3EC8FABB5EA90571A662BD862">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:date id="2824CE73DC9D5B85A38791B43D882D43">2021</mods:date>
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<treatment id="F1E5B4CD8E9955169A748D682313B230" ID-GBIF-Taxon="182473839" LSID="urn:lsid:zoobank.org:act:834A4E7E-D5A7-42F5-B786-930927D0CC43" httpUri="http://treatment.plazi.org/id/F1E5B4CD8E9955169A748D682313B230" lastPageNumber="151" pageId="0" pageNumber="151" scope_family="Polymitarcyidae" scope_order="Ephemeroptera">
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<subSubSection id="BF783BA923691E932662786675818A08" pageId="0" pageNumber="151" type="nomenclature">
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<paragraph id="6EF4A6519C9134C85D96851CCD1CBE1A" pageId="0" pageNumber="151">
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<taxonomicName id="31128766A8CBD8E109A487362B1B2F8E" LSID="https://zoobank.org/834A4E7E-D5A7-42F5-B786-930927D0CC43" authority="Molineri & Dias & Zúñiga, 2021" authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis toro" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">Tortopsis toro</taxonomicName>
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<taxonomicNameLabel id="182A1B991F822295BD1D6CE6B2DE28D8" pageId="0" pageNumber="151">sp. nov.</taxonomicNameLabel>
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<figureCitation id="7451B5D41DBF338EF0AF449BCFE95EFF" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">Figs 1-6</figureCitation>
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<figureCitation id="3A895B2C14923291425E72A4B1E1034D" captionStart="Figures 7–14" captionStartId="F2" captionText="Figures 7 - 14. Tortopsis toro sp. nov., nymph. 7, labrum, dorsal view; 8, left mandible, inner view, 9, right mandible, inner view; 10, right mandible, detail of tusk, ventral view (arrow indicates stout spine near inner margin); 11, hypopharynx, dorsal view; 12, labium, ventral view; 13, left maxilla, ventral view (arrow indicates membranous gill); 14, right maxilla, ventral view." figureDoi="10.3897/asp.79.e62735.figures7-14" httpUri="https://binary.pensoft.net/fig/550218" pageId="0" pageNumber="151">, 7-14</figureCitation>
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<figureCitation id="EFD0008A8906AFCF4DA28D825A308BED" captionStart="Figures 15–18" captionStartId="F3" captionText="Figures 15 - 18. Tortopsis toro sp. nov., nymph. 15, fore leg, dorsal view; 16, fore tarsal claw; 17, middle leg, dorsal view; 18, hind leg, ventral view." figureDoi="10.3897/asp.79.e62735.figures15-18" httpUri="https://binary.pensoft.net/fig/550219" pageId="0" pageNumber="151">, 15-18</figureCitation>
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<figureCitation id="F155314E7DFD3D34B7CB6C5BCFF03EAD" captionStart="Figures 19–26" captionStartId="F4" captionText="Figures 19 - 26. Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted)." figureDoi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151">, 19-26</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="96AC0D0BA3937557660A5BE62C2056C4" pageId="0" pageNumber="151" type="type material">
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<paragraph id="DDA546977D7C7B7067081C0340C65E01" pageId="0" pageNumber="151">Type material.</paragraph>
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<paragraph id="180FA479676082FA0A55CDB6E13C48B8" pageId="0" pageNumber="151">
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Holotype male imago (CEBUC) from Colombia,
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<normalizedToken id="9F053B4352499887B41B250E4A35A730" originalValue="Caquetá">Caqueta</normalizedToken>
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, Curillo,
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<normalizedToken id="D09A28EFABFAD39B1E492CD5AC12018C" originalValue="río">rio</normalizedToken>
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<normalizedToken id="8D15C6B85B562ED141FA868008A3A183" originalValue="Caquetá">Caqueta</normalizedToken>
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, 220 m,
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<geoCoordinate id="E7797A9F6C1B9FAE8CDB19128F4B4272" degrees="01" direction="north" minutes="01" orientation="latitude" precision="15" seconds="44" value="1.0288888">
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01°01
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<normalizedToken id="EBF797B7EFCEC57D410ACBDBB9CBBFAF" originalValue="’44”">'44"</normalizedToken>
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N
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</geoCoordinate>
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;
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<geoCoordinate id="E04BEECA722C2F75CF65E2FE1E554FE1" degrees="75" direction="west" minutes="55" orientation="longitude" precision="15" seconds="9" value="-75.91917">
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75°55
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<normalizedToken id="B40A45E8DFAB3902C0FA3519B4B11D44" originalValue="’9”">'9"</normalizedToken>
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W
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</geoCoordinate>
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, 20.ix.2018, LG Dias, MC
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<normalizedToken id="36DD5086BB5407C78DFFED1941DACEB2" originalValue="Zúñiga">Zuniga</normalizedToken>
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, B Toro, JP Chaux and C Molineri cols. Paratypes: 1 reared male adult with nymphal exuviae (IBN); 5 male imagos, 3 female imagos and 13 nymphs (IBN); 10 males imagos, 5 females imagos and 10 nymphs (CEBUC); 5 male imagos, 3 female imagos and 10 nymphs (MUSENUV), same data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection id="F49A3250D07E7BC1DFD5A3C3230DA4CA" pageId="0" pageNumber="151" type="diagnosis">
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<paragraph id="62BF851A8E590A888F790416EAE1B28A" pageId="0" pageNumber="151">Diagnosis.</paragraph>
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<paragraph id="A4D8C3A8ECDAB6FA85FF85FF66A3A5A5" pageId="0" pageNumber="151">
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<taxonomicName id="774D8F40A642B73EC98C09C844931DA2" authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis toro" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="toro">
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<emphasis id="B75DC310E9ABF12DDF5BA3457DA0F942" italics="true" pageId="0" pageNumber="151">Tortopsis toro</emphasis>
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</taxonomicName>
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sp. nov., known from imagos of both sexes and nymphs, can be distinguished from all other species of the genus by: In adults, 1) forewing length ranges between 12.2-13.2 mm (male) and 15.5-17.0 mm (female); 2) hyaline wings, with brownish veins and costal margin shaded slightly gray; 3) parastyli almost straight in lateral view, slightly curved medially, with a longitudinal ventral furrow (Figs
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<figureCitation id="97354B4C16770E9D0AD239F449A38407" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">2-3</figureCitation>
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,
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<figureCitation id="ECB096ACBEFBEC9B1A280AFBB51B0A8D" captionStart="Figures 19–26" captionStartId="F4" captionText="Figures 19 - 26. Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted)." figureDoi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151">19-21</figureCitation>
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); 4) penis apically widened and flattened, with rounded semicircular apical spine (Fig.
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<figureCitation id="A5E5A0696FC4B06E4995B7468E13EDC2" captionStart="Figures 19–26" captionStartId="F4" captionText="Figures 19 - 26. Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted)." figureDoi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151">20</figureCitation>
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); 5) female parastyli receptors with rounded and elongated sockets (Fig.
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<figureCitation id="C820587D3B6273E905BC7013A9C5FDD8" captionStart="Figures 19–26" captionStartId="F4" captionText="Figures 19 - 26. Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted)." figureDoi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151">22</figureCitation>
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); 6) head shaded black among ocelli, occiput with very light gray anastomosed lines (Figs
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<figureCitation id="D97DB887E8218A5CDF5CE2374C207087" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">1</figureCitation>
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,
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<figureCitation id="886B7E754AF1F366A341F13FF6892485" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">6</figureCitation>
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); 7) coloration as in Fig.
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<figureCitation id="CF06C011285FF78319E5ECD932B6331D" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">1</figureCitation>
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. The nymphs can be separated from the others of the genus by: 1) mandibles at most with 2 stout small spines on inner margin of tusk (basally to large subdistal tubercle, see arrow in Fig.
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<figureCitation id="2E6E2B9A46B6F0D24D07FC4963C21318" captionStart="Figures 19–26" captionStartId="F4" captionText="Figures 19 - 26. Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted)." figureDoi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151">24</figureCitation>
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); 2) outer margin of tusk, just before distal spur, with a marked indentation (arrow in Fig.
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<figureCitation id="24AC72B7EC8F2E22DBEF78E9FBFEEF87" captionStart="Figures 19–26" captionStartId="F4" captionText="Figures 19 - 26. Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted)." figureDoi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151">25</figureCitation>
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); 3) head shaded with gray on occiput as in Fig.
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<figureCitation id="0DE4EE655F6CF15B992D55DC122BDC4A" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">6</figureCitation>
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;
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<figureCitation id="37D503BC1628C5B0670A8083350D413E" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">4</figureCitation>
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) developing fore wing buds pigmented with gray slightly along costal margin and base of longitudinal veins (similar to
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<taxonomicName id="E26400A4CBEC316C72C40A66ABAC3B23" lsidName="T. puella" pageId="0" pageNumber="151" rank="species" species="puella">
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<emphasis id="0CEB329C3B5BCD5109FB5A6188832C7E" italics="true" pageId="0" pageNumber="151">T. puella</emphasis>
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</taxonomicName>
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in
|
||
<bibRefCitation id="4A0FDEF27E109C3CFC00253F124FF4F3" DOI="https://doi.org/10.1080/01650420701694201" author="Molineri, C" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="151" pagination="7 - 19" refId="B13" refString="Molineri, C, 2008. The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae). Aquatic Insects 30: 7 - 19, DOI: https://doi.org/10.1080/01650420701694201" title="The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae)." url="https://doi.org/10.1080/01650420701694201" volume="30" year="2008">Molineri 2008</bibRefCitation>
|
||
: figure 16).
|
||
</paragraph>
|
||
<paragraph id="052D8785F93A93A4293971C770D72993" pageId="0" pageNumber="151">
|
||
<emphasis id="4D6B0B85FFF8C0074A707DDE45074A47" italics="true" pageId="0" pageNumber="151">Male imago</emphasis>
|
||
. Length (mm): body, 14.0-14.5; forewing, 12.2-13.2; hindwing, 5.9-6.1; foreleg, 5.3-5.9; cerci, 32.0-36.0. General coloration yellowish white shaded with gray dorsally (Fig.
|
||
<figureCitation id="4ED61C282C948CB1F144D86302602E33" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">1</figureCitation>
|
||
). Head. Coloration yellowish white except black area among ocelli (Fig.
|
||
<figureCitation id="307440F46802BDFDDAF4A339FC113C1B" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">1</figureCitation>
|
||
). Antennae: scape and pedicel yellowish white completely shaded with gray, flagellum hyaline. Thorax. Pronotum hyaline, shaded with gray dorsally (Fig.
|
||
<figureCitation id="D73F1F9913E3D34D50AF1D19CD13ECF6" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">1</figureCitation>
|
||
). Mesonotum whitish yellow shaded with gray along sutures but not medially. Metanotum whitish yellow shaded gray medially. Thoracic pleura and sterna whitish yellow. Legs yellowish white, shaded with grayish on fore tibia and tarsus, rest of leg (including femur and claws, not shaded). Wings. Membrane hyaline, shaded slightly gray along C and Sc areas; longitudinal and cross veins brownish. Abdomen. Whitish translucent with gray shading dorsally, except on medial band. Terga I-III shaded gray on submedial rounded areas; terga IV-VII similarly shaded but a longitudinal darker strip is visible submedially (Fig.
|
||
<figureCitation id="1510CD7D62F64FD0436C5CEBDB78372A" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">1</figureCitation>
|
||
); terga VIII-IX more widely shaded except on small marks and sigilla (Fig.
|
||
<figureCitation id="856611CD06AF1D944396790378F7A277" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">1</figureCitation>
|
||
); tergum X with whitish median area and blackish margins. Abdominal sterna pale, without dark shadings, except sternum IX with a small median gray mark on posterior margin. Genitalia (Figs
|
||
<figureCitation id="D85AC03EA11A818096CE6A0CAED1FBE2" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">2-3</figureCitation>
|
||
,
|
||
<figureCitation id="523D7DC27A4A822CD871769A9797801D" captionStart="Figures 19–26" captionStartId="F4" captionText="Figures 19 - 26. Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted)." figureDoi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151">19-21</figureCitation>
|
||
): whitish except parastylus yellowish. Parastylus almost straight in lateral view, slightly curved medially, with a longitudinal ventrolateral furrow along its entire length; penis distally widened, semicircular
|
||
<normalizedToken id="666D7BDCE5AF3A8BA65E3DD71C96184D" originalValue="“spine”">"spine"</normalizedToken>
|
||
at apex of penis. Caudal filament whitish translucent.
|
||
</paragraph>
|
||
<caption id="DBE06072BE0E2E0300FFA30197DB83F8" doi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151" start="Figures 1–6" startId="F1">
|
||
<paragraph id="D4A451DFD0F736B6C3C9D994456FDA46" pageId="0" pageNumber="151">
|
||
<emphasis id="7A2A090A0B540218103D87D91024EB05" bold="true" pageId="0" pageNumber="151">Figures 1-6.</emphasis>
|
||
<taxonomicName id="AF6237C4898BB38AB2A12E19E980FAC3" authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis toro" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="7FFB721C8D623672B89672EEA7EA4055" italics="true" pageId="0" pageNumber="151">Tortopsis toro</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="643EBB9B253A58E775DEC461C195342E" bold="true" pageId="0" pageNumber="151">sp. nov.</emphasis>
|
||
Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3151C24C2F30F8C637999BF1A65718CB" pageId="0" pageNumber="151">
|
||
<emphasis id="82D7845287D23DADFA8B573C4CCAF951" italics="true" pageId="0" pageNumber="151">Female imago</emphasis>
|
||
. Length (mm): body, 18.0-18.5; forewing, 15.5-17.0; hindwing, 6.5-7.5; cerci, 4.3-4.8. General coloration as in male except darker markings. Head. Coloration black between ocelli, with posterior and submedian grayish lines on occiput (similar to Fig.
|
||
<figureCitation id="7BF4023F985004641502F3ACA1372CA0" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">4</figureCitation>
|
||
). Thorax. Furcasterna shaded with light gray medially. Wings: Membrane hyaline slightly tinged with yellow. Abdomen. shaded more extensively with gray on terga. Parastyli receptors on sternum VIII with elongated and rounded sockets (Fig.
|
||
<figureCitation id="EF81B0A1B69150123E99331940D67F4B" captionStart="Figures 19–26" captionStartId="F4" captionText="Figures 19 - 26. Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted)." figureDoi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151">22</figureCitation>
|
||
). Light gray marks medially on sterna VIII-IX.
|
||
</paragraph>
|
||
<paragraph id="18443A754D1DD2F5D206D14CA3A9D08F" pageId="0" pageNumber="151">
|
||
<emphasis id="71378BBA675DC2025EC65715C4BDF29D" italics="true" pageId="0" pageNumber="151">Egg.</emphasis>
|
||
Suboval. Length, 410-420
|
||
<normalizedToken id="A9BFD53CD92ACD347CDEB72FA872AAC3" originalValue="μm">μm</normalizedToken>
|
||
; width, 320-340
|
||
<normalizedToken id="0B6416D9A138F3907B8CBF844FACBC6E" originalValue="μm">μm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9216A058D539DE0C31096EAB5ADF7981" pageId="0" pageNumber="151">
|
||
<emphasis id="7AB33C9DA96A886420869A5408528327" italics="true" pageId="0" pageNumber="151">Nymph</emphasis>
|
||
(mature). Length of body (from apex of tusks to apex of abdominal tergum X): female, 22.0-26.0 mm; male, 14.0-19.0 mm. General coloration yellowish white with gray markings dorsally (Figs
|
||
<figureCitation id="C5DC45F32E1BCC21A90D938484278A57" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">4-6</figureCitation>
|
||
). Head. Coloration whitish shaded gray among ocelli, occiput with very light gray anastomosed lines submedially as Fig.
|
||
<figureCitation id="D2CCEBAF8B99806B0A0A65F9A43205A4" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">6</figureCitation>
|
||
. Antennae and mouthparts whitish except apex of tusk orangeish, and spines yellowish. Finger-like gill present near base of maxillae (arrow in Fig.
|
||
<figureCitation id="1271310A45340B82EC4C8F8A5ACAEAA1" captionStart="Figures 7–14" captionStartId="F2" captionText="Figures 7 - 14. Tortopsis toro sp. nov., nymph. 7, labrum, dorsal view; 8, left mandible, inner view, 9, right mandible, inner view; 10, right mandible, detail of tusk, ventral view (arrow indicates stout spine near inner margin); 11, hypopharynx, dorsal view; 12, labium, ventral view; 13, left maxilla, ventral view (arrow indicates membranous gill); 14, right maxilla, ventral view." figureDoi="10.3897/asp.79.e62735.figures7-14" httpUri="https://binary.pensoft.net/fig/550218" pageId="0" pageNumber="151">13</figureCitation>
|
||
). Mandibular tusks with 0-2 stout spines on inner margin, basal to subdistal tubercle (see arrow in Fig.
|
||
<figureCitation id="4BFAB29F609DD964D7B70AA22FC72CDF" captionStart="Figures 19–26" captionStartId="F4" captionText="Figures 19 - 26. Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted)." figureDoi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151">24</figureCitation>
|
||
); outer margin of tusk with row of 18-26 stout spines (Figs
|
||
<figureCitation id="DA8AB31084534C3994C42E9B786EF787" captionStart="Figures 7–14" captionStartId="F2" captionText="Figures 7 - 14. Tortopsis toro sp. nov., nymph. 7, labrum, dorsal view; 8, left mandible, inner view, 9, right mandible, inner view; 10, right mandible, detail of tusk, ventral view (arrow indicates stout spine near inner margin); 11, hypopharynx, dorsal view; 12, labium, ventral view; 13, left maxilla, ventral view (arrow indicates membranous gill); 14, right maxilla, ventral view." figureDoi="10.3897/asp.79.e62735.figures7-14" httpUri="https://binary.pensoft.net/fig/550218" pageId="0" pageNumber="151">8-10</figureCitation>
|
||
,
|
||
<figureCitation id="9BF393A099ECC7F76FF6BE859646AF91" captionStart="Figures 19–26" captionStartId="F4" captionText="Figures 19 - 26. Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted)." figureDoi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151">24-25</figureCitation>
|
||
). Thorax. Pronotum, anterior ring shaded black almost completely including the anterolateral pointed projections, posterior ring patterned as in Fig.
|
||
<figureCitation id="86D0DEE889597DE2379753FDD0F8C3E8" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">6</figureCitation>
|
||
, both with a median pale line. Meso- and metanotum with gray and black marks, lighter on median area; wing buds whitish, pigmented with gray slightly along costal margin and base of longitudinal veins. Thoracic pleura and sterna whitish. Legs whitish, with yellowish setae and apex of tarsal claws yellowish (Figs
|
||
<figureCitation id="A479F4F87267C7D6B672911CF3BDEFFF" captionStart="Figures 15–18" captionStartId="F3" captionText="Figures 15 - 18. Tortopsis toro sp. nov., nymph. 15, fore leg, dorsal view; 16, fore tarsal claw; 17, middle leg, dorsal view; 18, hind leg, ventral view." figureDoi="10.3897/asp.79.e62735.figures15-18" httpUri="https://binary.pensoft.net/fig/550219" pageId="0" pageNumber="151">15-18</figureCitation>
|
||
). Abdomen. Color pattern as described for the adults. Gills: vestigial gills I translucent; gills II-VII well developed, formed by a pair of large whitish lamellae, the outer (dorsal) lamellae of each pair is shaded with black on a medio longitudinal band (thinner on gill II), the inner (ventral) lamellae of each pair show only a thin brownish line along trachea. Genital rudiments of males showing distally widened penis. Caudal filaments yellowish white.
|
||
</paragraph>
|
||
<caption id="BE425DE4E1D0F0E9DFB0E1610319FA42" doi="10.3897/asp.79.e62735.figures7-14" httpUri="https://binary.pensoft.net/fig/550218" pageId="0" pageNumber="151" start="Figures 7–14" startId="F2">
|
||
<paragraph id="799439816221367D605A29B7B2FBDCAB" pageId="0" pageNumber="151">
|
||
<emphasis id="A532023FE5BDA72C353F10F8F5A0064A" bold="true" pageId="0" pageNumber="151">Figures 7-14.</emphasis>
|
||
<taxonomicName id="B610B04103E683E9C52FC098BD1DA883" authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis toro" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="D70C71CE91FE24E6261323A56639EE4A" italics="true" pageId="0" pageNumber="151">Tortopsis toro</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E80F74BCAA238A5340EE321B606FCC85" bold="true" pageId="0" pageNumber="151">sp. nov.</emphasis>
|
||
, nymph. 7, labrum, dorsal view; 8, left mandible, inner view, 9, right mandible, inner view; 10, right mandible, detail of tusk, ventral view (arrow indicates stout spine near inner margin); 11, hypopharynx, dorsal view; 12, labium, ventral view; 13, left maxilla, ventral view (arrow indicates membranous gill); 14, right maxilla, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="A8EF386CEADA5A9C9F01A2152CE45809" pageId="0" pageNumber="151" type="etymology">
|
||
<paragraph id="5368F8FACBDE0509AC026426A6BC5BA1" pageId="0" pageNumber="151">Etymology.</paragraph>
|
||
<paragraph id="A701664905695D786F57C96AB60199AC" pageId="0" pageNumber="151">This species is dedicated to Dr. Beatriz Toro Restrepo, Universidad de Caldas, in recognition of her work in environmental education, friendship and contributions in fieldwork with aquatic insects of Colombia.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="361C4D67A8B3339805325C963575B440" pageId="0" pageNumber="151" type="distribution">
|
||
<paragraph id="AFF7DD1AC8DE2245D90473943C18277C" pageId="0" pageNumber="151">Distribution.</paragraph>
|
||
<paragraph id="ADE48DFF16E88AB795F1010FAEA38721" pageId="0" pageNumber="151">Known only from the type locality.</paragraph>
|
||
<caption id="ABD1CB1B7B39B415A0C9CEA4A5F102A0" doi="10.3897/asp.79.e62735.figures15-18" httpUri="https://binary.pensoft.net/fig/550219" pageId="0" pageNumber="151" start="Figures 15–18" startId="F3">
|
||
<paragraph id="1DFD285FC9A9D43109BC67CBB143B8B1" pageId="0" pageNumber="151">
|
||
<emphasis id="55CA2836BD4E158A940C9984C026906E" bold="true" pageId="0" pageNumber="151">Figures 15-18.</emphasis>
|
||
<taxonomicName id="561E2C6B41B998177EAB757D29F878EB" authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis toro" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B45A64EF0664331F5C1A38BBDAD97A9F" italics="true" pageId="0" pageNumber="151">Tortopsis toro</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7642D403F24657B63FCCAE1AA7F39CA" bold="true" pageId="0" pageNumber="151">sp. nov.</emphasis>
|
||
, nymph. 15, fore leg, dorsal view; 16, fore tarsal claw; 17, middle leg, dorsal view; 18, hind leg, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5E38A9A7932C8FB87A929743FCF51D8D" doi="10.3897/asp.79.e62735.figures19-26" httpUri="https://binary.pensoft.net/fig/550220" pageId="0" pageNumber="151" start="Figures 19–26" startId="F4">
|
||
<paragraph id="FD00F8A0F767695C4E91D356771C7BA6" pageId="0" pageNumber="151">
|
||
<emphasis id="97A783A82E375BDE9052C588FD7B56DB" bold="true" pageId="0" pageNumber="151">Figures 19-26.</emphasis>
|
||
<taxonomicName id="BC7F9E05E46AD195702D4C412795F375" authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis toro" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="4F6446EE5B77B73315E3C09A6FE64DC1" italics="true" pageId="0" pageNumber="151">Tortopsis toro</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FDC71799833AD788F04FB92017C2815C" bold="true" pageId="0" pageNumber="151">sp. nov.</emphasis>
|
||
, imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |